522 lines
63 KiB
XML
522 lines
63 KiB
XML
<document id="20B8ADFCBC6BBCBEF851C18DC469663D" ID-DOI="10.5281/zenodo.5739687" ID-ISSN="0374-1036" ID-Zenodo-Dep="5739687" ID-ZooBank="0CC4ABA4-70DA-42BD-9922-45316034E5BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638252473029" checkinUser="marcus" docAuthor="Král, David & Šípek, Petr" docDate="2013" docId="E2052558FFDBFFA0FE2AAC960E12FE15" docLanguage="en" docName="ActaEntMusNatPra.2013.53.2.633-648.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Aphodius (Gilletianus) rajawatorum Král & Šípek 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="643" masterDocId="1E3C5D20FFDDFFAAFFA3AE2E0C29FF81" masterDocTitle="Aphodiinae (Coleoptera: Scarabaeidae) of the Goa, Maharashtra and Rajasthan (India) with description of Aphodius (Gilletianus) rajawatorum sp. nov." masterLastPageNumber="648" masterPageNumber="633" pageNumber="639" updateTime="1699435488041" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="033FA43F1096C049A34B49A82596145E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E8F7FBD33CAD4E9F9B8FED6184849011">Aphodiinae (Coleoptera: Scarabaeidae) of the Goa, Maharashtra and Rajasthan (India) with description of Aphodius (Gilletianus) rajawatorum sp. nov.</mods:title>
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<mods:namePart id="D9F99878E4B05C15D6C5A25EC7EECB45">Král, David</mods:namePart>
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<mods:roleTerm id="BA58B0DE58F41336CC233DC7211F1373">Author</mods:roleTerm>
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<mods:namePart id="05A6641D644C73BA2FADB145663C736A">Šípek, Petr</mods:namePart>
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<mods:typeOfResource id="89BDF0C1A80A3348CA3344471D8E77CD">text</mods:typeOfResource>
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<mods:title id="1A18A480F16710468E0A8BA38779D3D6">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="58307CFC4D9BF1D032C87A93BE0580FA">
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<mods:date id="FB874478634E655F0B040209D29DD9D7">2013</mods:date>
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<mods:number id="D3E31F2D5F91EE25DFA7EF102C295B10">2013-11-15</mods:number>
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<mods:number id="6D2285A2E3FD23ABCF3FD8B74A0C3D9A">53</mods:number>
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<mods:identifier id="AAB624B15FC78E1C827FDDD416BD32D7" type="DOI">10.5281/zenodo.5739687</mods:identifier>
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<mods:identifier id="C5B4F454BCFF38BB5850858256E82B5C" type="ISSN">0374-1036</mods:identifier>
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<treatment id="E2052558FFDBFFA0FE2AAC960E12FE15" LSID="urn:lsid:plazi:treatment:E2052558FFDBFFA0FE2AAC960E12FE15" httpUri="http://treatment.plazi.org/id/E2052558FFDBFFA0FE2AAC960E12FE15" lastPageId="10" lastPageNumber="643" pageId="6" pageNumber="639">
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<subSubSection id="22B6C7C5FFDBFFACFE2AAC960F5CFD50" box="[393,885,696,721]" pageId="6" pageNumber="639" type="nomenclature">
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<paragraph id="6A13944EFFDBFFACFE2AAC960F5CFD50" blockId="6.[393,885,696,721]" box="[393,885,696,721]" pageId="6" pageNumber="639">
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<heading id="315B2322FFDBFFACFE2AAC960F5CFD50" bold="true" box="[393,885,696,721]" centered="true" fontSize="8" level="2" pageId="6" pageNumber="639" reason="3">
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<emphasis id="58D8485CFFDBFFACFE2AAC960F5CFD50" bold="true" box="[393,885,696,721]" pageId="6" pageNumber="639">
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<taxonomicName id="ADACEFCDFFDBFFACFE2AAC960F30FD50" authority="Král & Šípek, 2013" authorityName="Král & Šípek" authorityYear="2013" box="[393,793,696,721]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="639" phylum="Arthropoda" rank="species" species="rajawatorum" status="sp. nov." subGenus="Gilletianus">
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<emphasis id="58D8485CFFDBFFACFE2AAC960DDBFD50" bold="true" box="[393,498,696,721]" italics="true" pageId="6" pageNumber="639">Aphodius</emphasis>
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(
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||
<emphasis id="58D8485CFFDBFFACFDA1AC960E55FD50" bold="true" box="[514,636,696,721]" italics="true" pageId="6" pageNumber="639">Gilletianus</emphasis>
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)
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<emphasis id="58D8485CFFDBFFACFD28AC970F30FD50" bold="true" box="[651,793,697,721]" italics="true" pageId="6" pageNumber="639">rajawatorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="43EBF527FFDBFFACFC83AC970F5CFD50" box="[800,885,697,721]" pageId="6" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="22B6C7C5FFDBFFACFD8FACF60EFBFD6A" box="[556,722,728,747]" pageId="6" pageNumber="639" type="description">
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<paragraph id="6A13944EFFDBFFACFD8FACF60EFBFD6A" blockId="6.[556,722,728,747]" box="[556,722,728,747]" pageId="6" pageNumber="639">
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(
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<figureCitation id="F29788CBFFDBFFACFD91ACF60E41FD6A" box="[562,616,728,747]" captionStart="Fig" captionStartId="2.[858,891,862,881]" captionTargetBox="[149,838,219,959]" captionTargetId="figure-208@2.[149,839,219,959]" captionTargetPageId="2" captionText="Fig. 1. Sketch map of India with marked localities visited (states of Goa, Maharashtra and Rajasthan)." figureDoi="http://doi.org/10.5281/zenodo.5739691" httpUri="https://zenodo.org/record/5739691/files/figure.png" pageId="6" pageNumber="639">Figs 1</figureCitation>
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,
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||
<figureCitation id="F29788CBFFDBFFACFDD0ACF60E80FD6A" box="[627,681,728,747]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="6" pageNumber="639">2A–D</figureCitation>
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,
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<figureCitation id="F29788CBFFDBFFACFD12ACF60EE2FD6A" box="[689,715,728,747]" captionStart="Fig" captionStartId="9.[113,144,988,1007]" captionTargetBox="[127,1038,223,954]" captionTargetId="figure-185@9.[113,1088,214,966]" captionTargetPageId="9" captionText="Fig. 3. Head, pronotum and protibia, JJ, dorsal aspect. A – Aphodius (Gilletianus) fukiensis Balthasar, 1952 (China, Fujian: Kuatun; syntype, NMPC), B – A. (G.) rajawatorum sp. nov. (holotype), C – A. (G.) segmentaroides A. Schmidt, 1909 (India, Rajasthan: 50 km W of Agra, Bharatpur, NMPC), D – A. (G.) therondi Balthasar, 1963 (Vietnam: Tonkin, Tam Dao, paratype, NMPC); arrows show diagnostic characters mentioned in the description of the new species and in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5739697" httpUri="https://zenodo.org/record/5739697/files/figure.png" pageId="6" pageNumber="639">3B</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="22B6C7C5FFDBFFACFF34AD2C0F30FC20" pageId="6" pageNumber="639" type="materials_examined">
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<paragraph id="6A13944EFFDBFFACFF34AD2C0D03FCAE" blockId="6.[151,1127,768,930]" pageId="6" pageNumber="639">
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<emphasis id="58D8485CFFDBFFACFF34AD2C0D3AFC94" bold="true" box="[151,275,769,789]" pageId="6" pageNumber="639">
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<typeStatus id="B5172AECFFDBFFACFF34AD2C0CEDFC94" box="[151,196,770,789]" pageId="6" pageNumber="639">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
NW
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<collectingCountry id="12BBD4DEFFDBFFACFEE5AD2F0D5DFC95" box="[326,372,769,788]" name="India" pageId="6" pageNumber="639">India</collectingCountry>
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||
,
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||
<collectingRegion id="A8685AACFFDBFFACFEDCAD2F0E00FC95" box="[383,553,769,788]" country="India" name="Rajasthan" pageId="6" pageNumber="639">Rajasthan province</collectingRegion>
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||
,
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||
<quantity id="AD5439ABFFDBFFACFD97AD2F0E45FC95" box="[564,620,769,788]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="639" unit="km" value="50.0">50 km</quantity>
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||
W of Agra, Bharatpur env.,
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||
<geoCoordinate id="0F98F289FFDBFFACFCCCAD2C0FF8FC94" box="[879,977,768,789]" degrees="27" direction="north" minutes="12.42" orientation="latitude" pageId="6" pageNumber="639" precision="9" value="27.207">
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27
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||
<emphasis id="58D8485CFFDBFFACFC27AD2E0FA4FC95" box="[900,909,768,788]" italics="true" pageId="6" pageNumber="639">°</emphasis>
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12.42′N
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||
</geoCoordinate>
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||
<geoCoordinate id="0F98F289FFDBFFACFC7AAD2C0812FC94" box="[985,1083,768,789]" degrees="77" direction="east" minutes="30.48" orientation="longitude" pageId="6" pageNumber="639" precision="9" value="77.508">
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77
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||
<emphasis id="58D8485CFFDBFFACFC4DAD2E0FDEFC95" box="[1006,1015,768,788]" italics="true" pageId="6" pageNumber="639">°</emphasis>
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||
30.48′E
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||
</geoCoordinate>
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,
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||
<quantity id="AD5439ABFFDBFFACFBE5AD2C0C81FCB1" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="639" unit="m" value="220.0">220 m</quantity>
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a. s. l. (
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||
<figureCitation id="F29788CBFFDBFFACFF48AD320D34FCAE" box="[235,285,796,815]" captionStart="Fig" captionStartId="2.[858,891,862,881]" captionTargetBox="[149,838,219,959]" captionTargetId="figure-208@2.[149,839,219,959]" captionTargetPageId="2" captionText="Fig. 1. Sketch map of India with marked localities visited (states of Goa, Maharashtra and Rajasthan)." figureDoi="http://doi.org/10.5281/zenodo.5739691" httpUri="https://zenodo.org/record/5739691/files/figure.png" pageId="6" pageNumber="639">Fig. 1</figureCitation>
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).
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||
</paragraph>
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||
<paragraph id="6A13944EFFDBFFACFF34AD100F30FC20" blockId="6.[151,1127,768,930]" pageId="6" pageNumber="639">
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||
<emphasis id="58D8485CFFDBFFACFF34AD100D32FCD3" bold="true" box="[151,283,830,850]" pageId="6" pageNumber="639">Type material.</emphasis>
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||
<typeStatus id="B5172AECFFDBFFACFEBCAD110D5BFCD1" box="[287,370,831,850]" pageId="6" pageNumber="639" type="holotype">HOLOTYPE</typeStatus>
|
||
and
|
||
<materialsCitation id="DAC49E13FFDBFFACFE3AAD6C0F9BFCED" collectionCode="AGRA" country="India" county="Nos" elevation="220" latitude="27.207" location="Expedition" longLatPrecision="12" longitude="77.508" municipality="Bharatpur" pageId="6" pageNumber="639" specimenCount="1" stateProvince="Rajasthan" typeStatus="paratype">
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||
<typeStatus id="B5172AECFFDBFFACFE3AAD6C0DC2FCD1" box="[409,491,834,848]" pageId="6" pageNumber="639" type="paratype">PARATYPES</typeStatus>
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<collectingCounty id="8372ECC2FFDBFFACFE53AD110E3BFCD3" box="[496,530,831,850]" pageId="6" pageNumber="639">Nos</collectingCounty>
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||
1–4 (all JJ), labelled: ‘
|
||
<collectingCountry id="12BBD4DEFFDBFFACFD7EAD100F35FCD0" box="[733,796,830,849]" name="India" pageId="6" pageNumber="639">INDIA</collectingCountry>
|
||
bor. occ. /
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||
<collectingRegion id="A8685AACFFDBFFACFCD4AD10086AFCD0" box="[887,1091,830,849]" country="India" name="Rajasthan" pageId="6" pageNumber="639">RAJASTHAN province</collectingRegion>
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||
/
|
||
<quantity id="AD5439ABFFDBFFACFBF2AD100C9BFCED" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="639" unit="km" value="50.0">50 km</quantity>
|
||
W. of
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||
<collectionCode id="0CBD0C8BFFDBFFACFF49AD770D0CFCED" box="[234,293,857,876]" pageId="6" pageNumber="639">AGRA</collectionCode>
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/
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||
<collectingMunicipality id="8A770E34FFDBFFACFE95AD770DA2FCED" box="[310,395,857,876]" pageId="6" pageNumber="639">Bharatpur</collectingMunicipality>
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env.:
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||
<quantity id="AD5439ABFFDBFFACFE1DAD770DC6FCED" box="[446,495,857,876]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="639" unit="m" value="220.0">
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<elevation id="E181737DFFDBFFACFE1DAD770DC6FCED" box="[446,495,857,876]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="639" unit="m" value="220.0">220m</elevation>
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||
</quantity>
|
||
//
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||
<collectingCountry id="12BBD4DEFFDBFFACFDA6AD770E69FCED" box="[517,576,857,876]" name="India" pageId="6" pageNumber="639">INDIA</collectingCountry>
|
||
2002
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||
<location id="6F73C295FFDBFFACFDD5AD770EFDFCED" LSID="urn:lsid:plazi:treatment:E2052558FFDBFFA0FE2AAC960E12FE15:6F73C295FFDBFFACFDD5AD770EFDFCED" box="[630,724,857,876]" country="India" county="Nos" latitude="27.207" longLatPrecision="12" longitude="77.508" municipality="Bharatpur" name="Expedition" pageId="6" pageNumber="639" stateProvince="Rajasthan">Expedition</location>
|
||
/
|
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<geoCoordinate id="0F98F289FFDBFFACFD47AD770F60FCED" box="[740,841,856,876]" degrees="27" direction="north" minutes="12.42" orientation="latitude" pageId="6" pageNumber="639" precision="9" value="27.207">
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||
27
|
||
<emphasis id="58D8485CFFDBFFACFD59AD760F2AFCED" box="[762,771,856,876]" italics="true" pageId="6" pageNumber="639">°</emphasis>
|
||
12.42’N
|
||
</geoCoordinate>
|
||
<geoCoordinate id="0F98F289FFDBFFACFCEDAD770F9BFCED" box="[846,946,856,876]" degrees="77" direction="east" minutes="30.48" orientation="longitude" pageId="6" pageNumber="639" precision="9" value="77.508">
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||
77
|
||
<emphasis id="58D8485CFFDBFFACFCC0AD760F45FCED" box="[867,876,856,876]" italics="true" pageId="6" pageNumber="639">°</emphasis>
|
||
30.48’E
|
||
</geoCoordinate>
|
||
</materialsCitation>
|
||
;
|
||
<date id="1E12B28EFFDBFFACFC18AD77084EFCED" box="[955,1127,857,876]" pageId="6" pageNumber="639" value="2002-08-31" valueMax="2002-09-05" valueMin="2002-08-31">31.VIII.–5. IX.2002</date>
|
||
/ P. Šípek & M. Fikáček lgt. [printed] //
|
||
<taxonomicName id="ADACEFCDFFDBFFACFE45AD5A0E83FC06" authority="" authorityName="Balthasar" authorityYear="1933" box="[486,682,884,903]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="639" phylum="Arthropoda" rank="subGenus" subGenus="Gilletianus">Aphodius (Gilletianus)</taxonomicName>
|
||
/ rajawatorum
|
||
<taxonomicNameLabel id="43EBF527FFDBFFACFC89AD5A0F47FC06" box="[810,878,884,903]" pageId="6" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, /
|
||
<typeStatus id="B5172AECFFDBFFACFC23AD5A0FD6FC06" box="[896,1023,884,903]" pageId="6" pageNumber="639" type="holotype">HOLOTYPUS</typeStatus>
|
||
[or]
|
||
<collectionCode id="0CBD0C8BFFDBFFACFB8BAD5A0CF2FC20" pageId="6" pageNumber="639">PARA- TYPUS</collectionCode>
|
||
No. x J / David Král & Petr Šípek det. 2011 [red label, printed]’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22B6C7C5FFDBFFA3FF34AD940FE2FAB2" lastPageId="9" lastPageNumber="642" pageId="6" pageNumber="639" type="description">
|
||
<paragraph id="6A13944EFFDBFFACFF34AD940867FBB3" blockId="6.[151,1127,953,1586]" pageId="6" pageNumber="639">
|
||
<emphasis id="58D8485CFFDBFFACFF34AD940D49FC53" bold="true" box="[151,352,953,978]" pageId="6" pageNumber="639">
|
||
Description.
|
||
<emphasis id="58D8485CFFDBFFACFE8BAD970D49FC53" bold="true" box="[296,352,953,978]" italics="true" pageId="6" pageNumber="639">Male</emphasis>
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="B5172AECFFDBFFACFECDAD940DF9FC53" box="[366,464,954,978]" pageId="6" pageNumber="639" type="holotype">holotype</typeStatus>
|
||
; left metatarsomeres 2–5 missing in the specimen). Total body length 3.3 mm. Body elongately oval, moderately convex, dorsal surface shining; colour brown, anterior and lateral parts of head, sides of pronotum, apical elytron declivity, and ventral surface lighter; subapical darker spots of elytron faintly marked (
|
||
<figureCitation id="F29788CBFFDBFFACFC3FAA34083DFBB3" box="[924,1044,1050,1074]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="6" pageNumber="639">Figs 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="F29788CBFFDBFFACFBBCAA340868FBB3" box="[1055,1089,1050,1074]" captionStart="Fig" captionStartId="9.[113,144,988,1007]" captionTargetBox="[127,1038,223,954]" captionTargetId="figure-185@9.[113,1088,214,966]" captionTargetPageId="9" captionText="Fig. 3. Head, pronotum and protibia, JJ, dorsal aspect. A – Aphodius (Gilletianus) fukiensis Balthasar, 1952 (China, Fujian: Kuatun; syntype, NMPC), B – A. (G.) rajawatorum sp. nov. (holotype), C – A. (G.) segmentaroides A. Schmidt, 1909 (India, Rajasthan: 50 km W of Agra, Bharatpur, NMPC), D – A. (G.) therondi Balthasar, 1963 (Vietnam: Tonkin, Tam Dao, paratype, NMPC); arrows show diagnostic characters mentioned in the description of the new species and in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5739697" httpUri="https://zenodo.org/record/5739697/files/figure.png" pageId="6" pageNumber="639">3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFDBFFACFF16AA150D39FB50" blockId="6.[151,1127,953,1586]" pageId="6" pageNumber="639">
|
||
<emphasis id="58D8485CFFDBFFACFF16AA150CC6FBD3" box="[181,239,1083,1106]" italics="true" pageId="6" pageNumber="639">Head</emphasis>
|
||
(
|
||
<figureCitation id="F29788CBFFDBFFACFF5CAA140D50FBD3" box="[255,377,1082,1106]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="6" pageNumber="639">Figs 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="F29788CBFFDBFFACFE24AA140D8FFBD3" box="[391,422,1082,1106]" captionStart="Fig" captionStartId="9.[113,144,988,1007]" captionTargetBox="[127,1038,223,954]" captionTargetId="figure-185@9.[113,1088,214,966]" captionTargetPageId="9" captionText="Fig. 3. Head, pronotum and protibia, JJ, dorsal aspect. A – Aphodius (Gilletianus) fukiensis Balthasar, 1952 (China, Fujian: Kuatun; syntype, NMPC), B – A. (G.) rajawatorum sp. nov. (holotype), C – A. (G.) segmentaroides A. Schmidt, 1909 (India, Rajasthan: 50 km W of Agra, Bharatpur, NMPC), D – A. (G.) therondi Balthasar, 1963 (Vietnam: Tonkin, Tam Dao, paratype, NMPC); arrows show diagnostic characters mentioned in the description of the new species and in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5739697" httpUri="https://zenodo.org/record/5739697/files/figure.png" pageId="6" pageNumber="639">3B</figureCitation>
|
||
) semicircular; clypeus almost semicircular, anterior margin only weakly sinuate, not considerably upturned, sides regularly round towards small, right-angled genae, distinctly exceeding large eyes; frontal suture slightly impressed; surface punctures double, fine, superficial, evenly distributed, large punctures about three times larger than small ones.
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFDBFFACFF16AAF50F4CFAB3" blockId="6.[151,1127,953,1586]" pageId="6" pageNumber="639">
|
||
<emphasis id="58D8485CFFDBFFACFF16AAF50D36FB73" box="[181,287,1243,1266]" italics="true" pageId="6" pageNumber="639">Pronotum</emphasis>
|
||
(
|
||
<figureCitation id="F29788CBFFDBFFACFE92AAF40D84FB73" box="[305,429,1242,1266]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="6" pageNumber="639">Figs 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="F29788CBFFDBFFACFE18AAF40DF2FB73" box="[443,475,1242,1266]" captionStart="Fig" captionStartId="9.[113,144,988,1007]" captionTargetBox="[127,1038,223,954]" captionTargetId="figure-185@9.[113,1088,214,966]" captionTargetPageId="9" captionText="Fig. 3. Head, pronotum and protibia, JJ, dorsal aspect. A – Aphodius (Gilletianus) fukiensis Balthasar, 1952 (China, Fujian: Kuatun; syntype, NMPC), B – A. (G.) rajawatorum sp. nov. (holotype), C – A. (G.) segmentaroides A. Schmidt, 1909 (India, Rajasthan: 50 km W of Agra, Bharatpur, NMPC), D – A. (G.) therondi Balthasar, 1963 (Vietnam: Tonkin, Tam Dao, paratype, NMPC); arrows show diagnostic characters mentioned in the description of the new species and in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5739697" httpUri="https://zenodo.org/record/5739697/files/figure.png" pageId="6" pageNumber="639">3B</figureCitation>
|
||
) more or less rectangular, sides regularly round, distinctly bordered, posterior angles obtuse, base without border, surface punctation double, coarse punctures relatively densely distributed, concentrated laterobasally.
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFDBFFACFF16AB150E95FAD3" blockId="6.[151,1127,953,1586]" box="[181,700,1338,1362]" pageId="6" pageNumber="639">
|
||
<emphasis id="58D8485CFFDBFFACFF16AB150D75FAD3" box="[181,348,1339,1362]" italics="true" pageId="6" pageNumber="639">Scutellar shield</emphasis>
|
||
narrowly triangular, impunctate.
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFDBFFACFF16AB750D90FA70" blockId="6.[151,1127,953,1586]" pageId="6" pageNumber="639">
|
||
<emphasis id="58D8485CFFDBFFACFF16AB750CDEFAF3" box="[181,247,1371,1394]" italics="true" pageId="6" pageNumber="639">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="F29788CBFFDBFFACFEA7AB740D7CFAF3" box="[260,341,1370,1394]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="6" pageNumber="639">Fig. 2A</figureCitation>
|
||
) suboval, slightly widened toward apex, minutely macrosetaceous apically, striae finely impressed and punctate, punctures separated by more than once their diameter, slightly crenate interval margins, intervals moderately convex in basal half, toward apex becoming completely flat, sutural interval almost flat in whole length, surface punctation superficial, fine and sparse.
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFDBFFACFF16ABD50D22F9B3" blockId="6.[151,1127,953,1586]" pageId="6" pageNumber="639">
|
||
<emphasis id="58D8485CFFDBFFACFF16ABD50D51F993" box="[181,376,1531,1554]" italics="true" pageId="6" pageNumber="639">Metaventral disc</emphasis>
|
||
(
|
||
<figureCitation id="F29788CBFFDBFFACFE2FABD40DCEF993" box="[396,487,1530,1554]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="6" pageNumber="639">Fig. 2B</figureCitation>
|
||
) shiny, flat, glabrous, with longitudinal line distinctly impressed.
|
||
</paragraph>
|
||
<caption id="3ED3C4C6FFDAFFADFFD2AEF50F9CFF6E" box="[113,949,219,239]" pageId="7" pageNumber="640" startId="7.[113,160,219,238]">
|
||
<paragraph id="6A13944EFFDAFFADFFD2AEF50F9CFF6E" blockId="7.[113,949,219,239]" box="[113,949,219,239]" pageId="7" pageNumber="640">
|
||
Table 2. Character matrix distinguishing
|
||
<taxonomicName id="ADACEFCDFFDAFFADFE6CAEF20E07FF6F" authorityName="Balthasar" authorityYear="1933" box="[463,558,220,238]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="642" phylum="Arthropoda" rank="subGenus" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFDAFFADFE6CAEF20E07FF6F" box="[463,558,220,238]" italics="true" pageId="7" pageNumber="640">Gilletianus</emphasis>
|
||
</taxonomicName>
|
||
species close to
|
||
<emphasis id="58D8485CFFDAFFADFD1EAEF20EE3FF6F" box="[701,714,220,238]" italics="true" pageId="7" pageNumber="640">A</emphasis>
|
||
.
|
||
<taxonomicName id="ADACEFCDFFDAFFADFD76AEF20F45FF6F" authority="Král & Šípek, 2013" authorityName="Král & Šípek" authorityYear="2013" box="[725,876,220,239]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="640" phylum="Arthropoda" rank="species" species="rajawatorum" status="sp. nov." subGenus="Gilletianus">
|
||
(
|
||
<emphasis id="58D8485CFFDAFFADFD7FAEF20EC2FF6F" box="[732,747,220,238]" italics="true" pageId="7" pageNumber="640">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFDAFFADFD5EAEF20F45FF6F" box="[765,876,220,238]" italics="true" pageId="7" pageNumber="640">rajawatorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="43EBF527FFDAFFADFCD1AEF20F9CFF6E" box="[882,949,220,239]" pageId="7" pageNumber="640" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6A13944EFFDAFFADFFDAAF230FC1FB31" pageId="7" pageNumber="640">
|
||
<table id="18AC66EEFFDA0055FFDAAF220810FA9A" box="[121,1081,268,1307]" gridcols="5" gridrows="11" pageId="7" pageNumber="640">
|
||
<tr id="D49C960CFFDA0055FFDAAF220810FED4" box="[121,1081,268,341]" gridrow="0" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAF220D6DFED4" box="[121,324,268,341]" gridcol="0" gridrow="0" pageId="7" pageNumber="640">character in male / species</th>
|
||
<th id="974DFF70FFDA0055FEFFAF220E21FED4" box="[348,520,268,341]" gridcol="1" gridrow="0" pageId="7" pageNumber="640">
|
||
<taxonomicName id="ADACEFCDFFDAFFADFECAAF230DD1FEBB" authority="Balthasar, 1952" authorityName="Balthasar" authorityYear="1952" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="640" phylum="Arthropoda" rank="species" species="fukiensis" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFDAFFADFECAAF230DD1FEBB" bold="true" italics="true" pageId="7" pageNumber="640">
|
||
A. (
|
||
<emphasis id="58D8485CFFDAFFADFE2AAF230DB1FEA1" bold="true" box="[393,408,269,288]" italics="true" pageId="7" pageNumber="640">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFDAFFADFE09AF220DD0FEA1" bold="true" box="[426,505,268,288]" italics="true" pageId="7" pageNumber="640">fukiensis</emphasis>
|
||
Balthasar, 1952
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="974DFF70FFDA0055FDB5AF220EE9FED4" box="[534,704,268,341]" gridcol="2" gridrow="0" pageId="7" pageNumber="640">
|
||
<emphasis id="58D8485CFFDAFFADFD82AF230E8AFEBB" bold="true" italics="true" pageId="7" pageNumber="640">
|
||
A. (
|
||
<emphasis id="58D8485CFFDAFFADFDE2AF230E79FEA1" bold="true" box="[577,592,269,288]" italics="true" pageId="7" pageNumber="640">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFDAFFADFDC0AF230E73FEBA" bold="true" italics="true" pageId="7" pageNumber="640">rajawato- rum</emphasis>
|
||
sp. nov.
|
||
</emphasis>
|
||
</th>
|
||
<th id="974DFF70FFDA0055FD72AF220F55FED4" box="[721,892,268,341]" gridcol="3" gridrow="0" pageId="7" pageNumber="640">
|
||
<emphasis id="58D8485CFFDAFFADFD7AAF230F5CFED4" bold="true" italics="true" pageId="7" pageNumber="640">
|
||
A. (
|
||
<emphasis id="58D8485CFFDAFFADFD5AAF230F21FEA1" bold="true" box="[761,776,269,288]" italics="true" pageId="7" pageNumber="640">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFDAFFADFCB9AF230F68FEBB" bold="true" italics="true" pageId="7" pageNumber="640">segmenta- roides</emphasis>
|
||
A. Schmidt, 1909
|
||
</emphasis>
|
||
</th>
|
||
<th id="974DFF70FFDA0055FC2FAF220810FED4" box="[908,1081,268,341]" gridcol="4" gridrow="0" pageId="7" pageNumber="640">
|
||
<taxonomicName id="ADACEFCDFFDAFFADFC3FAF230801FEBB" authority="Balthasar, 1963" authorityName="Balthasar" authorityYear="1963" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="640" phylum="Arthropoda" rank="species" species="therondi" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFDAFFADFC3FAF230801FEBB" bold="true" italics="true" pageId="7" pageNumber="640">
|
||
A. (
|
||
<emphasis id="58D8485CFFDAFFADFC1FAF230FE2FEA1" bold="true" box="[956,971,269,288]" italics="true" pageId="7" pageNumber="640">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFDAFFADFC7EAF220801FEA1" bold="true" box="[989,1064,268,288]" italics="true" pageId="7" pageNumber="640">therondi</emphasis>
|
||
Balthasar, 1963
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAF4E0810FE79" box="[121,1081,352,504]" gridrow="1" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAF4E0D6DFE79" box="[121,324,352,504]" gridcol="0" gridrow="1" pageId="7" pageNumber="640">clypeus outline</th>
|
||
<td id="974DFF70FFDA0055FEFFAF4E0E21FE79" box="[348,520,352,504]" gridcol="1" gridrow="1" pageId="7" pageNumber="640">trapezoidal, anterior margin distinctly sinuate, considerab- ly upturned, sides almost straight (Fig. 3A)</td>
|
||
<td id="974DFF70FFDA0055FDB5AF4E0EE9FE79" box="[534,704,352,504]" gridcol="2" gridrow="1" pageId="7" pageNumber="640">semicircular, ante- rior margin only weakly sinuate, not considerably uptur- ned, sides round (Figs 2A, 3B)</td>
|
||
<td id="974DFF70FFDA0055FD72AF4E0F55FE79" box="[721,892,352,504]" gridcol="3" gridrow="1" pageId="7" pageNumber="640">trapezoidal, anterior margin very weakly sinuate, considerab- ly upturned, sides almost straight (Fig. 3C)</td>
|
||
<td id="974DFF70FFDA0055FC2FAF4E0810FE79" box="[908,1081,352,504]" gridcol="4" gridrow="1" pageId="7" pageNumber="640">trapezoidal, anterior margin very weakly sinuate, considerab- ly upturned, sides almost straight (Fig. 3D)</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAC2D0810FDB0" box="[121,1081,515,561]" gridrow="2" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAC2D0D6DFDB0" box="[121,324,515,561]" gridcol="0" gridrow="2" pageId="7" pageNumber="640">clypeus surface pun- ctation</th>
|
||
<td id="974DFF70FFDA0055FEFFAC2D0E21FDB0" box="[348,520,515,561]" gridcol="1" gridrow="2" pageId="7" pageNumber="640">simple (Fig. 3A)</td>
|
||
<td id="974DFF70FFDA0055FDB5AC2D0EE9FDB0" box="[534,704,515,561]" gridcol="2" gridrow="2" pageId="7" pageNumber="640">double (Figs 2A, 3B)</td>
|
||
<td id="974DFF70FFDA0055FD72AC2D0F55FDB0" box="[721,892,515,561]" gridcol="3" gridrow="2" pageId="7" pageNumber="640">simple (Fig. 3C)</td>
|
||
<td id="974DFF70FFDA0055FC2FAC2D0810FDB0" box="[908,1081,515,561]" gridcol="4" gridrow="2" pageId="7" pageNumber="640">double (Fig. 3D)</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAC120810FD1E" box="[121,1081,572,671]" gridrow="3" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAC120D6DFD1E" box="[121,324,572,671]" gridcol="0" gridrow="3" pageId="7" pageNumber="640">pronotum outline</th>
|
||
<td id="974DFF70FFDA0055FEFFAC120E21FD1E" box="[348,520,572,671]" gridcol="1" gridrow="3" pageId="7" pageNumber="640">sides round, pos- terior angle round (Fig. 3A)</td>
|
||
<td id="974DFF70FFDA0055FDB5AC120EE9FD1E" box="[534,704,572,671]" gridcol="2" gridrow="3" pageId="7" pageNumber="640">sides round, poste- rior angle angular (Figs 2A, 3B)</td>
|
||
<td id="974DFF70FFDA0055FD72AC120F55FD1E" box="[721,892,572,671]" gridcol="3" gridrow="3" pageId="7" pageNumber="640">sides almost parallel, posterior angle round (Fig. 3C)</td>
|
||
<td id="974DFF70FFDA0055FC2FAC120810FD1E" box="[908,1081,572,671]" gridcol="4" gridrow="3" pageId="7" pageNumber="640">sides in posterior third slightly sinua- te, posterior angle angular (Fig. 3D)</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAC840810FCC3" box="[121,1081,682,834]" gridrow="4" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAC840D6DFCC3" box="[121,324,682,834]" gridcol="0" gridrow="4" pageId="7" pageNumber="640">pronotum surface punctation</th>
|
||
<td id="974DFF70FFDA0055FEFFAC840E21FCC3" box="[348,520,682,834]" gridcol="1" gridrow="4" pageId="7" pageNumber="640">double, coarse punctures relatively sparsely distributed, concetrated lateral- ly (Fig. 3A)</td>
|
||
<td id="974DFF70FFDA0055FDB5AC840EE9FCC3" box="[534,704,682,834]" gridcol="2" gridrow="4" pageId="7" pageNumber="640">double, coarse punctures relatively densely distributed, concetrated latero- basally (Fig. 3B)</td>
|
||
<td id="974DFF70FFDA0055FD72AC840F55FCC3" box="[721,892,682,834]" gridcol="3" gridrow="4" pageId="7" pageNumber="640">double, coarse punctures relatively densely distributed, concetrated lateral- ly (Fig. 3C)</td>
|
||
<td id="974DFF70FFDA0055FC2FAC840810FCC3" box="[908,1081,682,834]" gridcol="4" gridrow="4" pageId="7" pageNumber="640">double, coarse punctures relatively sparsely distributed on whole surface except for disc (Fig. 3D)</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAD630810FCE1" box="[121,1081,845,864]" gridrow="5" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAD630D6DFCE1" box="[121,324,845,864]" gridcol="0" gridrow="5" pageId="7" pageNumber="640">elytron surface</th>
|
||
<td id="974DFF70FFDA0055FEFFAD630E21FCE1" box="[348,520,845,864]" gridcol="1" gridrow="5" pageId="7" pageNumber="640">shining</td>
|
||
<td id="974DFF70FFDA0055FDB5AD630EE9FCE1" box="[534,704,845,864]" gridcol="2" gridrow="5" pageId="7" pageNumber="640">shining</td>
|
||
<td id="974DFF70FFDA0055FD72AD630F55FCE1" box="[721,892,845,864]" gridcol="3" gridrow="5" pageId="7" pageNumber="640">opaque</td>
|
||
<td id="974DFF70FFDA0055FC2FAD630810FCE1" box="[908,1081,845,864]" gridcol="4" gridrow="5" pageId="7" pageNumber="640">shining</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAD450810FC32" box="[121,1081,875,947]" gridrow="6" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAD450D6DFC32" box="[121,324,875,947]" gridcol="0" gridrow="6" pageId="7" pageNumber="640">elytron macrosetation</th>
|
||
<td id="974DFF70FFDA0055FEFFAD450E21FC32" box="[348,520,875,947]" gridcol="1" gridrow="6" pageId="7" pageNumber="640">entirely glabrous</td>
|
||
<td id="974DFF70FFDA0055FDB5AD450EE9FC32" box="[534,704,875,947]" gridcol="2" gridrow="6" pageId="7" pageNumber="640">minutely macrose- taceous apically</td>
|
||
<td id="974DFF70FFDA0055FD72AD450F55FC32" box="[721,892,875,947]" gridcol="3" gridrow="6" pageId="7" pageNumber="640">distinctly macro- setaceous laterally and in apical half</td>
|
||
<td id="974DFF70FFDA0055FC2FAD450810FC32" box="[908,1081,875,947]" gridcol="4" gridrow="6" pageId="7" pageNumber="640">entirely glabrous</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAD900810FB86" box="[121,1081,958,1031]" gridrow="7" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAD900D6DFB86" box="[121,324,958,1031]" gridcol="0" gridrow="7" pageId="7" pageNumber="640">shape and punctation of sutural elytron interval</th>
|
||
<td id="974DFF70FFDA0055FEFFAD900E21FB86" box="[348,520,958,1031]" gridcol="1" gridrow="7" pageId="7" pageNumber="640">almost flat, with very sparse minute punctures</td>
|
||
<td id="974DFF70FFDA0055FDB5AD900EE9FB86" box="[534,704,958,1031]" gridcol="2" gridrow="7" pageId="7" pageNumber="640">moderately convex, impunctate</td>
|
||
<td id="974DFF70FFDA0055FD72AD900F55FB86" box="[721,892,958,1031]" gridcol="3" gridrow="7" pageId="7" pageNumber="640">almost flat, with sparse minute punctures</td>
|
||
<td id="974DFF70FFDA0055FC2FAD900810FB86" box="[908,1081,958,1031]" gridcol="4" gridrow="7" pageId="7" pageNumber="640">almost flat, with very sparse minute punctures</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAA3F0810FBBE" box="[121,1081,1041,1087]" gridrow="8" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAA3F0D6DFBBE" box="[121,324,1041,1087]" gridcol="0" gridrow="8" pageId="7" pageNumber="640">shape of protibia</th>
|
||
<td id="974DFF70FFDA0055FEFFAA3F0E21FBBE" box="[348,520,1041,1087]" gridcol="1" gridrow="8" pageId="7" pageNumber="640">not narrowed basal- ly (Fig. 3A)</td>
|
||
<td id="974DFF70FFDA0055FDB5AA3F0EE9FBBE" box="[534,704,1041,1087]" gridcol="2" gridrow="8" pageId="7" pageNumber="640">narrowed basally (Fig. 3B)</td>
|
||
<td id="974DFF70FFDA0055FD72AA3F0F55FBBE" box="[721,892,1041,1087]" gridcol="3" gridrow="8" pageId="7" pageNumber="640">not narrowed basal- ly (Fig. 3C)</td>
|
||
<td id="974DFF70FFDA0055FC2FAA3F0810FBBE" box="[908,1081,1041,1087]" gridcol="4" gridrow="8" pageId="7" pageNumber="640">not narrowed basal- ly (Fig. 3D)</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAA640810FB13" box="[121,1081,1098,1170]" gridrow="9" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAA640D6DFB13" box="[121,324,1098,1170]" gridcol="0" gridrow="9" pageId="7" pageNumber="640">shape and direction of terminal protibial spur</th>
|
||
<td id="974DFF70FFDA0055FEFFAA640E21FB13" box="[348,520,1098,1170]" gridcol="1" gridrow="9" pageId="7" pageNumber="640">broadly lanceolate, bent strongly down- ward</td>
|
||
<td id="974DFF70FFDA0055FDB5AA640EE9FB13" box="[534,704,1098,1170]" gridcol="2" gridrow="9" pageId="7" pageNumber="640">simply thickened, not strongly bent downward</td>
|
||
<td id="974DFF70FFDA0055FD72AA640F55FB13" box="[721,892,1098,1170]" gridcol="3" gridrow="9" pageId="7" pageNumber="640">simply thickened, not strongly bent downward</td>
|
||
<td id="974DFF70FFDA0055FC2FAA640810FB13" box="[908,1081,1098,1170]" gridcol="4" gridrow="9" pageId="7" pageNumber="640">broadly lanceolate, not strongly bent downward</td>
|
||
</tr>
|
||
<tr id="D49C960CFFDA0055FFDAAAB30810FA9A" box="[121,1081,1181,1307]" gridrow="10" pageId="7" pageNumber="640">
|
||
<th id="974DFF70FFDA0055FFDAAAB30D6DFA9A" box="[121,324,1181,1307]" gridcol="0" gridrow="10" pageId="7" pageNumber="640">known distribution</th>
|
||
<td id="974DFF70FFDA0055FEFFAAB30E21FA9A" box="[348,520,1181,1307]" gridcol="1" gridrow="10" pageId="7" pageNumber="640">China: Fujian; Thailand</td>
|
||
<td id="974DFF70FFDA0055FDB5AAB30EE9FA9A" box="[534,704,1181,1307]" gridcol="2" gridrow="10" pageId="7" pageNumber="640">India: Rajasthan</td>
|
||
<td id="974DFF70FFDA0055FD72AAB30F55FA9A" box="[721,892,1181,1307]" gridcol="3" gridrow="10" pageId="7" pageNumber="640">India: Karnata- ka, Rajasthan, Uttarakhand; Laos; Myanmar; Thai- land; Vietnam</td>
|
||
<td id="974DFF70FFDA0055FC2FAAB30810FA9A" box="[908,1081,1181,1307]" gridcol="4" gridrow="10" pageId="7" pageNumber="640">N Vietnam</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFDAFFADFF33AB6D0EC2F9BB" blockId="7.[113,1089,1346,1594]" pageId="7" pageNumber="640">
|
||
<emphasis id="58D8485CFFDAFFADFF33AB6D0CE0FADB" box="[144,201,1347,1370]" italics="true" pageId="7" pageNumber="640">Legs.</emphasis>
|
||
Femora shining, glabrous (
|
||
<figureCitation id="F29788CBFFDAFFADFE53AB6C0E6CFADB" box="[496,581,1346,1370]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="7" pageNumber="640">Fig. 2B</figureCitation>
|
||
). Protibia considerably narrow, almost parallel, moreover narrowed in basal quarter, with three triangular teeth externally (
|
||
<figureCitation id="F29788CBFFDAFFADFC2CAB4C082EFAFB" box="[911,1031,1378,1402]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="7" pageNumber="640">Figs 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="F29788CBFFDAFFADFBB1AB4C081AFAFB" box="[1042,1075,1378,1402]" captionStart="Fig" captionStartId="9.[113,144,988,1007]" captionTargetBox="[127,1038,223,954]" captionTargetId="figure-185@9.[113,1088,214,966]" captionTargetPageId="9" captionText="Fig. 3. Head, pronotum and protibia, JJ, dorsal aspect. A – Aphodius (Gilletianus) fukiensis Balthasar, 1952 (China, Fujian: Kuatun; syntype, NMPC), B – A. (G.) rajawatorum sp. nov. (holotype), C – A. (G.) segmentaroides A. Schmidt, 1909 (India, Rajasthan: 50 km W of Agra, Bharatpur, NMPC), D – A. (G.) therondi Balthasar, 1963 (Vietnam: Tonkin, Tam Dao, paratype, NMPC); arrows show diagnostic characters mentioned in the description of the new species and in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5739697" httpUri="https://zenodo.org/record/5739697/files/figure.png" pageId="7" pageNumber="640">3C</figureCitation>
|
||
), ventral edge unarmed, terminal spur simply thickened, not strongly bent downward. External carinae and terminal edge of meso- and metatibia fimbriate with macrosetae strongly unequal in length. Tarses long and thin; basimesotarsite hardly longer than exterior terminal spur of mesotibiae, basimetatarsite approximately one quarter longer than exterior terminal spur of metatibiae and equal in lenght to next three tarsites combined (
|
||
<figureCitation id="F29788CBFFDAFFADFC80A82C0FB6F99B" box="[803,927,1538,1562]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="7" pageNumber="640">Figs 2A–B</figureCitation>
|
||
). Claws feebly arcuate, almost as long as two preceding tarsites combined.
|
||
</paragraph>
|
||
<caption id="3ED3C4C6FFD5FFA2FF34A82A0ED3F9B3" ID-DOI="http://doi.org/10.5281/zenodo.5739693" ID-Zenodo-Dep="5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="8" pageNumber="641" startId="8.[151,182,1540,1559]" targetBox="[195,1074,219,1490]" targetPageId="8">
|
||
<paragraph id="6A13944EFFD5FFA2FF34A82A0ED3F9B3" blockId="8.[151,1127,1540,1586]" pageId="8" pageNumber="641">
|
||
Fig. 2.
|
||
<taxonomicName id="ADACEFCDFFD5FFA2FF75A82B0E26F996" authority="Král & Šípek, 2013" authorityName="Král & Šípek" authorityYear="2013" box="[214,527,1540,1559]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="641" phylum="Arthropoda" rank="species" species="rajawatorum" status="sp. nov." subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD5FFA2FF75A82B0D0EF996" box="[214,295,1541,1559]" italics="true" pageId="8" pageNumber="641">Aphodius</emphasis>
|
||
(
|
||
<emphasis id="58D8485CFFD5FFA2FE97A82B0DBAF996" box="[308,403,1541,1559]" italics="true" pageId="8" pageNumber="641">Gilletianus</emphasis>
|
||
)
|
||
<emphasis id="58D8485CFFD5FFA2FE03A82B0E26F996" box="[416,527,1541,1559]" italics="true" pageId="8" pageNumber="641">rajawatorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="43EBF527FFD5FFA2FDB5A82A0E75F996" box="[534,604,1540,1559]" pageId="8" pageNumber="641" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3ED3C4C6FFD4FFA3FFD2ADF20D55FBDB" ID-DOI="http://doi.org/10.5281/zenodo.5739697" ID-Zenodo-Dep="5739697" httpUri="https://zenodo.org/record/5739697/files/figure.png" pageId="9" pageNumber="642" startId="9.[113,144,988,1007]" targetBox="[127,1038,223,954]" targetPageId="9">
|
||
<paragraph id="6A13944EFFD4FFA3FFD2ADF20D55FBDB" blockId="9.[113,1089,988,1114]" pageId="9" pageNumber="642">
|
||
Fig. 3. Head, pronotum and protibia, JJ, dorsal aspect. A –
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FD35ADF30868FC6E" authority="Balthasar, 1952" authorityName="Balthasar" authorityYear="1952" box="[662,1089,988,1007]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="species" species="fukiensis" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FD35ADF30ECEFC6E" box="[662,743,989,1007]" italics="true" pageId="9" pageNumber="642">Aphodius</emphasis>
|
||
(
|
||
<emphasis id="58D8485CFFD4FFA3FD55ADF30F7CFC6E" box="[758,853,989,1007]" italics="true" pageId="9" pageNumber="642">Gilletianus</emphasis>
|
||
)
|
||
<emphasis id="58D8485CFFD4FFA3FCC6ADF30F99FC6E" box="[869,944,989,1007]" italics="true" pageId="9" pageNumber="642">fukiensis</emphasis>
|
||
Balthasar, 1952
|
||
</taxonomicName>
|
||
(China, Fujian: Kuatun; syntype, NMPC), B –
|
||
<emphasis id="58D8485CFFD4FFA3FE5EADD60E23FB8B" box="[509,522,1016,1034]" italics="true" pageId="9" pageNumber="642">A</emphasis>
|
||
.
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FDB6ADD90E82FB8B" authority="Král & Šípek, 2013" authorityName="Král & Šípek" authorityYear="2013" box="[533,683,1015,1034]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="species" species="rajawatorum" status="sp. nov." subGenus="Gilletianus">
|
||
(
|
||
<emphasis id="58D8485CFFD4FFA3FDBFADD60E02FB8B" box="[540,555,1016,1034]" italics="true" pageId="9" pageNumber="642">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFD4FFA3FD9FADD60E82FB8B" box="[572,683,1016,1034]" italics="true" pageId="9" pageNumber="642">rajawatorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="43EBF527FFD4FFA3FD12ADD90EDAFB8B" box="[689,755,1015,1034]" pageId="9" pageNumber="642" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(holotype), C –
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FCDFADD60D23FBA5" authority="A. Schmidt, 1909" authorityName="A. Schmidt" authorityYear="1909" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="species" species="segmentaroides" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FCDFADD60FA0FB8B" box="[892,905,1016,1034]" italics="true" pageId="9" pageNumber="642">A</emphasis>
|
||
. (
|
||
<emphasis id="58D8485CFFD4FFA3FC38ADD60F83FB8B" box="[923,938,1016,1034]" italics="true" pageId="9" pageNumber="642">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFD4FFA3FC18ADD60869FB8B" box="[955,1088,1016,1034]" italics="true" pageId="9" pageNumber="642">segmentaroides</emphasis>
|
||
A. Schmidt, 1909
|
||
</taxonomicName>
|
||
(India, Rajasthan: 50 km W of Agra, Bharatpur, NMPC), D –
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FC89AA3C0868FBA5" authority="Balthasar, 1963" authorityName="Balthasar" authorityYear="1963" box="[810,1089,1041,1061]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="640" phylum="Arthropoda" rank="species" species="therondi" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FC89AA3C0F1EFBA5" box="[810,823,1042,1060]" italics="true" pageId="9" pageNumber="642">A</emphasis>
|
||
. (
|
||
<emphasis id="58D8485CFFD4FFA3FCE9AA3C0F70FBA5" box="[842,857,1042,1060]" italics="true" pageId="9" pageNumber="642">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFD4FFA3FCCEAA3C0F9CFBA5" box="[877,949,1042,1060]" italics="true" pageId="9" pageNumber="642">therondi</emphasis>
|
||
Balthasar, 1963
|
||
</taxonomicName>
|
||
(Vietnam: Tonkin, Tam Dao, paratype, NMPC); arrows show diagnostic characters mentioned in the description of the new species and in Table 2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6A13944EFFD4FFA3FF33AAB20FEFFB32" blockId="9.[113,1089,1179,1587]" box="[144,966,1179,1203]" pageId="9" pageNumber="642">
|
||
<emphasis id="58D8485CFFD4FFA3FF33AAB20CC4FB32" box="[144,237,1180,1203]" italics="true" pageId="9" pageNumber="642">Ventrites</emphasis>
|
||
shining, impunctate, only with few long erect macrosetae (
|
||
<figureCitation id="F29788CBFFD4FFA3FCC6AAB50F90FB32" box="[869,953,1179,1203]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="9" pageNumber="642">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFD4FFA3FF33AA920E75FB72" blockId="9.[113,1089,1179,1587]" pageId="9" pageNumber="642">
|
||
<emphasis id="58D8485CFFD4FFA3FF33AA920CD1FB52" box="[144,248,1212,1235]" italics="true" pageId="9" pageNumber="642">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="F29788CBFFD4FFA3FEA6AA950D57FB52" box="[261,382,1211,1235]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="9" pageNumber="642">Figs 2C–D</figureCitation>
|
||
). Parameres simple, shorter than phallobasis, with tuft of long, fine macrosetae subapically, almost acute apically.
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFD4FFA3FF33AAD40D48FA92" blockId="9.[113,1089,1179,1587]" box="[144,353,1274,1299]" pageId="9" pageNumber="642">
|
||
<emphasis id="58D8485CFFD4FFA3FF33AAD40CC3FA92" bold="true" box="[144,234,1274,1299]" pageId="9" pageNumber="642">
|
||
<emphasis id="58D8485CFFD4FFA3FF33AAD40CCAFA92" bold="true" box="[144,227,1274,1299]" italics="true" pageId="9" pageNumber="642">Female</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFD4FFA3FFD2AB350FE2FAB2" blockId="9.[113,1089,1179,1587]" box="[113,971,1307,1332]" pageId="9" pageNumber="642">
|
||
<emphasis id="58D8485CFFD4FFA3FFD2AB350CD9FAB2" bold="true" box="[113,240,1307,1331]" pageId="9" pageNumber="642">Variability.</emphasis>
|
||
<typeStatus id="B5172AECFFD4FFA3FF54AB320D76FAB5" box="[247,351,1308,1332]" pageId="9" pageNumber="642" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="F29788CBFFD4FFA3FECDAB350D96FAB2" box="[366,447,1307,1331]" captionStart="Fig" captionStartId="8.[151,182,1540,1559]" captionTargetBox="[195,1074,219,1490]" captionTargetId="figure-10@8.[164,1128,214,1496]" captionTargetPageId="8" captionText="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." figureDoi="http://doi.org/10.5281/zenodo.5739693" httpUri="https://zenodo.org/record/5739693/files/figure.png" pageId="9" pageNumber="642">Fig. 2B</figureCitation>
|
||
) vary only slightly in body length (3.2–3.3 mm).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22B6C7C5FFD4FFA3FFD2AB15081BF9B2" pageId="9" pageNumber="642" type="diagnosis">
|
||
<paragraph id="6A13944EFFD4FFA3FFD2AB15081BF9B2" blockId="9.[113,1089,1179,1587]" pageId="9" pageNumber="642">
|
||
<emphasis id="58D8485CFFD4FFA3FFD2AB150D47FAD2" bold="true" box="[113,366,1339,1363]" pageId="9" pageNumber="642">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FED4AB120DF5FAD2" authorityName="Hellwig" authorityYear="1798" box="[375,476,1340,1363]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="58D8485CFFD4FFA3FED4AB120DF5FAD2" box="[375,476,1340,1363]" italics="true" pageId="9" pageNumber="642">Aphodius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FE45AB150E8FFAD2" authority="Král & Šípek, 2013" authorityName="Král & Šípek" authorityYear="2013" box="[486,678,1339,1363]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="species" species="rajawatorum" status="sp. nov." subGenus="Gilletianus">
|
||
(
|
||
<emphasis id="58D8485CFFD4FFA3FE4DAB120E28FAD2" box="[494,513,1340,1363]" italics="true" pageId="9" pageNumber="642">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFD4FFA3FDB8AB120E8FFAD2" box="[539,678,1340,1363]" italics="true" pageId="9" pageNumber="642">rajawatorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="43EBF527FFD4FFA3FD0CAB120F2CFAD2" box="[687,773,1340,1363]" pageId="9" pageNumber="642" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is classified to the subgenus
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FFD2AB720DBAFAF2" authority="Balthasar, 1933" authorityName="Balthasar" authorityYear="1933" box="[113,403,1371,1395]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="subGenus" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FFD2AB720CCEFAF2" box="[113,231,1372,1395]" italics="true" pageId="9" pageNumber="642">Gilletianus</emphasis>
|
||
<bibRefCitation id="0E3DE9BFFFD4FFA3FF4FAB750DBAFAF2" author="BALTHASAR V." box="[236,403,1371,1395]" pageId="9" pageNumber="642" pagination="139 - 146" refId="ref6470" refString="BALTHASAR V. 1933: Neue Aphodius-Arten aus dem palaarktischen Asien mit Uebersichten der Untergattungen Volinus und Calamosternus. Koleopterologische Rundschau 19: 139 - 146." type="journal article" year="1933">Balthasar, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
, as the subgenus was diagnosed by
|
||
<bibRefCitation id="0E3DE9BFFFD4FFA3FCA1AB720FEAFAF2" author="BALTHASAR V." box="[770,963,1371,1395]" pageId="9" pageNumber="642" pagination="139 - 146" refId="ref6470" refString="BALTHASAR V. 1933: Neue Aphodius-Arten aus dem palaarktischen Asien mit Uebersichten der Untergattungen Volinus und Calamosternus. Koleopterologische Rundschau 19: 139 - 146." type="journal article" year="1933">BALTHASAR (1933)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="0E3DE9BFFFD4FFA3FC6FAB720CDBFA12" author="DELLACASA G. & BORDAT P. & DELLACASA M." pageId="9" pageNumber="642" pagination="1 - 482" refId="ref6909" refString="DELLACASA G., BORDAT P. & DELLACASA M. 2001: A revisional essay of world genus-group taxa of Aphodiinae. Memorie della Societa Entomologica Italiana 79 (2000): 1 - 482." type="journal article" year="2001">DELLACASA et al. (2001)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="0E3DE9BFFFD4FFA3FF58AB520E74FA12" author="DELLACASA M. & DELLACASA G." box="[251,605,1403,1427]" pageId="9" pageNumber="642" pagination="27 - 30" refId="ref7115" refString="DELLACASA M. & DELLACASA G. 2006 b: Aphodius reichei Harold, 1859: neotype designation and placement in the genus Gilletianus Balthasar, 1933. Kogane (Tokyo) 7: 27 - 30." type="journal article" year="2006">DELLACASA & DELLACASA (2006b)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="0E3DE9BFFFD4FFA3FD2DAB550F61FA12" author="OCHI T. & KON M. & KAWAHARA M." box="[654,840,1403,1427]" pageId="9" pageNumber="642" pagination="21 - 32" refId="ref7516" refString="OCHI T., KON M. & KAWAHARA M. 2010: Four new species of the genus Gilletianus from Southeast Asia (Coleoptera, Scarabaeidae). Kogane (Tokyo) 11: 21 - 32." type="journal article" year="2010">OCHI et al. (2010)</bibRefCitation>
|
||
. Within this subgenus it is close to
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FF43ABB20E13FA32" authority="Balthasar, 1952" authorityName="Balthasar" authorityYear="1952" box="[224,570,1435,1459]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="640" phylum="Arthropoda" rank="species" species="fukiensis" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FF43ABB20CD1FA32" box="[224,248,1436,1459]" italics="true" pageId="9" pageNumber="642">A.</emphasis>
|
||
(
|
||
<emphasis id="58D8485CFFD4FFA3FEA5ABB20D30FA32" box="[262,281,1436,1459]" italics="true" pageId="9" pageNumber="642">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFD4FFA3FE8CABB20DA5FA32" box="[303,396,1436,1459]" italics="true" pageId="9" pageNumber="642">fukiensis</emphasis>
|
||
Balthasar, 1952
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FDE7ABB20FDEFA32" authority="A. Schmidt, 1909" authorityName="A. Schmidt" authorityYear="1909" box="[580,1015,1435,1459]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="species" species="segmentaroides" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FDE7ABB20E7DFA32" box="[580,596,1436,1459]" italics="true" pageId="9" pageNumber="642">A</emphasis>
|
||
. (
|
||
<emphasis id="58D8485CFFD4FFA3FDCAABB20EADFA32" box="[617,644,1436,1459]" italics="true" pageId="9" pageNumber="642">G.</emphasis>
|
||
)
|
||
<emphasis id="58D8485CFFD4FFA3FD31ABB20F11FA32" box="[658,824,1436,1459]" italics="true" pageId="9" pageNumber="642">segmentaroides</emphasis>
|
||
A. Schmidt, 1909
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FB89ABB20D81FA52" authority="Balthasar, 1963" authorityName="Balthasar" authorityYear="1963" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="640" phylum="Arthropoda" rank="species" species="therondi" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FB89ABB20813FA32" box="[1066,1082,1436,1459]" italics="true" pageId="9" pageNumber="642">A</emphasis>
|
||
. (
|
||
<emphasis id="58D8485CFFD4FFA3FFD9AB920CA4FA52" box="[122,141,1468,1491]" italics="true" pageId="9" pageNumber="642">G</emphasis>
|
||
.)
|
||
<emphasis id="58D8485CFFD4FFA3FF00AB920CD5FA52" box="[163,252,1468,1491]" italics="true" pageId="9" pageNumber="642">therondi</emphasis>
|
||
Balthasar,
|
||
<quantity id="AD5439ABFFD4FFA3FED0AB950D81FA52" box="[371,424,1467,1491]" metricMagnitude="1" metricUnit="m" metricValue="4.98602" pageId="9" pageNumber="642" unit="in" value="1963.0">1963</quantity>
|
||
</taxonomicName>
|
||
in having distinctly narrow, almost parallel protibia, only very weakly widened in distal part (
|
||
<figureCitation id="F29788CBFFD4FFA3FE16ABF50E3FFA72" box="[437,534,1499,1523]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[151,182,1540,1559]" captionStartId-1="9.[113,144,988,1007]" captionTargetBox-0="[195,1074,219,1490]" captionTargetBox-1="[127,1038,223,954]" captionTargetId-0="figure-10@8.[164,1128,214,1496]" captionTargetId-1="figure-185@9.[113,1088,214,966]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 2. Aphodius (Gilletianus) rajawatorum sp. nov., holotype, male. A – habitus in dorsal aspect; B – habitus in ventral aspect; C–D – aedeagus in dorsal (C) and left lateral (D) aspect." captionText-1="Fig. 3. Head, pronotum and protibia, JJ, dorsal aspect. A – Aphodius (Gilletianus) fukiensis Balthasar, 1952 (China, Fujian: Kuatun; syntype, NMPC), B – A. (G.) rajawatorum sp. nov. (holotype), C – A. (G.) segmentaroides A. Schmidt, 1909 (India, Rajasthan: 50 km W of Agra, Bharatpur, NMPC), D – A. (G.) therondi Balthasar, 1963 (Vietnam: Tonkin, Tam Dao, paratype, NMPC); arrows show diagnostic characters mentioned in the description of the new species and in Table 2." figureDoi-0="http://doi.org/10.5281/zenodo.5739693" figureDoi-1="http://doi.org/10.5281/zenodo.5739697" httpUri-0="https://zenodo.org/record/5739693/files/figure.png" httpUri-1="https://zenodo.org/record/5739697/files/figure.png" pageId="9" pageNumber="642">Figs 2–3</figureCitation>
|
||
). The remaining
|
||
<taxonomicName id="ADACEFCDFFD4FFA3FD6AABF20F69FA72" authorityName="Balthasar" authorityYear="1933" box="[713,832,1500,1523]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="642" phylum="Arthropoda" rank="subGenus" subGenus="Gilletianus">
|
||
<emphasis id="58D8485CFFD4FFA3FD6AABF20F69FA72" box="[713,832,1500,1523]" italics="true" pageId="9" pageNumber="642">Gilletianus</emphasis>
|
||
</taxonomicName>
|
||
species known from the
|
||
<collectingRegion id="A8685AACFFD4FFA3FFD2ABD50D34F992" box="[113,285,1531,1555]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="9" pageNumber="642">Oriental Region</collectingRegion>
|
||
possess protibia more or less distinctly triangularly widened distad. These four species are distinguishable from each other by diagnostic characters given in
|
||
<tableCitation id="272EA1F5FFD4FFA3FC7FA8350807F9B2" box="[988,1070,1563,1587]" captionStart="Table 2" captionStartId="7.[113,160,219,238]" captionText="Table 2. Character matrix distinguishing Gilletianus species close to A. (G.) rajawatorum sp. nov." pageId="9" pageNumber="642">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22B6C7C5FFD7FFA0FF34AEF20F28FEF2" pageId="10" pageNumber="643" type="etymology">
|
||
<paragraph id="6A13944EFFD7FFA0FF34AEF20D93FEB2" blockId="10.[151,1127,219,404]" pageId="10" pageNumber="643">
|
||
<emphasis id="58D8485CFFD7FFA0FF34AEF20D31FF75" bold="true" box="[151,280,220,244]" pageId="10" pageNumber="643">Etymology.</emphasis>
|
||
The new species is named in honour of the Rajawat family from Gola-Ka-Bas (
|
||
<collectingCountry id="12BBD4DEFFD7FFA0FF3CAED50CF2FE92" box="[159,219,251,275]" name="India" pageId="10" pageNumber="643">India</collectingCountry>
|
||
:
|
||
<collectingRegion id="A8685AACFFD7FFA0FF46AED50D79FE92" box="[229,336,251,275]" country="India" name="Rajasthan" pageId="10" pageNumber="643">Rajasthan</collectingRegion>
|
||
) who not only hosted me (PŠ) in their house, but also became our second family far from our homes.
|
||
</paragraph>
|
||
<paragraph id="6A13944EFFD7FFA0FF34AF150F28FEF2" blockId="10.[151,1127,219,404]" pageId="10" pageNumber="643">
|
||
<emphasis id="58D8485CFFD7FFA0FF34AF150D9EFED2" bold="true" box="[151,439,315,339]" pageId="10" pageNumber="643">Collecting circumstances.</emphasis>
|
||
Collected from cow dung pad. For brief characteristics of locality see the Localities visited section, localities No. 5a and 5b.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22B6C7C5FFD7FFA0FF34AF550E12FE15" box="[151,571,379,404]" pageId="10" pageNumber="643" type="distribution">
|
||
<paragraph id="6A13944EFFD7FFA0FF34AF550E12FE15" blockId="10.[151,1127,219,404]" box="[151,571,379,404]" pageId="10" pageNumber="643">
|
||
<emphasis id="58D8485CFFD7FFA0FF34AF550D03FE12" bold="true" box="[151,298,379,403]" pageId="10" pageNumber="643">Distribution.</emphasis>
|
||
<collectingCountry id="12BBD4DEFFD7FFA0FE93AF550D42FE12" box="[304,363,379,403]" name="India" pageId="10" pageNumber="643">India</collectingCountry>
|
||
:
|
||
<collectingRegion id="A8685AACFFD7FFA0FED6AF550DF7FE12" box="[373,478,379,403]" country="India" name="Rajasthan" pageId="10" pageNumber="643">Rajasthan</collectingRegion>
|
||
(
|
||
<figureCitation id="F29788CBFFD7FFA0FE4FAF550E02FE15" box="[492,555,379,404]" captionStart="Fig" captionStartId="2.[858,891,862,881]" captionTargetBox="[149,838,219,959]" captionTargetId="figure-208@2.[149,839,219,959]" captionTargetPageId="2" captionText="Fig. 1. Sketch map of India with marked localities visited (states of Goa, Maharashtra and Rajasthan)." figureDoi="http://doi.org/10.5281/zenodo.5739691" httpUri="https://zenodo.org/record/5739691/files/figure.png" pageId="10" pageNumber="643">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |