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<document id="B2A6D8C8DF8BC9C8C46F898DBAFD8420" ID-CLB-Dataset="3746" ID-DOI="10.11646/zootaxa.5060.3.6" ID-GBIF-Dataset="bdcf9ea4-269b-4084-ba96-c8f391588323" ID-ISSN="1175-5326" ID-Zenodo-Dep="5635912" ID-ZooBank="D87E8F4A-DDBC-4F72-AE15-8E3719664581" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1635758555739" checkinUser="plazi" docAuthor="Shah, Bismillah, Naveed, Hassan, Webb, Michael D. &amp; Duan, Yani" docDate="2021" docId="E36C771FFFE1FFAAFF7DFB340A300783" docLanguage="en" docName="zootaxa.5050.3.6.pdf" docOrigin="Zootaxa 5060 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maiestas tareni" docType="treatment" docVersion="6" lastPageNumber="408" masterDocId="1F550F67FFE6FFADFFEAFFE90B210163" masterDocTitle="Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records" masterLastPageNumber="416" masterPageNumber="401" pageNumber="408" updateTime="1699467297445" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2CEF37B4E337DE6C1CF4954A53E0BAFD">Taxonomic review of the grassland leafhopper genus Maiestas Distant (Hemiptera: Cicadellidae: Deltocephalinae: Deltocephalini) from Pakistan with description of a new species and two new records</mods:title>
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<mods:namePart id="6F6D96E76A536250DD8D91850E386EA8">Shah, Bismillah</mods:namePart>
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<mods:affiliation id="23A5769876492C1860A2DA75A534E43B">Anhui Province Key Laboratory of Integrated Pest Management on Crops, Key Laboratory of Biology and Sustainable Management of Plant Diseases and Pests of Anhui Higher Education Institutes, School of Plant Protection, Anhui Agricultural University, Hefei, Anhui 230036, China. &amp; bismillahshah 1990 @ yahoo. com; https: // orcid. org / 0000 - 0002 - 8407 - 8627</mods:affiliation>
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<mods:namePart id="1985AE114EB104B614968DECC795DDD8">Naveed, Hassan</mods:namePart>
<mods:affiliation id="D1DE6B332226DFAA6D09D8689C6950F4">College of Life Sciences, Leshan Normal University, Leshan, Sichuan 614004, China.</mods:affiliation>
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<mods:namePart id="3E126A3DFB7ED4A911E54A1633B69373">Webb, Michael D.</mods:namePart>
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<mods:affiliation id="4BEA0666F9B21C0A83B2F3025FB1E3F5">Department of Life Sciences, Insects, the Natural History Museum, South Kensington, London, SW 7 5 BD, UK m. webb @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
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<mods:namePart id="0973D2235ED8E9400DBCF99F58021A21">Duan, Yani</mods:namePart>
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<mods:affiliation id="2C363799FE096C87F1A82D39417DFFD3">Anhui Province Key Laboratory of Integrated Pest Management on Crops, Key Laboratory of Biology and Sustainable Management of Plant Diseases and Pests of Anhui Higher Education Institutes, School of Plant Protection, Anhui Agricultural University, Hefei, Anhui 230036, China. &amp; duanyani @ hotmail. com; https: // orcid. org / 0000 - 0002 - 5952 - 0778</mods:affiliation>
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<treatment id="E36C771FFFE1FFAAFF7DFB340A300783" ID-DOI="http://doi.org/10.5281/zenodo.5635944" ID-GBIF-Taxon="189798222" ID-Zenodo-Dep="5635944" LSID="urn:lsid:plazi:treatment:E36C771FFFE1FFAAFF7DFB340A300783" httpUri="http://treatment.plazi.org/id/E36C771FFFE1FFAAFF7DFB340A300783" lastPageNumber="408" pageId="7" pageNumber="408">
<subSubSection id="23DF9582FFE1FFAAFF7DFB3409540594" box="[151,629,1245,1271]" pageId="7" pageNumber="408" type="nomenclature">
<paragraph id="6B7AC609FFE1FFAAFF7DFB3409540594" blockId="7.[151,629,1245,1308]" box="[151,629,1245,1271]" pageId="7" pageNumber="408">
<heading id="30327165FFE1FFAAFF7DFB3409540594" bold="true" box="[151,629,1245,1271]" fontSize="11" level="1" pageId="7" pageNumber="408" reason="1">
<taxonomicName id="ACC5BD8AFFE1FFAAFF7DFB3409540594" ID-CoL="8RLWH" authority="(Dash &amp; Viraktamath)" baseAuthorityName="Dash &amp; Viraktamath" baseAuthorityYear="1998" box="[151,629,1245,1271]" class="Insecta" family="Cicadellidae" genus="Maiestas" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tareni">
<emphasis id="59B11A1BFFE1FFAAFF7DFB3409540594" bold="true" box="[151,629,1245,1271]" pageId="7" pageNumber="408">
<emphasis id="59B11A1BFFE1FFAAFF7DFB340A730594" bold="true" box="[151,338,1245,1271]" italics="true" pageId="7" pageNumber="408">Maiestas tareni</emphasis>
(Dash &amp; Viraktamath)
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="23DF9582FFE1FFAAFF7DFAE80BCA047F" box="[151,235,1281,1308]" pageId="7" pageNumber="408" type="description">
<paragraph id="6B7AC609FFE1FFAAFF7DFAE80BCA047F" blockId="7.[151,629,1245,1308]" box="[151,235,1281,1308]" pageId="7" pageNumber="408">
(
<figureCitation id="F3FEDA8CFFE1FFAAFF4AFAE80BC3047F" box="[160,226,1281,1308]" captionStart="FIGURE 5" captionStartId="9.[151,250,1749,1773]" captionTargetBox="[217,1365,194,1721]" captionTargetId="figure-19@9.[205,1383,181,1725]" captionTargetPageId="9" captionText="FIGURE 5. Maiestas tareni (Dash &amp; Viraktamath). AH: this study, A: habitus, dorsal view; B: habitus, lateral view; C: head and thorax, dorsal view; D: face; E: male genital capsule, lateral view; F: subgenital plate, valve and styles, dorsal view; G &amp; H: connective and aedeagus, dorsal and lateral view; IN: after Zhang &amp; Duan, 2011, I: habitus, dorsal view; J: male pygofer lobe, lateral view; K: subgenital plate, ventral view; L: style, dorsal view; M &amp; N: connective and aedeagus, dorsal and lateral view; OT: after Khatri &amp; Webb, 2010, O: habitus, dorsal view; P: head and thorax, dorsal view; Q: subgenital plate, ventral view; R: style, dorsal view; S &amp; T: connective and aedeagus, dorsal and lateral view; UZ: after Dash &amp; Viraktamath, 1995, U: head and thorax, dorsal view; V: face; W: subgenital plates and valve, dorsal view; X: style, dorsal view; Y &amp; Z: connective and aedeagus, dorsal and lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.5635926" httpUri="https://zenodo.org/record/5635926/files/figure.png" pageId="7" pageNumber="408">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="23DF9582FFE1FFAAFF7DFAAF0A9504F8" pageId="7" pageNumber="408" type="reference_group">
<paragraph id="6B7AC609FFE1FFAAFF7DFAAF0EBC041E" blockId="7.[151,1437,1350,1435]" pageId="7" pageNumber="408">
<treatmentCitationGroup id="4BD5E127FFE1FFAAFF7DFAAF0EBC041E" pageId="7" pageNumber="408">
<treatmentCitation id="EA64E018FFE1FFAAFF7DFAAF0811043D" author="Dash, P. C. &amp; Viraktamath, C. A." box="[151,816,1350,1374]" page="76" pageId="7" pageNumber="408" year="1995">
<taxonomicName id="ACC5BD8AFFE1FFAAFF7DFAAF0811043D" ID-CoL="8RSCN" authority="Dash &amp; Viraktamath, 1995: 74 - 76" authorityName="Dash &amp; Viraktamath" authorityPageNumber="74 - 76" authorityYear="1995" box="[151,816,1350,1374]" class="Insecta" family="Cicadellidae" genus="Deltocephalus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tareni" subGenus="Recilia">
<emphasis id="59B11A1BFFE1FFAAFF7DFAAF0A08043D" box="[151,297,1350,1374]" italics="true" pageId="7" pageNumber="408">Deltocephalus</emphasis>
(
<emphasis id="59B11A1BFFE1FFAAFED2FAAF0AA1043D" box="[312,384,1350,1374]" italics="true" pageId="7" pageNumber="408">Recilia</emphasis>
)
<emphasis id="59B11A1BFFE1FFAAFE65FAAF0AEA043E" box="[399,459,1350,1373]" italics="true" pageId="7" pageNumber="408">tareni</emphasis>
<bibRefCitation id="0F54BBF8FFE1FFAAFE3BFAAF0811043D" author="Dash, P. C. &amp; Viraktamath, C. A." box="[465,816,1350,1374]" pageId="7" pageNumber="408" pagination="71 - 78" refId="ref7341" refString="Dash, P. C. &amp; Viraktamath, C. A. (1995) Two new species of grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) from Orissa, India. Hexapoda, 7, 71 - 78." type="journal article" year="1995">Dash &amp; Viraktamath, 1995: 7476</bibRefCitation>
</taxonomicName>
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, figs 115;
<treatmentCitation id="EA64E018FFE1FFAAFC4DFAAF0FC1043D" author="Dash, P. C. &amp; Viraktamath, C. A." box="[935,1248,1350,1374]" page="16" pageId="7" pageNumber="408" year="1998">
<bibRefCitation id="0F54BBF8FFE1FFAAFC4DFAAF0FC1043D" author="Dash, P. C. &amp; Viraktamath, C. A." box="[935,1248,1350,1374]" pageId="7" pageNumber="408" pagination="1 - 59" refId="ref7387" refString="Dash, P. C. &amp; Viraktamath, C. A. (1998) A review of the Indian and Nepalese grass feeding leafhopper genus Deltocephalus (Recilia) (Homoptera: Cicadellidae) with description of new species. Hexapoda, 10 (1 &amp; 2), 1 - 59." type="journal article" year="1998">Dash &amp; Viraktamath, 1998: 16</bibRefCitation>
</treatmentCitation>
, figs 7884.
<treatmentCitation id="EA64E018FFE1FFAAFF7DFA8C09A4041E" author="Webb, M. D. &amp; Viraktamath, C. A." box="[151,645,1380,1405]" page="22" pageId="7" pageNumber="408" year="2009">
<taxonomicName id="ACC5BD8AFFE1FFAAFF7DFA8C09A4041E" ID-CoL="8RLWH" authority=", Webb &amp; Viraktamath, 2009: 22" authorityName="Webb &amp; Viraktamath" authorityPageNumber="22" authorityYear="2009" box="[151,645,1380,1405]" class="Insecta" family="Cicadellidae" genus="Maiestas" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tareni" status="comb. nov.">
<emphasis id="59B11A1BFFE1FFAAFF7DFA8C0A1D041F" box="[151,316,1381,1404]" italics="true" pageId="7" pageNumber="408">Maiestas tareni,</emphasis>
<bibRefCitation id="0F54BBF8FFE1FFAAFEA8FA8D09A4041E" author="Webb, M. D. &amp; Viraktamath, C. A." box="[322,645,1380,1405]" pageId="7" pageNumber="408" pagination="1 - 64" refId="ref8291" refString="Webb, M. D. &amp; Viraktamath, C. A. (2009) Annotated check-list, generic key and new species of Old World Deltocephalini leafhoppers with nomenclatorial changes in the Deltocephalus group and other Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa, 2163 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 2163.1.1" type="journal article" year="2009">Webb &amp; Viraktamath, 2009: 22</bibRefCitation>
</taxonomicName>
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,
<taxonomicNameLabel id="4282A760FFE1FFAAFD7AFA8D0825041F" box="[656,772,1380,1404]" pageId="7" pageNumber="408" rank="species">comb. nov.</taxonomicNameLabel>
;
<treatmentCitation id="EA64E018FFE1FFAAFCE4FA8D0F32041E" author="Khatri I. &amp; Webb, M. D." box="[782,1043,1380,1405]" page="11" pageId="7" pageNumber="408" year="2010">
<bibRefCitation id="0F54BBF8FFE1FFAAFCE4FA8D0F32041E" author="Khatri I. &amp; Webb, M. D." box="[782,1043,1380,1405]" pageId="7" pageNumber="408" pagination="1 - 47" refId="ref7812" refString="Khatri I., &amp; Webb, M. D. (2010) The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa, 2365 (1), 1 - 47. https: // doi. org / 10.11646 / zootaxa. 2365.1.1" type="journal article" year="2010">Khatri &amp; Webb, 2010: 11</bibRefCitation>
</treatmentCitation>
, plate 2d, fig. 11;
<bibRefCitation id="0F54BBF8FFE1FFAAFB27FA8D0EBC041E" author="Naveed, H. &amp; Sohail, K. &amp; Islam, W. &amp; Zhang, Y. &amp; Bu, W. J." box="[1229,1437,1380,1405]" pageId="7" pageNumber="408" pagination="283 - 292" refId="ref8042" refString="Naveed, H., Sohail, K., Islam, W., Zhang, Y. &amp; Bu, W. J. (2019) A review of the leafhopper tribe Deltocephalini (Hemiptera: Cicadellidae: Deltocephalinae) from Pakistan. Revista Chilena de Entomologia, 45 (2), 283 - 292. https: // doi. org / 10.35249 / rche. 45.2.19.16" type="journal article" year="2019">
Naveed
<emphasis id="59B11A1BFFE1FFAAFAC8FA8C0E6E041F" box="[1314,1359,1380,1404]" italics="true" pageId="7" pageNumber="408">et al</emphasis>
., 2019:
</bibRefCitation>
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</paragraph>
<paragraph id="6B7AC609FFE1FFAAFF2DFA6A0A9504F8" blockId="7.[151,1437,1350,1435]" box="[199,436,1411,1435]" pageId="7" pageNumber="408">290, figs 2GI, 3NO).</paragraph>
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<emphasis id="59B11A1BFFE1FFAAFF7DFA200A5E0480" bold="true" box="[151,383,1481,1507]" pageId="7" pageNumber="408">Material examined.</emphasis>
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:
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(
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;
<materialsCitation id="DBADCC54FFE1FFAAFE0AFA040F140764" ID-GBIF-Occurrence="3396018310" box="[480,1077,1517,1544]" collectingDate="1985-08" collectionCode="BMNH" collectorName="C. &amp; L. O'Brien" country="Pakistan" pageId="7" pageNumber="408" specimenCount="1" specimenCount-male="1" stateProvince="Punjab">
<specimenCount id="7DC30D80FFE1FFAAFE0AFA0409220764" box="[480,515,1517,1543]" pageId="7" pageNumber="408" type="male">1♂</specimenCount>
Rawalpindi,
<date id="1F7BE0C9FFE1FFAAFD73FA040820076B" box="[665,769,1517,1544]" pageId="7" pageNumber="408" value="1985-08">
<collectingDate id="0F3F1921FFE1FFAAFD73FA040820076B" box="[665,769,1517,1544]" pageId="7" pageNumber="408" value="1985-08">viii.1985</collectingDate>
</date>
,
<collectorName id="C630A3DFFFE1FFAAFCE6FA0408E50764" box="[780,964,1517,1544]" pageId="7" pageNumber="408">C. &amp; L. OBrien</collectorName>
(
<collectionCode id="0DD45ECCFFE1FFAAFC39FA040F0C0764" box="[979,1069,1517,1543]" pageId="7" pageNumber="408">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="23DF9582FFE1FFAAFF2DF9F809470748" box="[199,614,1553,1579]" pageId="7" pageNumber="408" type="distribution">
<paragraph id="6B7AC609FFE1FFAAFF2DF9F809470748" blockId="7.[151,1437,1481,1760]" box="[199,614,1553,1579]" pageId="7" pageNumber="408">
<emphasis id="59B11A1BFFE1FFAAFF2DF9F80A400748" bold="true" box="[199,353,1553,1579]" pageId="7" pageNumber="408">Distribution.</emphasis>
<collectingCountry id="13D28699FFE1FFAAFE8DF9F80AE80748" box="[359,457,1553,1579]" name="Pakistan" pageId="7" pageNumber="408">Pakistan</collectingCountry>
,
<collectingCountry id="13D28699FFE1FFAAFE3EF9F8093B0748" box="[468,538,1553,1579]" name="China" pageId="7" pageNumber="408">China</collectingCountry>
,
<collectingCountry id="13D28699FFE1FFAAFDCFF9F809430748" box="[549,610,1553,1579]" name="India" pageId="7" pageNumber="408">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="23DF9582FFE1FFAAFF2DF9DC0A300783" pageId="7" pageNumber="408" type="discussion">
<paragraph id="6B7AC609FFE1FFAAFF2DF9DC0A300783" blockId="7.[151,1437,1481,1760]" pageId="7" pageNumber="408">
<emphasis id="59B11A1BFFE1FFAAFF2DF9DC0A1A072C" bold="true" box="[199,315,1589,1615]" pageId="7" pageNumber="408">Remarks.</emphasis>
This species can be distinguished by its relatively straight and stout style apical apophysis with serrated inner margin and by the aedeagus in lateral view evenly tapered from base to apex and relatively straight. Some slight differences are shown between the specimens studied here (
<figureCitation id="F3FEDA8CFFE1FFAAFC32F9940F7B07FB" box="[984,1114,1661,1688]" captionStart="FIGURE 5" captionStartId="9.[151,250,1749,1773]" captionTargetBox="[217,1365,194,1721]" captionTargetId="figure-19@9.[205,1383,181,1725]" captionTargetPageId="9" captionText="FIGURE 5. Maiestas tareni (Dash &amp; Viraktamath). AH: this study, A: habitus, dorsal view; B: habitus, lateral view; C: head and thorax, dorsal view; D: face; E: male genital capsule, lateral view; F: subgenital plate, valve and styles, dorsal view; G &amp; H: connective and aedeagus, dorsal and lateral view; IN: after Zhang &amp; Duan, 2011, I: habitus, dorsal view; J: male pygofer lobe, lateral view; K: subgenital plate, ventral view; L: style, dorsal view; M &amp; N: connective and aedeagus, dorsal and lateral view; OT: after Khatri &amp; Webb, 2010, O: habitus, dorsal view; P: head and thorax, dorsal view; Q: subgenital plate, ventral view; R: style, dorsal view; S &amp; T: connective and aedeagus, dorsal and lateral view; UZ: after Dash &amp; Viraktamath, 1995, U: head and thorax, dorsal view; V: face; W: subgenital plates and valve, dorsal view; X: style, dorsal view; Y &amp; Z: connective and aedeagus, dorsal and lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.5635926" httpUri="https://zenodo.org/record/5635926/files/figure.png" pageId="7" pageNumber="408">Figs 5AH</figureCitation>
,
<collectingRegion id="A90108EBFFE1FFAAFB82F9940F9907FB" box="[1128,1208,1661,1688]" country="Pakistan" name="Punjab" pageId="7" pageNumber="408">Punjab</collectingRegion>
) and between some published figures (
<figureCitation id="F3FEDA8CFFE1FFAAFE82F9480AF807DF" box="[360,473,1697,1724]" captionStart="FIGURE 5" captionStartId="9.[151,250,1749,1773]" captionTargetBox="[217,1365,194,1721]" captionTargetId="figure-19@9.[205,1383,181,1725]" captionTargetPageId="9" captionText="FIGURE 5. Maiestas tareni (Dash &amp; Viraktamath). AH: this study, A: habitus, dorsal view; B: habitus, lateral view; C: head and thorax, dorsal view; D: face; E: male genital capsule, lateral view; F: subgenital plate, valve and styles, dorsal view; G &amp; H: connective and aedeagus, dorsal and lateral view; IN: after Zhang &amp; Duan, 2011, I: habitus, dorsal view; J: male pygofer lobe, lateral view; K: subgenital plate, ventral view; L: style, dorsal view; M &amp; N: connective and aedeagus, dorsal and lateral view; OT: after Khatri &amp; Webb, 2010, O: habitus, dorsal view; P: head and thorax, dorsal view; Q: subgenital plate, ventral view; R: style, dorsal view; S &amp; T: connective and aedeagus, dorsal and lateral view; UZ: after Dash &amp; Viraktamath, 1995, U: head and thorax, dorsal view; V: face; W: subgenital plates and valve, dorsal view; X: style, dorsal view; Y &amp; Z: connective and aedeagus, dorsal and lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.5635926" httpUri="https://zenodo.org/record/5635926/files/figure.png" pageId="7" pageNumber="408">Figs 5IT</figureCitation>
) and the original description (
<figureCitation id="F3FEDA8CFFE1FFAAFCC0F948088807DF" box="[810,937,1697,1724]" captionStart="FIGURE 5" captionStartId="9.[151,250,1749,1773]" captionTargetBox="[217,1365,194,1721]" captionTargetId="figure-19@9.[205,1383,181,1725]" captionTargetPageId="9" captionText="FIGURE 5. Maiestas tareni (Dash &amp; Viraktamath). AH: this study, A: habitus, dorsal view; B: habitus, lateral view; C: head and thorax, dorsal view; D: face; E: male genital capsule, lateral view; F: subgenital plate, valve and styles, dorsal view; G &amp; H: connective and aedeagus, dorsal and lateral view; IN: after Zhang &amp; Duan, 2011, I: habitus, dorsal view; J: male pygofer lobe, lateral view; K: subgenital plate, ventral view; L: style, dorsal view; M &amp; N: connective and aedeagus, dorsal and lateral view; OT: after Khatri &amp; Webb, 2010, O: habitus, dorsal view; P: head and thorax, dorsal view; Q: subgenital plate, ventral view; R: style, dorsal view; S &amp; T: connective and aedeagus, dorsal and lateral view; UZ: after Dash &amp; Viraktamath, 1995, U: head and thorax, dorsal view; V: face; W: subgenital plates and valve, dorsal view; X: style, dorsal view; Y &amp; Z: connective and aedeagus, dorsal and lateral view respectively." figureDoi="http://doi.org/10.5281/zenodo.5635926" httpUri="https://zenodo.org/record/5635926/files/figure.png" pageId="7" pageNumber="408">Figs 5UZ</figureCitation>
), suggesting that more than one species may be present.
</paragraph>
</subSubSection>
</treatment>
</document>