290 lines
33 KiB
XML
290 lines
33 KiB
XML
<document id="269A62C3606E64C8F585DAC7F614CB5C" ID-CLB-Dataset="30668" ID-DOI="http://dx.doi.org/10.3897/nl.41.21584" ID-GBIF-Dataset="cdeba65e-5c0f-4b24-baa7-75a1273594e1" ID-Pensoft-Pub="2367-5365-1-99" ID-ZooBank="C2D8F31797FF44338A6385AF8488783D" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2367-5365-1-99" ModsDocOrigin="Nota Lepidopterologica 41 (1)" ModsDocTitle="New data on the taxonomy, morphology and distribution of Naardaineffectalis (Walker, 1859) (Lepidoptera, Erebidae, Hypeninae)" checkinTime="1519429591526" checkinUser="pensoft" docAuthor="Toth, Balazs" docDate="2018" docId="E7263586C77072F295F499021FD1FFBA" docLanguage="en" docName="NotaLepidopt 41(1): 99-106" docOrigin="Nota Lepidopterologica 41 (1)" docSource="http://dx.doi.org/10.3897/nl.41.21584" docTitle="Naarda ineffectalis Walker 1859" docType="treatment" docVersion="7" lastPageNumber="103" masterDocId="FFEC2654FF8AFFFC9E03FFB0FFDDFFEF" masterDocTitle="New data on the taxonomy, morphology and distribution of Naardaineffectalis (Walker, 1859) (Lepidoptera, Erebidae, Hypeninae)" masterLastPageNumber="106" masterPageNumber="99" pageNumber="100" updateTime="1701381345164" updateUser="ExternalLinkService">
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<mods:title id="F4E496FA255DAE79B285584F2E411491">New data on the taxonomy, morphology and distribution of Naardaineffectalis (Walker, 1859) (Lepidoptera, Erebidae, Hypeninae)</mods:title>
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<mods:namePart id="F646CCF4F5DB50C1284890C510C08052">Toth, Balazs</mods:namePart>
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<mods:title id="AA33156BFCD61F1366ECD8031A63FBC3">Nota Lepidopterologica</mods:title>
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<mods:date id="75891611F75317402B258ADF0AC661F8">2018</mods:date>
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<treatment id="E7263586C77072F295F499021FD1FFBA" ID-GBIF-Taxon="141124769" LSID="urn:lsid:plazi:treatment:E7263586C77072F295F499021FD1FFBA" httpUri="http://treatment.plazi.org/id/E7263586C77072F295F499021FD1FFBA" lastPageId="4" lastPageNumber="103" pageId="1" pageNumber="100">
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<subSubSection id="1F069611F352C9D44C7EE0943986CCBA" pageId="1" pageNumber="100" type="nomenclature">
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<paragraph id="C385F61F159ECDD8980EF782BC6D99AD" pageId="1" pageNumber="100">
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<taxonomicName id="D89141B759CB518F2FC843642D4DC5AB" ID-CoL="45JBN" authority="Walker, 1859" authorityName="Walker" authorityYear="1859" class="Insecta" family="Erebidae" genus="Naarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naarda ineffectalis" order="Lepidoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="ineffectalis">Naarda ineffectalis (Walker, 1859)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="F939C01FEA954C761B905D41D25068F9" pageId="1" pageNumber="100" type="reference_group">
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<paragraph id="61B304C07DDC6203F79DECC7A6BC4E1A" pageId="1" pageNumber="100">
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<taxonomicName id="2BDA50E318B8CCD84D94DAA54C3EDEEE" class="Insecta" family="Erebidae" genus="Hypena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypena ineffectalis" order="Lepidoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="ineffectalis">Hypena ineffectalis</taxonomicName>
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Walker, 1859 - List of specimens of lepidopterous insects in the Collection of the British Museum 16: 85. Type locality: Ceylon [= Sri Lanka]. Holotype, coll. NHMUK.
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</paragraph>
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<paragraph id="0AE73C226FB06BDE737D7AF6D661C6DE" pageId="1" pageNumber="100">
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<taxonomicName id="F4A76C9A406FB656C69F79A10C4C0B3E" class="Insecta" family="Erebidae" genus="Ptyophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptyophora ochreistigma" order="Lepidoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="ochreistigma">Ptyophora ochreistigma</taxonomicName>
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Hampson, 1893 - Illustrations of typical specimens of
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<taxonomicName id="6327437BD96387A1E4A66D47ADD33D83" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="1" pageNumber="100" rank="order">Lepidoptera</taxonomicName>
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<taxonomicName id="92CDE2EF519746068A8187A1180FCF25" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Heterocera" order="Lepidoptera" pageId="1" pageNumber="100" rank="suborder" suborder="Heterocera">Heterocera</taxonomicName>
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in the Collection of the British Museum 2: 124. Type locality: Nawalapitya, Ceylon [= Sri Lanka]. Holotype, coll. NHMUK.
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</paragraph>
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</subSubSection>
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<subSubSection id="EE1B96A28A2A29A267BFCCD5ACEAFCC1" pageId="1" pageNumber="100" type="type material">
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<paragraph id="90ADB6C69E9C02165AF46F0579890C27" pageId="1" pageNumber="100">Type material examined.</paragraph>
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<paragraph id="CD1A2E44E1E70D878295865AD9C3C0E7" pageId="1" pageNumber="100">Holotype (Figs 1, 13). ♀ (see below): Ceylon; Abdomen missing (coll. NHMUK).</paragraph>
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<caption id="691285A7186522146200E48051C3F3BB" pageId="1" pageNumber="100">
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<paragraph id="2C0114BD1AD66828FC4749971FE2E549" pageId="1" pageNumber="100">
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Figures 1-16. 1-12. Adults. 1.
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<taxonomicName id="7BFC54633207C6B1723780EE499FF26A" class="Insecta" family="Erebidae" genus="Naarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naarda ineffectalis" order="Lepidoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="ineffectalis">Naarda ineffectalis</taxonomicName>
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(Walker, 1859) female, holotype (coll. NHMUK). 2. ditto, female, Sri Lanka; slide No. BM 21618♀ (coll. NHMUK). 3. ditto, female, Thailand; slide No. OP1624f (coll. OP). 4. ditto, male, Korea; slide No. TB568m (coll. MU). 5. ditto, female, Japan; slide No. TB1583f (coll. NHMUK). 6. ditto, male, Cambodia; slide No. TB832m (coll. HNHM). 7. ditto, female, Cambodia; slide No. TB827f (coll. HNHM). 8. ditto, male, Borneo, Sarawak (coll. ZMUC). 9. ditto, female, Borneo, Sarawak (coll. ZMUC). 10. ditto, male, Sumatra; slide No. TB421m (coll. MF in ZMUC). 11.
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<taxonomicName id="E7ED987FE00BD7AB5ADD9A9519959206" lsidName="N. notata" pageId="1" pageNumber="100" rank="species" species="notata">N. notata</taxonomicName>
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(Hampson, 1891), male, Sri Lanka; slide No. BM 20090♂ (coll. NHMUK). 12. ditto, female, syntype; slide No. BM 21613 (coll. NHMUK). 13-16. Labial palps. 13.
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<taxonomicName id="89BE8B1652AD4EEB27493C3705B505BF" lsidName="N. ineffectalis" pageId="1" pageNumber="100" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
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female, holotype (coll. NHMUK). 14. ditto, female, Sumatra (coll. MF in ZMUC). 15. ditto, male, Sumatra (coll. MF in ZMUC). 16.
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<taxonomicName id="89FA138C6E49AD8D8BA7C09972EA1D89" lsidName="N. notata" pageId="1" pageNumber="100" rank="species" species="notata">N. notata</taxonomicName>
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female, syntype, Sri Lanka; slide No. BM 21613♀ (coll. NHMUK). Figures are to scale.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="D3DC0E497DE5BDEE3A53A8B11027ED81" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100" type="materials_examined">
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<paragraph id="3D49FBADFA9B3B6F37DE1AC1C0430A8D" pageId="1" pageNumber="100">Additional material examined.</paragraph>
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<paragraph id="18117CAA56A9A0D07B2363887452A0EA" pageId="1" pageNumber="100">
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Sri Lanka. 1 ♀: Colombo, viii.1908. (coll. NHMUK), 1 ♀: Colombo, 22.ii.1902. (coll. NHMUK), 1 ♀: Panquelba, i.1904. (coll. NHMUK), 1 ♀: Haputale, vi.1899 (coll. NHMUK), 1 ♀: Gampola, viii.1907; Abdomen missing (coll. NHMUK), 1 ♂: Nawalapitya; Ceylon; Coll. Green; 91.-26., slide No. BM Noct. 2812 (holotype of
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<taxonomicName id="CC360B539A5A8F12D4FEFCF316372163" class="Insecta" family="Erebidae" genus="Ptyophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptyophora ochreistigma" order="Lepidoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="ochreistigma">Ptyophora ochreistigma</taxonomicName>
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) (coll. NHMUK), 1 ♀ (Figs 2, 14): Haldumulla; xii.1909; ex. Coll. G.C. Alston; Joicey Bequest. Brit. Mus. 1934-120; slide No. RL7689♀ = BM Noct. 21618♀ (coll. NHMUK), 1 ♀: Coll. Green; 29-26; Abdomen missing (coll. NHMUK), 4 ♀: Maskeliya; January; ex. Coll. G.C. Alston.; Joicey Bequest. Brit. Mus. 1934-120, slide No. BM Noct. 2944 (coll. NHMUK), 1 ♀: data as previous specimen, but October (coll. NHMUK), 1 ♀: data as previous specimen, but March (coll. NHMUK), 1 ♀: data as previous specimen, but April (coll. NHMUK), 1 ♀: Negombo; 1973.iii.23.; leg.
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<normalizedToken id="A7E06C6CA27F391DDBA199268F3B5AAF" originalValue="Hüttler">Huettler</normalizedToken>
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B.; slide No. RL10784f (coll. HNHM).
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</paragraph>
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<paragraph id="AF2D6877C2A17FC6FD3476B3B9341448" pageId="2" pageNumber="101">
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<pageBreakToken id="FA43A89B4BE287F452E57BCAA2A16D7D" pageId="2" pageNumber="101" start="start">Thailand</pageBreakToken>
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. 1 ♀ (Fig. 3): N. Thailand, Mae Hong Son prov., Pai,
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<geoCoordinate id="ADDE87A86B84365F8105442F7C52296F" direction="north" orientation="latitude" precision="9" value="19.358334">19°21.50'N</geoCoordinate>
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,
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<geoCoordinate id="8E0E43A08233D94DFD8F40D2BD357C60" direction="east" orientation="longitude" precision="9" value="98.44683">98°26.81'E</geoCoordinate>
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, ~500 m a.s.l., at light, 15-24.xii.2010., leg. K. Tomkovich; slide No. OP1624f (coll. OP).
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</paragraph>
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<paragraph id="D7E741EA184A848E274E4C7C9E9C37A2" pageId="2" pageNumber="101">
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Cambodia. 2 ♂, 1 ♀: Mondolkiri prov.; Seima Biodiversity Conservation Area, road
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<normalizedToken id="7BFC8665EECAF02A861149B12F7080EC" originalValue="Seima–O’Rang">Seima-O'Rang</normalizedToken>
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;
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<geoCoordinate id="F87D45F5BD3597AC5C5A25B8FC9691FB" direction="north" orientation="latitude" precision="9" value="12.200334">12°12.02'N</geoCoordinate>
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,
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<geoCoordinate id="A295D3D17725FB278B0B1B189A296808" direction="east" orientation="longitude" precision="9" value="107.033">107°01.98'E</geoCoordinate>
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, 300 m; No. 117, 30.i.2006, at light; leg. G. Csorba & G. Ronkay; slide Nos TB793m, TB794f, TB813m (coll. HNHM), 2 ♂, 5 ♀: Mondolkiri prov.; Seima Biodiversity Conservation Area, road
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<normalizedToken id="B117827E6AE3CDB309420B4FA21A74AC" originalValue="Seima–O’Rang">Seima-O'Rang</normalizedToken>
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;
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<geoCoordinate id="65E1F5B6BAFCAD826D1D1A40D9B637BA" direction="north" orientation="latitude" precision="9" value="12.262167">12°15.73'N</geoCoordinate>
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,
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<geoCoordinate id="7ABA26A1D5C634768C2B302F62854A1D" direction="east" orientation="longitude" precision="9" value="107.06367">107°03.82'E</geoCoordinate>
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, 360 m; No. 88, 27-29.i.2006, at light; leg. G. Csorba & G. Ronkay; slide Nos TB827f (Fig. 7), TB830f, TB831f, TB832m (Fig. 6) (coll. HNHM).
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</paragraph>
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<paragraph id="7514F479F310134E12C1BC756B109484" pageId="2" pageNumber="101">
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Japan. 1 ♀ (Fig. 5): Mifune; prov. Higo; Kyushu; 20.x.1855; A.E. Wileman; 1378;
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<normalizedToken id="6006B580DB787ECDB4B88613ADDBF132" originalValue="“Gnaephila”">"Gnaephila"</normalizedToken>
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[sic, hand-written]; Wileman Coll. B.M. 1929-261; NHMUK 010914474; slide No. TB1583f (coll. NHMUK).
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</paragraph>
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<paragraph id="60B6868CA978FD4307EC589309A2DC61" pageId="2" pageNumber="101">
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Korea. 1 ♂ (Fig. 4): JN: Muan, Sangma-ri; 8.vi.2006. Sei-Woong Choi;
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<geoCoordinate id="F070665EFA139EF2B244E3D8C51A066A" direction="north" orientation="latitude" precision="925" value="34.916668">34°55'N</geoCoordinate>
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,
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<geoCoordinate id="CC01BFFEEE3944C2F73EF10A20367B3C" direction="east" orientation="longitude" precision="925" value="126.416664">126°25'E</geoCoordinate>
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, 55 m; slide No. TB568m (coll. MU).
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</paragraph>
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<paragraph id="6D5B2C39BE935FC30AD4AFC3BAB67229" pageId="2" pageNumber="101">
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Sumatra. 3 ♂: North Sumatra, Pematang Siantar, 460 m. 10-14.ii.2002. leg. M. Fibiger & K. Larsen; slide Nos TB412m, TB413m, TB421m (Figs 10, 15) (coll. MF, currently in ZMUC), 1 ♂: North-Sumatra, Hutu Padang (Asahan), River Silau, 48 km SE P. Siantar, Near Sialangoman, 220 m, 13.ii.2002. leg. M. Fibiger & K. Larsen; slide No. TB416m (coll. MF, currently in ZMUC), 1 ♀: North-Sumatra; H.W. II., 28 km S. P. Siantar; Near Tigadoluk, 1050 m;
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<geoCoordinate id="764A956BFE4BF9970A08C8F83E0FE3B0" direction="north" orientation="latitude" precision="9" value="2.7645">2°45.87'N</geoCoordinate>
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,
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<geoCoordinate id="62C44127AAE25470BF0E8966C2DDE95D" direction="east" orientation="longitude" precision="9" value="99.97217">99°58.33'E</geoCoordinate>
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; 4.iii.2002., leg. K. Larsen & M. Fibiger; slide No. TB649f (coll. MF, currently in ZMUC), 1 ♀: West-Sumatra, Bukittinggi, 980 m,
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<geoCoordinate id="17BDDF2CCDA1EC742C915B7E459BA43C" direction="south" orientation="latitude" precision="9" value="-0.25833333">00°15.50'S</geoCoordinate>
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,
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<geoCoordinate id="01204DED2C7EAC1EDE0E6B466C1B9970" direction="east" orientation="longitude" precision="9" value="100.35367">100°21.22'E</geoCoordinate>
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, 19-23.ii.2002., leg. M. Fibiger & K. Larsen; slide No. TB414f (coll. HNHM), 1 ♀: West-Sumatra, Batang Palupuh, 880 m, 7 km N. Bukittinggi,
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<geoCoordinate id="B6B4B882B4D49BDBE7A8E8B5634F39EC" direction="south" orientation="latitude" precision="9" value="-0.24283333">00°14.57'S</geoCoordinate>
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,
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<geoCoordinate id="9EEA3AF2D730C506E28EC3794CE5A207" direction="east" orientation="longitude" precision="9" value="100.15366">100°09.22'E</geoCoordinate>
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, 21.ii.2002. leg. M. Fibiger & K. Larsen; slide No. TB681f (coll. MF, currently in ZMUC), 1 ♀: West-Sumatra, Penjabungan Tonga,
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<geoCoordinate id="172A71A2BCEF0E4A1B6E12527B1A9AEB" direction="north" orientation="latitude" precision="9" value="0.8105">00°48.63'N</geoCoordinate>
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,
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<geoCoordinate id="8AA104ED509332647BE9571D93D81EF0" direction="east" orientation="longitude" precision="9" value="99.568665">99°34.12'E</geoCoordinate>
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, 200 m, 18.ii.2002. leg. M. Fibiger & K. Larsen; slide No. TB418f (coll. MF, currently in ZMUC).
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</paragraph>
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<paragraph id="1175B246E0B6B55632079ED70705DF95" pageId="2" pageNumber="101">Borneo. 1 ♂, 1 ♀: Sarawak, Semongok; 12 mi. S Kuching; 26.iv.1974; at light, A. Earnshaw; slide Nos TB425m, TB544f (coll. ZMUC), 1 ♂ (Fig. 8): data as previous specimens, but 12.viii.1974 (coll. ZMUC), 2 ♂: data as previous specimen, but 24.x.1974; slide Nos TB539m, TB545m (coll. HNHM), 3 ♀: data as previous specimens, but 25.x.1974; slide Nos TB538f, TB546f (coll. ZMUC), 1 ♂: data as previous specimens, but 1.xii.1974; slide No. TB543m (coll. ZMUC), 1 ♀: data as previous specimens, but 6.xii.1974 (coll. ZMUC), 2 ♀ (Fig. 9): data as previous specimen, but 24.xii.1974; slide No. TB683f (coll. ZMUC), 1 ♀: data as previous specimens, but 1.i.1975 (coll. ZMUC).</paragraph>
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</subSubSection>
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<subSubSection id="A39AAFA5CB2CEDF06017B771BE778781" pageId="2" pageNumber="101" type="taxonomic comments">
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<paragraph id="DA2DB84A9E143D52BC90A04587897407" pageId="2" pageNumber="101">Taxonomic comments.</paragraph>
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<paragraph id="3C3C51F2B620E2F6B7AAB0A5F0526331" pageId="2" pageNumber="101">
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<bibRefCitation id="E9296F34DC8AF307C6CDEDE24CCB04F3" author="Holloway, JD" journalOrPublisher="Malayan Nature Journal" pageId="5" pageNumber="104" pagination="1 - 268" title="The moths of Borneo: family Noctuidae, subfamilies Rivulinae, Phytometrinae, Herminiinae, Hypeninae, Hypenodinae." volume="60" year="2008">Holloway (2008)</bibRefCitation>
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synonymised
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<taxonomicName id="46A78A7731156B0EA23163C2488E726D" lsidName="N. ochreistigma" pageId="2" pageNumber="101" rank="species" species="ochreistigma">N. ochreistigma</taxonomicName>
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(Hampson, 1893) and
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<taxonomicName id="D20794746EA96805F8420E4B2AB23B08" lsidName="N. ineffectalis" pageId="2" pageNumber="101" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
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on the basis of a topotypic male specimen from the latter taxon having the same genitalia as those of the
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<taxonomicName id="055862B1FAB1742EDF526A2A0A5B78EA" lsidName="N. ochreistigma" pageId="2" pageNumber="101" rank="species" species="ochreistigma">N. ochreistigma</taxonomicName>
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holotype.
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</paragraph>
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<paragraph id="79BA138A4C29280CA0CA6DA367EFBDBF" pageId="2" pageNumber="101">
|
||
Examination of the head structure of the
|
||
<taxonomicName id="D79AA242ACB13832D2E30F8FD5A4101D" lsidName="N. ineffectalis" pageId="2" pageNumber="101" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
holotype specimen (Figs 1, 13) led me to the striking conclusion that it was, in contrast to
|
||
<bibRefCitation id="A96913976338B4FECE4B4F89B1954124" author="Hampson, GF" journalOrPublisher="Taylor & Francis, London" pageId="5" pageNumber="104" title="Illustrations of typical specimens of LepidopteraHeterocera in the Collection of the British Museum IX: The Macrolepidoptera Heterocera of Ceylon." year="1893">Hampson (1893)</bibRefCitation>
|
||
, actually a female. Additionally, the characters of the antenna and labial palps match well with those of
|
||
<taxonomicName id="FC6A884379F10130817E6174A1B73753" lsidName="N. ochreistigma" pageId="2" pageNumber="101" rank="species" species="ochreistigma">N. ochreistigma</taxonomicName>
|
||
females. These results support
|
||
<normalizedToken id="8B3F4E387FBBBBC14D9F3198EBA88034" originalValue="Holloway’s">Holloway's</normalizedToken>
|
||
synonymisation. Unfortunately the genitalia of the holotype specimen of
|
||
<taxonomicName id="F087AF37B28FFEC8249711FC4AA0543F" lsidName="N. ineffectalis" pageId="2" pageNumber="101" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
cannot be studied because its abdomen is missing.
|
||
</paragraph>
|
||
<paragraph id="10CA55460B9893D610C94D645E838CCF" pageId="2" pageNumber="101">
|
||
<taxonomicName id="7E97579D33DB9450545ABA545A6D3317" class="Insecta" family="Erebidae" genus="Naarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naarda ineffectalis" order="Lepidoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="ineffectalis">Naarda ineffectalis</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="63D90B46E192E17988C95C2C464175C1" lsidName="N. notata" pageId="2" pageNumber="101" rank="species" species="notata">N. notata</taxonomicName>
|
||
in its shape of labial palps, overall size, ground colour and pattern of wings as well as in the genitalia of both sexes (Figs 1-20). These differences are detailed below, in the Diagnosis. I consider
|
||
<taxonomicName id="B74BAC08E68C914B8860CDAF8C3B3FB4" class="Insecta" family="Erebidae" genus="Naarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naarda notata" order="Lepidoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="notata">Naarda notata</taxonomicName>
|
||
(Hampson, 1891), corroborating
|
||
<bibRefCitation id="DDA04D894AC0AD6044F84CC7761D6853" author="Holloway, JD" journalOrPublisher="Malayan Nature Journal" pageId="5" pageNumber="104" pagination="1 - 268" title="The moths of Borneo: family Noctuidae, subfamilies Rivulinae, Phytometrinae, Herminiinae, Hypeninae, Hypenodinae." volume="60" year="2008">Holloway (2008)</bibRefCitation>
|
||
, as a distinct species; stat. rev.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6E93106C69088B11F984B42EB9F5DAB8" lastPageId="3" lastPageNumber="102" pageId="2" pageNumber="101" type="description">
|
||
<paragraph id="699D9AEABAE4C7C6579CF6939F94B022" pageId="2" pageNumber="101">Description of the female.</paragraph>
|
||
<paragraph id="BC539388C74CDF22A2224D307ADE0B88" lastPageId="3" lastPageNumber="102" pageId="2" pageNumber="101">
|
||
Wingspan 12-15 mm, length of forewing 6-8 mm. Antenna typical of females in the genus: filiform and sparsely ciliate, with two setae on each segment as long as the diameter of the flagellum, cilia half as long as diameter of the flagellum. Labial palps four times
|
||
<pageBreakToken id="C48AC7C02FA16D8E29BBC97BE883E95B" pageId="3" pageNumber="102" start="start">longer</pageBreakToken>
|
||
than diameter of eye (longer than in male, see Figs 13-15); 3rd segment conspicuous, 2nd segment broad, domed. Scale-hood of vertex broad-based and long. Forewing brownish grey in colour; subterminal and postmedial lines parallel, slightly sinuous, the subterminal easily fading during ageing, the latter relatively broad; reniform stigma quite small, angular, elongated, pale yellow with two black dots inside, lower dot is larger than the upper one; orbicular rounded, as yellow as reniform. Hindwing slightly paler than forewing except for the area between vein Cu2 and dorsum being as dark as forewing; transverse lines like those of forewing but postmedial line often more conspicuous. Pattern of wings slightly less conspicuous in females than in males.
|
||
</paragraph>
|
||
<paragraph id="8E5CD3F29E8034958FF66BFBB20D4FE9" pageId="3" pageNumber="102">Female genitalia (Fig. 19). Corpus bursae ovoid, its anterior half densely covered by tiny grains, posterior half slightly wrinkled in longitudinal direction. Near the mouth of ductus bursae a broad and long, helical cervix arises. Ductus bursae smooth, broad, relatively short, nearly entirely covered by the triangular colliculum. Sternum A7 very narrow, sinus absent.</paragraph>
|
||
<paragraph id="DEB6D56B4752279F637AE5CA1736B0B5" pageId="3" pageNumber="102">Apophyses anteriores as long as apophyses posteriores; ovipositor lobes angular.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="335EF794AFA2620F5E30F5B5557D7BA4" lastPageId="4" lastPageNumber="103" pageId="3" pageNumber="102" type="diagnosis">
|
||
<paragraph id="218AED4DBB3A27CECE7B920487092517" pageId="3" pageNumber="102">Diagnosis.</paragraph>
|
||
<paragraph id="BBF6A75E3400512FF4714FE331C61FE4" lastPageId="4" lastPageNumber="103" pageId="3" pageNumber="102">
|
||
<taxonomicName id="E979D914469AC3415C12B96F1FC78C39" class="Insecta" family="Erebidae" genus="Naarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naarda ineffectalis" order="Lepidoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="ineffectalis">Naarda ineffectalis</taxonomicName>
|
||
externally resembles several sympatric congeners, including
|
||
<taxonomicName id="C3D1B3CFE3BC9C399F1C0895E84F6951" lsidName="N. huettleri" pageId="3" pageNumber="102" rank="species" species="huettleri">N. huettleri</taxonomicName>
|
||
<normalizedToken id="C415CF2DDAE63EBA304E07338DA9ED4E" originalValue="Tóth">Toth</normalizedToken>
|
||
& Ronkay, 2015 and
|
||
<taxonomicName id="D6F1B2033439648432ECC76D4D112EE1" lsidName="N. imitata" pageId="3" pageNumber="102" rank="species" species="imitata">N. imitata</taxonomicName>
|
||
<normalizedToken id="1B805F919D9FECB3DE8C1EE561A87DCF" originalValue="Tóth">Toth</normalizedToken>
|
||
& Ronkay, 2015 and tends to become worn, so that dissection of genitalia is usually needed for proper identification. Characteristic features of the male genitalia (Fig. 17) are as follows: uncus with double tips, juxta large and scobinate, saccus long, valva short, triangular, aedeagus stout, cornutus very large, vesica longitudinally wrinkled
|
||
<pageBreakToken id="62F6747584BE6FFFEFBCF3A119B42905" pageId="4" pageNumber="103" start="start">between</pageBreakToken>
|
||
cornutus and carina. The female genitalia of
|
||
<taxonomicName id="4DBC569CF0CF703ADD9E53324230385B" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
can be distinguished from those of all other
|
||
<taxonomicName id="B17DF7BDF69532652FA65CAB9B359105" class="Insecta" family="Erebidae" genus="Naarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naarda" order="Lepidoptera" pageId="4" pageNumber="103" phylum="Arthropoda" rank="genus">Naarda</taxonomicName>
|
||
species by presence of the large, helical cervix.
|
||
</paragraph>
|
||
<caption id="BBC4E54A8AF30EE2C05CCC10F10F1CA1" pageId="4" pageNumber="103">
|
||
<paragraph id="FC28873896271861CB71251363B40A01" pageId="4" pageNumber="103">
|
||
Figures 17-20. 17-18. Male genitalia. 17.
|
||
<taxonomicName id="AC4F2A7DAFF6F466797A987BE7720A2C" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
, aedeagus below: TB412m, clasping apparatus: TB416m (coll. MF in ZMUC). 18.
|
||
<taxonomicName id="A8F3B32C7EFB19AB0C68B6EB3361306C" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
, aedeagus below; slide No. BM 20090♂ (coll. NHMUK). 19-20. Female genitalia. 19.
|
||
<taxonomicName id="BFFDA2040570817889B8B88CD623153E" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
; slide No. TB414f (coll. HNHM). 20.
|
||
<taxonomicName id="405A8B01435862C22ABEFF888937D7B0" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
, syntype; slide No. BM 21613♀ (coll. NHMUK). Scale bar: 1 mm. Figures are to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="85A3DD50E7BECB0E26ADDC806DFFD026" pageId="4" pageNumber="103">
|
||
Compared to
|
||
<taxonomicName id="1EEB78D01262FED90FD5D867DDADAA7C" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
the former species has shorter and dorsally more convex labial palps; the wingspan of
|
||
<taxonomicName id="25521167DE5CC06CC66310B8FB29BD96" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
is only two-thirds of
|
||
<taxonomicName id="4D3CDEBC218A4BE4FE5D23AE08988171" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
; the ground colour of the wings is darker, greyer, especially in the basal field; on the forewing the reniform stigma is narrower and lighter, and the transverse lines are less prominent than those of
|
||
<taxonomicName id="B092C38F719CECAA04E61A4A37D1DC63" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
and
|
||
<taxonomicName id="3906FD3CA176BB3309819D5D4DB40B1A" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
lacks the blackish patch laterally from the reniform stigma, which is present in the other species. The hindwing of
|
||
<taxonomicName id="80C0ABC5AA6214032D7768C02084C034" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
is only slightly lighter than the forewing while in
|
||
<taxonomicName id="04DCD855A160717DBDA7099902DBBB24" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
the hindwing is much lighter than the forewing, and the ground colour becomes white towards the costa. In the male genitalia, the uncus of
|
||
<taxonomicName id="2CDFB5496B9F136B79B902922E46A151" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
is slightly bifurcate, but that of
|
||
<taxonomicName id="E9FF12C525141F540BB0B126808A2481" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
is simple; the juxta is larger, the valva is much broader and shorter and the cornutus of aedeagus is much larger in
|
||
<taxonomicName id="F15C42C56C3F249312E78EE71E88C186" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
than in
|
||
<taxonomicName id="D414E7B106072F2B656DA9CACFC0A22B" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
. The corpus bursae in the female genitalia of
|
||
<taxonomicName id="7843687C8CC5D46F6AC76E7C25876D6C" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
is bigger in proportion to the whole genitalia, the cervix is much longer and the ductus bursae is shorter than those of
|
||
<taxonomicName id="B5DD05421D6DE45BB3161FB3C0CD10EB" lsidName="N. notata" pageId="4" pageNumber="103" rank="species" species="notata">N. notata</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1A77C76A35DCAABF2DA1958A499DCEAE" pageId="4" pageNumber="103" type="distribution">
|
||
<paragraph id="7F74B36ECAE9D135C0D9762EA23F0529" pageId="4" pageNumber="103">Distribution.</paragraph>
|
||
<paragraph id="256EBE07E60FDFF984A55C248C6FB057" pageId="4" pageNumber="103">
|
||
From Sri Lanka to Sumatra and Borneo (Sarawak and Brunei:
|
||
<bibRefCitation id="16EBC850374DEB28A1C11FDEEB2B1A1F" author="Holloway, JD" journalOrPublisher="Malayan Nature Journal" pageId="5" pageNumber="104" pagination="1 - 268" title="The moths of Borneo: family Noctuidae, subfamilies Rivulinae, Phytometrinae, Herminiinae, Hypeninae, Hypenodinae." volume="60" year="2008">Holloway 2008</bibRefCitation>
|
||
). On the Asian mainland in Cambodia and Thailand, with isolated, unusually northern, records from Japan and Korea (Fig. 21). As both latter specimens were collected near large ports, accidental introduction by transport cannot be excluded.
|
||
</paragraph>
|
||
<caption id="920D8D40B0B2C1A47802E61419BEE5E8" pageId="4" pageNumber="103">
|
||
<paragraph id="945583AB62A3314AAABE183F70CA2069" pageId="4" pageNumber="103">
|
||
Figure 21. Distribution of
|
||
<taxonomicName id="19E5896E3FEE028F4D35A6AD67980F4E" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
. Previous records marked with dots, new records with diamonds.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5A54590A4E9CD9CACAD6637D16518F7E" pageId="4" pageNumber="103">New for the faunas of Cambodia, Thailand, Japan, Korea and Sumatra (thus for entire Asian mainland and also for the Palaearctic Region).</paragraph>
|
||
<subSection id="0A804DB53A3EF176B1B588818B082E3A" pageId="4" pageNumber="103" type="key to the naarda species of korea and main islands of japan">
|
||
<paragraph id="80CA74C50EDBCCFF3851EC7656610EEB" pageId="4" pageNumber="103">
|
||
Key to the
|
||
<taxonomicName id="54BCFE6CE3BD93C3A4B1CEC660124DED" class="Insecta" family="Erebidae" genus="Naarda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Naarda" order="Lepidoptera" pageId="4" pageNumber="103" phylum="Arthropoda" rank="genus">Naarda</taxonomicName>
|
||
species of Korea and main islands of Japan
|
||
</paragraph>
|
||
<paragraph id="A3061FD9EED514B7BF4454CD7534E110" pageId="4" pageNumber="103">
|
||
<table id="3E6D2895924C227234869C5D8B8F57E4" pageId="4" pageNumber="103">
|
||
<tr id="F0211A3BEF8270BFBC6240540806808E" pageId="4" pageNumber="103">
|
||
<td id="F1A25A7733F52D99B5A8E1E6EDC3E30E" colspan="1" pageId="4" pageNumber="103" rowspan="1">
|
||
<taxonomicName id="8735A9F56A2230D3F4827523DD444170" lsidName="N. maculifera" pageId="4" pageNumber="103" rank="species" species="maculifera">N. maculifera</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="05240608DE8802EA2D97B9854FCBCAFD" pageId="4" pageNumber="103">
|
||
<td id="225480A02CCA46B9ACAEF065EE8D82E4" colspan="1" pageId="4" pageNumber="103" rowspan="1">
|
||
<taxonomicName id="84EB14F49FC52231FE05392FA4DA7761" lsidName="N. hallasana" pageId="4" pageNumber="103" rank="species" species="hallasana">N. hallasana</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="37714E82FC2A998D7431E1784CB76A6F" pageId="4" pageNumber="103">
|
||
<td id="E3A523F9DF126019447E35432261FF85" colspan="1" pageId="4" pageNumber="103" rowspan="1">
|
||
<taxonomicName id="8F948DB011C7FA232A77F9C1A13C3168" lsidName="N. ineffectalis" pageId="4" pageNumber="103" rank="species" species="ineffectalis">N. ineffectalis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |