treatments-xml/data/DA/7C/87/DA7C87ECFFC4F31885B8F8FAB7269DE4.xml

<document id="46065EE6C270E5B798604F204BE283B3" ID-CLB-Dataset="31392" ID-DOI="10.11646/zootaxa.4083.3.6" ID-GBIF-Dataset="72fb038a-09d2-4777-96ea-ced44b6fcc40" ID-ISSN="1175-5326" ID-Zenodo-Dep="1053864" ID-ZooBank="06CBA73B-1D2F-46B5-9A38-E37F91BE1C43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510925067464" checkinUser="plazi" docAuthor="Shepard, William D. &amp; Barr, Cheryl B." docDate="2016" docId="DA7C87ECFFC4F31885B8F8FAB7269DE4" docLanguage="en" docName="zootaxa.4083.3.6.pdf" docOrigin="Zootaxa 4083 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoelmis guarani Shepard &amp; Barr 2016, sp. n." docType="treatment" docVersion="7" lastPageNumber="429" masterDocId="2645FF94FFC5F314852FFB0EB6559D31" masterDocTitle="Neoelmis guarani Shepard &amp; Barr, a sexually dimorphic new species from Paraguay (Insecta: Coleoptera: Elmidae: Elminae)" masterLastPageNumber="430" masterPageNumber="418" pageNumber="419" updateTime="1698500797002" updateUser="ExternalLinkService">
<mods:mods id="45E2497816D980C49135005F5DA09541" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DDC837AB2FD719CE18446CA1C6D0941E">
<mods:title id="BD3FFC1DF39C391C646C0287FB9595FA">Neoelmis guarani Shepard &amp; Barr, a sexually dimorphic new species from Paraguay (Insecta: Coleoptera: Elmidae: Elminae)</mods:title>
</mods:titleInfo>
<mods:name id="F19FA325932DD719C3351E4CAC520211" type="personal">
<mods:role id="2A9EC2AE132FFBAFE6C0DB910275EBEA">
<mods:roleTerm id="2425EE88ED3BD4B413A7F54948CC5570">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="889276148A0C38044AA404C64945A631">Shepard, William D.</mods:namePart>
</mods:name>
<mods:name id="19DD8A5F61F0AA4B78F9CF234B1FCDDF" type="personal">
<mods:role id="9341EE1A063C6FA851F0369325FD7853">
<mods:roleTerm id="80505B1233BADFA5B2F7D3534F6871BB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39A5DAFB535532235B1D2E402195D9B1">Barr, Cheryl B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="529A6BC87B693F414E0C155DDB9A0C59">text</mods:typeOfResource>
<mods:relatedItem id="06AB69DA198B386871AF88C0045E38CE" type="host">
<mods:titleInfo id="A07064B5E9AC34E5D52C4CA3EAE27482">
<mods:title id="6758351FC4DA88D396DE43B6F7BCBB4A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5C9565C01F22413B8E7252CC64E5B43D">
<mods:date id="F2D641B4C13BCCA3B765B58FBA2843AE">2016</mods:date>
<mods:detail id="F7C8269B67FFCB76D76C59D81DD3EAD3" type="pubDate">
<mods:number id="197B0A2878F2400103940DC9228BB5F2">2016-02-22</mods:number>
</mods:detail>
<mods:detail id="CD217D31E83FD9F076230AD83C49021D" type="volume">
<mods:number id="DFC2D8B8C1F5E57F9E86AE5B6FF1B3D1">4083</mods:number>
</mods:detail>
<mods:detail id="A4E3AB8685777AD273D1FE48220B9015" type="issue">
<mods:number id="8A36F13AA75BE625D99CEF3E228549D7">3</mods:number>
</mods:detail>
<mods:extent id="D2C915F11B31E5C3564F3F0834907DAB" unit="page">
<mods:start id="5F594BE9BBEB46FE711B1D225AEF4638">418</mods:start>
<mods:end id="B4BA42BBC6E39F93CD34C90622654BDD">430</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E473E21962B75D2C7F0DC43EFCFB326F">journal article</mods:classification>
<mods:identifier id="96FF4427587528F527AB347E825F1013" type="CLB-Dataset">31392</mods:identifier>
<mods:identifier id="3A5FCDD3BAD0A978194C984912952895" type="DOI">10.11646/zootaxa.4083.3.6</mods:identifier>
<mods:identifier id="42E1EDDA345C012E27F9FEE6E9A680C0" type="GBIF-Dataset">72fb038a-09d2-4777-96ea-ced44b6fcc40</mods:identifier>
<mods:identifier id="7F49BD491B080471CFC38A992C23FD8D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BCC576A9344C05D709BB227CD592AB46" type="Zenodo-Dep">1053864</mods:identifier>
<mods:identifier id="5A5B7B394AD044077FE9DF5DBB2678E0" type="ZooBank">06CBA73B-1D2F-46B5-9A38-E37F91BE1C43</mods:identifier>
</mods:mods>
<treatment id="DA7C87ECFFC4F31885B8F8FAB7269DE4" ID-DOI="http://doi.org/10.5281/zenodo.6082103" ID-GBIF-Taxon="137453366" ID-Zenodo-Dep="6082103" LSID="urn:lsid:plazi:treatment:DA7C87ECFFC4F31885B8F8FAB7269DE4" httpUri="http://treatment.plazi.org/id/DA7C87ECFFC4F31885B8F8FAB7269DE4" lastPageId="12" lastPageNumber="429" pageId="1" pageNumber="419">
<subSubSection id="1ACF6571FFC4F31585B8F8FAB7589901" pageId="1" pageNumber="419" type="nomenclature">
<paragraph id="526A36FAFFC4F31585B8F8FAB7ED993E" blockId="1.[151,440,1012,1072]" box="[151,440,1012,1039]" pageId="1" pageNumber="419">
<heading id="09228196FFC4F31585B8F8FAB7ED993E" bold="true" box="[151,440,1012,1039]" fontSize="11" level="1" pageId="1" pageNumber="419" reason="1">
<emphasis id="60A1EAE8FFC4F31585B8F8FAB7ED993E" bold="true" box="[151,440,1012,1039]" pageId="1" pageNumber="419">
<taxonomicName id="95D54D79FFC4F31585B8F8FAB73B993F" ID-CoL="46GSB" authority="Shepard &amp; Barr, 2016" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[151,366,1012,1038]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="419" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="60A1EAE8FFC4F31585B8F8FAB73B993F" bold="true" box="[151,366,1012,1038]" italics="true" pageId="1" pageNumber="419">Neoelmis guarani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7B925793FFC4F3158459F8FBB7ED993E" box="[374,440,1013,1039]" pageId="1" pageNumber="419" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="526A36FAFFC4F31585B8FF19B7589901" blockId="1.[151,440,1012,1072]" box="[151,269,1047,1072]" pageId="1" pageNumber="419">
<figureCitation id="CAEE2A7FFFC4F31585B8FF19B7589901" box="[151,269,1047,1072]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 3 – 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStart-6="FIGURE 9" captionStart-7="FIGURE 10" captionStart-8="FIGURE 11" captionStart-9="FIGURE 12" captionStartId-0="2.[151,250,1973,1995]" captionStartId-1="3.[151,250,1973,1995]" captionStartId-2="4.[151,264,1934,1957]" captionStartId-3="5.[151,250,1311,1333]" captionStartId-4="6.[151,250,1948,1970]" captionStartId-5="7.[151,250,890,912]" captionStartId-6="8.[151,250,1886,1908]" captionStartId-7="9.[151,250,1147,1169]" captionStartId-8="10.[151,250,1928,1950]" captionStartId-9="11.[151,250,1178,1200]" captionTargetBox-0="[181,1334,264,1941]" captionTargetBox-1="[247,1333,193,1916]" captionTargetBox-2="[257,1336,276,1839]" captionTargetBox-3="[246,1227,194,1271]" captionTargetBox-4="[151,1436,193,1927]" captionTargetBox-5="[151,1436,193,868]" captionTargetBox-6="[151,1436,151,1864]" captionTargetBox-7="[151,1436,161,1125]" captionTargetBox-8="[151,1436,193,1906]" captionTargetBox-9="[151,1436,193,1157]" captionTargetId-0="figure@2.[171,1415,193,1943]" captionTargetId-1="figure@3.[247,1334,193,1916]" captionTargetId-2="figure@4.[180,1406,193,2007]" captionTargetId-3="figure@5.[230,1356,175,1306]" captionTargetId-4="figure@6.[151,1436,193,1927]" captionTargetId-5="figure@7.[151,1436,193,868]" captionTargetId-6="figure@8.[151,1436,151,1864]" captionTargetId-7="figure@9.[151,1436,161,1125]" captionTargetId-8="figure@10.[151,1436,193,1906]" captionTargetId-9="figure@11.[151,1436,193,1157]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." captionText-1="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." captionText-2="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." captionText-3="FIGURE 6. Neoelmis guarani, aedeagus, A—dorsal view, B—lateral view. Scale bar = 0.1 mm." captionText-4="FIGURE 7. Neoelmis guarani, female dorsal habitus; length 2.3 mm. Arrow indicates paired elytral protuberances." captionText-5="FIGURE 8. Neoelmis guarani, female lateral habitus; length 2.3 mm." captionText-6="FIGURE 9. Arroyo Naranjo, type locality of Neoelmis guarani." captionText-7="FIGURE 10. Arroyo Naranjo, type locality of Neoelmis guarani." captionText-8="FIGURE 11. Arroyo Mborebí, Neoelmis guarani collection site." captionText-9="FIGURE 12. Capilla Cue, Neoelmis guarani collection site." httpUri-0="https://zenodo.org/record/1053866/files/figure.png" httpUri-1="https://zenodo.org/record/1053868/files/figure.png" httpUri-2="https://zenodo.org/record/1053870/files/figure.png" httpUri-3="https://zenodo.org/record/1053872/files/figure.png" httpUri-4="https://zenodo.org/record/1053874/files/figure.png" httpUri-5="https://zenodo.org/record/1053876/files/figure.png" httpUri-6="https://zenodo.org/record/1053878/files/figure.png" httpUri-7="https://zenodo.org/record/1053880/files/figure.png" httpUri-8="https://zenodo.org/record/1053882/files/figure.png" httpUri-9="https://zenodo.org/record/1053884/files/figure.png" pageId="1" pageNumber="419">Figs. 1–12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1ACF6571FFC4F31385B8FF51B28E9A20" lastPageId="7" lastPageNumber="425" pageId="1" pageNumber="419" type="materials_examined">
<paragraph id="526A36FAFFC4F31585B8FF51B53499AC" blockId="1.[151,1437,1119,2009]" pageId="1" pageNumber="419">
<emphasis id="60A1EAE8FFC4F31585B8FF51B7CF9949" bold="true" box="[151,410,1119,1144]" pageId="1" pageNumber="419">
<typeStatus id="8D6E8858FFC4F31585B8FF51B7539949" box="[151,262,1119,1144]" pageId="1" pageNumber="419" type="holotype">Holotype</typeStatus>
, adult male.
</emphasis>
Body slender, 
<quantity id="952D9B1FFFC4F3158769FF6EB4C89946" box="[582,669,1120,1144]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="419" unit="mm" value="2.1">2.1 mm</quantity>
long (pronotum + elytra), 
<quantity id="952D9B1FFFC4F31586E1FF6EB2709946" box="[974,1061,1120,1144]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="419" unit="mm" value="0.7">0.7 mm</quantity>
wide at widest point; body color testaceous, appendages (antennae, palpi, legs) paler (
<figureCitation id="CAEE2A7FFFC4F31587C8FF8AB50699AC" box="[743,851,1156,1181]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 3 – 5" captionStartId-0="2.[151,250,1973,1995]" captionStartId-1="3.[151,250,1973,1995]" captionStartId-2="4.[151,264,1934,1957]" captionTargetBox-0="[181,1334,264,1941]" captionTargetBox-1="[247,1333,193,1916]" captionTargetBox-2="[257,1336,276,1839]" captionTargetId-0="figure@2.[171,1415,193,1943]" captionTargetId-1="figure@3.[247,1334,193,1916]" captionTargetId-2="figure@4.[180,1406,193,2007]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." captionText-1="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." captionText-2="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri-0="https://zenodo.org/record/1053866/files/figure.png" httpUri-1="https://zenodo.org/record/1053868/files/figure.png" httpUri-2="https://zenodo.org/record/1053870/files/figure.png" pageId="1" pageNumber="419">Figs. 1–5</figureCitation>
).
</paragraph>
<paragraph id="526A36FAFFC4F31585E8FFA9B5919844" blockId="1.[151,1437,1119,2009]" pageId="1" pageNumber="419">
Head with frons moderately setose, setae long and pale; genae covered with fine, adpressed, plastron setae; eyes large, protuberant; antennal ridges produced, especially anteriorly. Antennae filiform, each with 11 segments; antennomeres 1–2 twice as long as wide, antennomeres 3–10 more than twice as long as wide; antennomere 11 clavate, slightly curved, longer than antennomeres 9+10 (
<figureCitation id="CAEE2A7FFFC4F315866FFE1AB5C7981D" box="[832,914,1300,1325]" captionStart="FIGURE 1" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[181,1334,264,1941]" captionTargetId="figure@2.[171,1415,193,1943]" captionTargetPageId="2" captionText="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." httpUri="https://zenodo.org/record/1053866/files/figure.png" pageId="1" pageNumber="419">Figs. 1</figureCitation>
, 
<figureCitation id="CAEE2A7FFFC4F3158689FE1AB5E2981D" box="[934,951,1300,1324]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="1" pageNumber="419">2</figureCitation>
). Clypeus setose, transverse, apex feebly emarginate. Labrum transverse, apex arcuate; surface shiny with few setae; apicolateral margins with fringe of pale, dense setae. Maxillary and labial palpi each with three palpomeres.
</paragraph>
<paragraph id="526A36FAFFC4F31585E8FE8EB35D9BA5" blockId="1.[151,1437,1119,2009]" pageId="1" pageNumber="419">
Pronotum (
<figureCitation id="CAEE2A7FFFC4F3158469FE71B7D298A9" box="[326,391,1407,1432]" captionStart="FIGURE 1" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[181,1334,264,1941]" captionTargetId="figure@2.[171,1415,193,1943]" captionTargetPageId="2" captionText="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." httpUri="https://zenodo.org/record/1053866/files/figure.png" pageId="1" pageNumber="419">Fig. 1</figureCitation>
) elongate, 
<quantity id="952D9B1FFFC4F3158726FE71B43598A6" box="[521,608,1407,1432]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="419" unit="mm" value="0.6">0.6 mm</quantity>
long, 
<quantity id="952D9B1FFFC4F3158787FE71B4AA98A6" box="[680,767,1407,1432]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="419" unit="mm" value="0.5">0.5 mm</quantity>
wide at widest point about 1/3 distance from base; surface shiny, smooth, and sparsely clothed with pale, recumbent setae except for central disc where finely granulate and more densely setose; lateral margins bisinuate and irregularly crenulate; apicolateral angles acute, produced; posterior border bisinuate, notched medially to receive scutellum; prominent, wide transverse sulcus at 1/3 distance from apex, deeper fovea at midline with longitudinal groove which tapers posteriorly; two prominent, sinuate, sublateral carinae; broad, shallow depression between each sublateral carina and lateral margin, with a large fovea ventrally adjacent to transverse sulcus. Hypomeron with shallow depression on each side at basal 1/3 and a large fovea ventrally adjacent to fovea between lateral margin and sublateral carina; without plastron setae.
</paragraph>
<paragraph id="526A36FAFFC4F31585E8FD91B5719B86" blockId="1.[151,1437,1119,2009]" box="[199,804,1695,1720]" pageId="1" pageNumber="419">Scutellum subovoid, longer than wide, feebly convex.</paragraph>
<paragraph id="526A36FAFFC4F31585E8FDCAB5C79A5C" blockId="1.[151,1437,1119,2009]" pageId="1" pageNumber="419">
Elytra (
<figureCitation id="CAEE2A7FFFC4F3158430FDCAB7369BED" box="[287,355,1732,1757]" captionStart="FIGURE 1" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[181,1334,264,1941]" captionTargetId="figure@2.[171,1415,193,1943]" captionTargetPageId="2" captionText="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." httpUri="https://zenodo.org/record/1053866/files/figure.png" pageId="1" pageNumber="419">Fig. 1</figureCitation>
) elongate, 
<quantity id="952D9B1FFFC4F31584C4FDCAB4109BED" box="[491,581,1732,1757]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="419" unit="mm" value="1.5">1.5 mm</quantity>
long, 
<quantity id="952D9B1FFFC4F31587BBFDCAB4BB9BED" box="[660,750,1732,1756]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="419" unit="mm" value="0.7">0.7 mm</quantity>
wide at widest point 1/3–1/2 distance from elytral apices; shallow linear punctures and setae present. Each elytron with a short, prominent, basal carina on third interval separating shallow lateral and medial depressions; two long sublateral carinae extending posteriorly from humeral angle, inner one extending to apical 1/3, outer one extending nearly to apex. Epipleuron with plastron setae, notched just before apex to receive tooth from abdominal ventrite 5.
</paragraph>
<paragraph id="526A36FAFFC4F31185E8FC77B5BC9B7C" blockId="1.[151,1437,1119,2009]" lastBlockId="5.[151,1437,1372,2009]" lastPageId="5" lastPageNumber="423" pageId="1" pageNumber="419">
Prosternum (
<figureCitation id="CAEE2A7FFFC4F3158471FC79B7FD9AA1" box="[350,424,1911,1936]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="1" pageNumber="419">Fig. 2</figureCitation>
) longer than wide, with flat, evenly and lightly setose, shiny disc bordered by two longitudinal carinae 3/4 length of prosternum, fading out anteriorly; episternum clothed with plastron setae; prosternal process wide, apex extending posterior to procoxae, margins raised and rimmed with narrow sulcus between. Mesoventrite (
<figureCitation id="CAEE2A7FFFC0F311848AFE52B7BD9844" box="[421,488,1372,1397]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="5" pageNumber="423">Fig. 2</figureCitation>
) shorter than wide; with a pair of large, posterolateral, ventrally projecting processes (
<figureCitation id="CAEE2A7FFFC0F31185B0FE8EB6BD98A8" box="[159,232,1408,1433]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="5" pageNumber="423">Fig. 2</figureCitation>
, arrow) adjacent to and extending below mesocoxae; processes with ventromedially oriented, flattened surfaces; area between processes excavated; mesanepisternum with plastron setae. Metaventrite (
<figureCitation id="CAEE2A7FFFC0F31181D9FEAAB369988C" box="[1270,1340,1444,1469]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="5" pageNumber="423">Fig. 2</figureCitation>
) shorter than prosternum, longer than mesosternum; disc flat, shiny, sparsely setose, with distinct discrimen; with two sublateral carinae extending from mesocoxae to metacoxae and small, carinate tubercle near each posterolateral margin; plastron setae present laterad to carinae and on metanepisternum. Pro- and mesocoxae globular; metacoxae transverse. Pro- and mesotrochanters of similar size, metatrochanters larger.
</paragraph>
<caption id="06AA6672FFC7F31685B8FCBBB5129AFB" box="[151,839,1972,1995]" httpUri="https://zenodo.org/record/1053866/files/figure.png" pageId="2" pageNumber="420" targetBox="[181,1334,264,1941]" targetPageId="2">
<paragraph id="526A36FAFFC7F31685B8FCBBB5129AFB" blockId="2.[151,839,1972,1995]" box="[151,839,1972,1995]" pageId="2" pageNumber="420">
<emphasis id="60A1EAE8FFC7F31685B8FCBBB7469AFB" bold="true" box="[151,275,1973,1995]" pageId="2" pageNumber="420">FIGURE 1.</emphasis>
<taxonomicName id="95D54D79FFC7F3168436FCBAB7989AFA" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[281,461,1972,1995]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="420" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFC7F3168436FCBAB7989AFA" box="[281,461,1972,1995]" italics="true" pageId="2" pageNumber="420">Neoelmis guarani</emphasis>
</taxonomicName>
, male dorsal habitus; length 2.1 mm.
</paragraph>
</caption>
<caption id="06AA6672FFC6F31785B8FCBBB73A9ADB" httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="3" pageNumber="421" targetBox="[247,1333,193,1916]" targetPageId="3">
<paragraph id="526A36FAFFC6F31785B8FCBBB73A9ADB" blockId="3.[151,1435,1973,2026]" pageId="3" pageNumber="421">
<emphasis id="60A1EAE8FFC6F31785B8FCBBB7469AFB" bold="true" box="[151,275,1973,1995]" pageId="3" pageNumber="421">FIGURE 2.</emphasis>
<taxonomicName id="95D54D79FFC6F3178435FCBBB79A9AFA" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[282,463,1973,1995]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="421" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFC6F3178435FCBBB79A9AFA" box="[282,463,1973,1995]" italics="true" pageId="3" pageNumber="421">Neoelmis guarani</emphasis>
</taxonomicName>
, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2.
</paragraph>
</caption>
<caption id="06AA6672FFC1F31085B8FC80B4CD9AF2" httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="4" pageNumber="422" targetBox="[257,1336,276,1839]" targetPageId="4">
<paragraph id="526A36FAFFC1F31085B8FC80B4CD9AF2" blockId="4.[151,1436,1934,1987]" pageId="4" pageNumber="422">
<emphasis id="60A1EAE8FFC1F31085B8FC80B76E9A95" bold="true" box="[151,315,1934,1957]" pageId="4" pageNumber="422">FIGURES 3–5.</emphasis>
<taxonomicName id="95D54D79FFC1F3108466FC81B7AA9A94" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[329,511,1935,1957]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="422" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFC1F3108466FC81B7AA9A94" box="[329,511,1935,1957]" italics="true" pageId="4" pageNumber="422">Neoelmis guarani</emphasis>
</taxonomicName>
, male. 
<emphasis id="60A1EAE8FFC1F310877CFC81B4309A95" bold="true" box="[595,613,1935,1956]" pageId="4" pageNumber="422">3.</emphasis>
Left prothoracic leg, femur, dorsal view. 
<emphasis id="60A1EAE8FFC1F3108137FC81B27E9A95" bold="true" box="[1048,1067,1935,1956]" pageId="4" pageNumber="422">4.</emphasis>
Left prothoracic leg, posteroventral view. 
<emphasis id="60A1EAE8FFC1F31085FCFCA0B6B39AF2" bold="true" box="[211,230,1966,1987]" pageId="4" pageNumber="422">5.</emphasis>
Left mesothoracic leg, anteroventral view.
</paragraph>
</caption>
<caption id="06AA6672FFC0F31185B8FE11B23D9804" box="[151,1128,1311,1333]" httpUri="https://zenodo.org/record/1053872/files/figure.png" pageId="5" pageNumber="423" targetBox="[246,1227,194,1271]" targetPageId="5">
<paragraph id="526A36FAFFC0F31185B8FE11B23D9804" blockId="5.[151,1128,1311,1333]" box="[151,1128,1311,1333]" pageId="5" pageNumber="423">
<emphasis id="60A1EAE8FFC0F31185B8FE11B7479805" bold="true" box="[151,274,1311,1333]" pageId="5" pageNumber="423">FIGURE 6.</emphasis>
<taxonomicName id="95D54D79FFC0F3118436FE11B7989804" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[281,461,1311,1333]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="423" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFC0F3118436FE11B7989804" box="[281,461,1311,1333]" italics="true" pageId="5" pageNumber="423">Neoelmis guarani</emphasis>
</taxonomicName>
, aedeagus, A—dorsal view, B—lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="526A36FAFFC0F31385E8FD56B7A79971" blockId="5.[151,1437,1372,2009]" lastBlockId="7.[151,1437,956,1809]" lastPageId="7" lastPageNumber="425" pageId="5" pageNumber="423">
Prothoracic leg (
<figureCitation id="CAEE2A7FFFC0F31184A3FD56B7A99B40" box="[396,508,1624,1649]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 3 – 5" captionStartId-0="2.[151,250,1973,1995]" captionStartId-1="3.[151,250,1973,1995]" captionStartId-2="4.[151,264,1934,1957]" captionTargetBox-0="[181,1334,264,1941]" captionTargetBox-1="[247,1333,193,1916]" captionTargetBox-2="[257,1336,276,1839]" captionTargetId-0="figure@2.[171,1415,193,1943]" captionTargetId-1="figure@3.[247,1334,193,1916]" captionTargetId-2="figure@4.[180,1406,193,2007]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." captionText-1="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." captionText-2="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri-0="https://zenodo.org/record/1053866/files/figure.png" httpUri-1="https://zenodo.org/record/1053868/files/figure.png" httpUri-2="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">Figs. 1–4</figureCitation>
) with profemur abruptly and deeply excavated on both anterior and posterior surfaces 1/3 distance from apex (
<figureCitation id="CAEE2A7FFFC0F3118727FD72B41E9BA4" box="[520,587,1660,1685]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[257,1336,276,1839]" captionTargetId="figure@4.[180,1406,193,2007]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">Fig. 3</figureCitation>
), excavations lined with short, stiff setae (
<figureCitation id="CAEE2A7FFFC0F311810AFD72B23F9BA4" box="[1061,1130,1660,1685]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[257,1336,276,1839]" captionTargetId="figure@4.[180,1406,193,2007]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">Fig. 4</figureCitation>
); anterior surface bearing a large, blunt, distally directed tooth at inner edge of excavation; plastron setae present except in excavations. Protibia (
<figureCitation id="CAEE2A7FFFC0F311842DFDCAB71C9BEC" box="[258,329,1732,1757]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[257,1336,276,1839]" captionTargetId="figure@4.[180,1406,193,2007]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">Fig. 4</figureCitation>
) slender, wider apically; posterior face with long setae present basally and apically and a deep excavation near apex; cleaning fringe of long, dense, pale setae on anterior surface. Protarsus (
<figureCitation id="CAEE2A7FFFC0F31181D1FDE6B3139A30" box="[1278,1350,1768,1793]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[257,1336,276,1839]" captionTargetId="figure@4.[180,1406,193,2007]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">Fig. 4</figureCitation>
) shiny, tarsomeres 1–4 with long setae ventrally, tarsomere 5 with a pair of long, projecting setae at apex; tarsomere 5 longer than tarsomeres 1–4 combined; claws simple. Mesothoracic leg (
<figureCitation id="CAEE2A7FFFC0F31186F3FC3EB2789A78" box="[988,1069,1840,1865]" captionStart="FIGURE 1" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[181,1334,264,1941]" captionTargetId="figure@2.[171,1415,193,1943]" captionTargetPageId="2" captionText="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." httpUri="https://zenodo.org/record/1053866/files/figure.png" pageId="5" pageNumber="423">Figs. 1</figureCitation>
, 
<figureCitation id="CAEE2A7FFFC0F3118111FC3FB21B9A78" box="[1086,1102,1841,1865]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="5" pageNumber="423">2</figureCitation>
, 
<figureCitation id="CAEE2A7FFFC0F3118171FC3EB2399A78" box="[1118,1132,1840,1865]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[257,1336,276,1839]" captionTargetId="figure@4.[180,1406,193,2007]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">5</figureCitation>
) with mesofemur (
<figureCitation id="CAEE2A7FFFC0F3118062FC3EB3C69A78" box="[1357,1427,1840,1865]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[257,1336,276,1839]" captionTargetId="figure@4.[180,1406,193,2007]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">Fig. 5</figureCitation>
) widened medially; a deep, longitudinal groove present on ventral surface to receive tibia when folded, posterior margin of groove with numerous, long, curved setae; plastron setae present except inside of groove. Mesotibia (
<figureCitation id="CAEE2A7FFFC0F31185B0FC92B6B49A84" box="[159,225,1948,1973]" captionStart="FIGURES 3 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[257,1336,276,1839]" captionTargetId="figure@4.[180,1406,193,2007]" captionTargetPageId="4" captionText="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri="https://zenodo.org/record/1053870/files/figure.png" pageId="5" pageNumber="423">Fig. 5</figureCitation>
) in lateral view with ventrally directed expansion at middle and shallow excavation on anterior face about 1/ 3 distance from apex; with anterior and posterior cleaning fringes (
<figureCitation id="CAEE2A7FFFC0F31186ACFCCEB5919AE8" box="[899,964,1984,2009]" captionStart="FIGURE 1" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[181,1334,264,1941]" captionTargetId="figure@2.[171,1415,193,1943]" captionTargetPageId="2" captionText="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." httpUri="https://zenodo.org/record/1053866/files/figure.png" pageId="5" pageNumber="423">Fig. 1</figureCitation>
) having long, tuft-like setae that extend beyond tibial apex. Mesotarsus similar to protarsus; right mesotarsus missing tarsomeres 3–5 due to breakage. Metathoracic leg (
<figureCitation id="CAEE2A7FFFC2F3138447F8EEB7E29EC8" box="[360,439,992,1017]" captionStart="FIGURE 1" captionStartId="2.[151,250,1973,1995]" captionTargetBox="[181,1334,264,1941]" captionTargetId="figure@2.[171,1415,193,1943]" captionTargetPageId="2" captionText="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." httpUri="https://zenodo.org/record/1053866/files/figure.png" pageId="7" pageNumber="425">Figs. 1</figureCitation>
, 
<figureCitation id="CAEE2A7FFFC2F31384E7F8EFB7829EC8" box="[456,471,993,1017]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="7" pageNumber="425">2</figureCitation>
) with metafemur unmodified. Metatibia with cleaning fringe on posterior surface at central 1/3; inner surface flat at apical 1/4, glabrous and shiny with a small peg near ventral margin. Metatarsus similar to pro- and mesotarsus.
</paragraph>
<caption id="06AA6672FFC3F31285B8FC92B37F9A83" box="[151,1322,1948,1970]" httpUri="https://zenodo.org/record/1053874/files/figure.png" pageId="6" pageNumber="424" targetBox="[151,1436,193,1927]" targetPageId="6">
<paragraph id="526A36FAFFC3F31285B8FC92B37F9A83" blockId="6.[151,1322,1948,1970]" box="[151,1322,1948,1970]" pageId="6" pageNumber="424">
<emphasis id="60A1EAE8FFC3F31285B8FC92B7479A83" bold="true" box="[151,274,1948,1970]" pageId="6" pageNumber="424">FIGURE 7.</emphasis>
<taxonomicName id="95D54D79FFC3F3128436FC92B7989A83" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[281,461,1948,1970]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="424" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFC3F3128436FC92B7989A83" box="[281,461,1948,1970]" italics="true" pageId="6" pageNumber="424">Neoelmis guarani</emphasis>
</taxonomicName>
, female dorsal habitus; length 2.3 mm. Arrow indicates paired elytral protuberances.
</paragraph>
</caption>
<caption id="06AA6672FFC2F31385B8F874B5099EA1" box="[151,860,890,912]" httpUri="https://zenodo.org/record/1053876/files/figure.png" pageId="7" pageNumber="425" targetBox="[151,1436,193,868]" targetPageId="7">
<paragraph id="526A36FAFFC2F31385B8F874B5099EA1" blockId="7.[151,860,890,912]" box="[151,860,890,912]" pageId="7" pageNumber="425">
<emphasis id="60A1EAE8FFC2F31385B8F874B7479EBE" bold="true" box="[151,274,890,912]" pageId="7" pageNumber="425">FIGURE 8.</emphasis>
<taxonomicName id="95D54D79FFC2F3138436F874B7989EA1" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[281,461,890,912]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="425" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFC2F3138436F874B7989EA1" box="[281,461,890,912]" italics="true" pageId="7" pageNumber="425">Neoelmis guarani</emphasis>
</taxonomicName>
, female lateral habitus; length 2.3 mm.
</paragraph>
</caption>
<paragraph id="526A36FAFFC2F31385E8FF42B5529850" blockId="7.[151,1437,956,1809]" pageId="7" pageNumber="425">
Abdomen (
<figureCitation id="CAEE2A7FFFC2F3138469FF42B7DF9954" box="[326,394,1100,1125]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="7" pageNumber="425">Fig. 2</figureCitation>
) with five ventrites; ventrites 1–4 decreasing in length posteriorly, ventrite 5 longer than all but ventrite 1. Ventrite 1 longest, with basomedial depression bordered by two short carinae which extend from posterior margin of metacoxae nearly to posterior margin of ventrite; surface shiny and sparsely setose between carinae, plastron setae present laterad to carinae. Ventrite 2 with a pair of processes (
<figureCitation id="CAEE2A7FFFC2F3138152FFB6B29099E0" box="[1149,1221,1208,1233]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1995]" captionTargetBox="[247,1333,193,1916]" captionTargetId="figure@3.[247,1334,193,1916]" captionTargetPageId="3" captionText="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." httpUri="https://zenodo.org/record/1053868/files/figure.png" pageId="7" pageNumber="425">Fig. 2</figureCitation>
, arrow) projecting ventrally from anteromedial border, size of each process less than half that of each mesoventral process; surface between processes shiny, plastron setae present elsewhere. Ventrites 3–5 covered with plastron setae; ventrite 5 with a dorsally projecting tooth on each lateral margin which serves to link with a notch in the epipleuron. Ventrite 5 removed from abdomen and mounted on a card point.
</paragraph>
<paragraph id="526A36FAFFC2F31385E8FE62B3339B24" blockId="7.[151,1437,956,1809]" pageId="7" pageNumber="425">
Aedeagus (
<figureCitation id="CAEE2A7FFFC2F3138466FE62B7C498B4" box="[329,401,1388,1413]" captionStart="FIGURE 6" captionStartId="5.[151,250,1311,1333]" captionTargetBox="[246,1227,194,1271]" captionTargetId="figure@5.[230,1356,175,1306]" captionTargetPageId="5" captionText="FIGURE 6. Neoelmis guarani, aedeagus, A—dorsal view, B—lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/1053872/files/figure.png" pageId="7" pageNumber="425">Fig. 6</figureCitation>
) in dorsal view with phallobase twice as long as parameres, widely open in basal half; parameres widely open at apical 2/3 exposing penis; penis stout, sides faintly arcuate at basal 1/2, then moderately constricted and nearly parallel-sided to apical 1/8, thereafter evenly converging to narrowly rounded apex which extends slightly past paramere apices. Penis in ventral view with fibula at middle. Aedeagus in lateral view with apices of parameres and penis acutely pointed. Aedeagus removed from abdomen and placed in genitalia vial.
</paragraph>
<paragraph id="526A36FAFFC2F31385E8FD2EB28E9A20" blockId="7.[151,1437,956,1809]" pageId="7" pageNumber="425">
<emphasis id="60A1EAE8FFC2F31385E8FD2EB7809B08" bold="true" box="[199,469,1568,1593]" pageId="7" pageNumber="425">
<typeStatus id="8D6E8858FFC2F31385E8FD2EB77B9B08" box="[199,302,1568,1593]" pageId="7" pageNumber="425" type="allotype">Allotype</typeStatus>
, adult female.
</emphasis>
Body slender, 
<quantity id="952D9B1FFFC2F3138752FD2FB4869B09" box="[637,723,1569,1593]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="425" unit="mm" value="2.3">2.3 mm</quantity>
long (pronotum + elytra), 
<quantity id="952D9B1FFFC2F31386D3FD2FB2079B09" box="[1020,1106,1569,1593]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="425" unit="mm" value="0.8">0.8 mm</quantity>
wide at widest point about 1/ 3 distance from elytral apices (
<figureCitation id="CAEE2A7FFFC2F31384DEFD4AB4609B6C" box="[497,565,1604,1629]" captionStart="FIGURE 7" captionStartId="6.[151,250,1948,1970]" captionTargetBox="[151,1436,193,1927]" captionTargetId="figure@6.[151,1436,193,1927]" captionTargetPageId="6" captionText="FIGURE 7. Neoelmis guarani, female dorsal habitus; length 2.3 mm. Arrow indicates paired elytral protuberances." httpUri="https://zenodo.org/record/1053874/files/figure.png" pageId="7" pageNumber="425">Fig. 7</figureCitation>
). Slightly larger but generally similar to male except as follows: Each elytron (
<figureCitation id="CAEE2A7FFFC2F31385B0FD66B6A49BB1" box="[159,241,1640,1665]" captionStart="FIGURE 7" captionStartId="6.[151,250,1948,1970]" captionTargetBox="[151,1436,193,1927]" captionTargetId="figure@6.[151,1436,193,1927]" captionTargetPageId="6" captionText="FIGURE 7. Neoelmis guarani, female dorsal habitus; length 2.3 mm. Arrow indicates paired elytral protuberances." httpUri="https://zenodo.org/record/1053874/files/figure.png" pageId="7" pageNumber="425">Figs. 7</figureCitation>
, 
<figureCitation id="CAEE2A7FFFC2F31385D0FD67B7589BB0" box="[255,269,1641,1665]" captionStart="FIGURE 8" captionStartId="7.[151,250,890,912]" captionTargetBox="[151,1436,193,868]" captionTargetId="figure@7.[151,1436,193,868]" captionTargetPageId="7" captionText="FIGURE 8. Neoelmis guarani, female lateral habitus; length 2.3 mm." httpUri="https://zenodo.org/record/1053876/files/figure.png" pageId="7" pageNumber="425">8</figureCitation>
) with a dorsally projecting protuberance in the third interval at about 1/3 distance from apex (
<figureCitation id="CAEE2A7FFFC2F313807FFD66B3C29BB1" box="[1360,1431,1640,1665]" captionStart="FIGURE 7" captionStartId="6.[151,250,1948,1970]" captionTargetBox="[151,1436,193,1927]" captionTargetId="figure@6.[151,1436,193,1927]" captionTargetPageId="6" captionText="FIGURE 7. Neoelmis guarani, female dorsal habitus; length 2.3 mm. Arrow indicates paired elytral protuberances." httpUri="https://zenodo.org/record/1053874/files/figure.png" pageId="7" pageNumber="425">Fig. 7</figureCitation>
, arrow), pointed and subtriangular in lateral view (
<figureCitation id="CAEE2A7FFFC2F31387D1FD82B51F9B94" box="[766,842,1676,1701]" captionStart="FIGURE 8" captionStartId="7.[151,250,890,912]" captionTargetBox="[151,1436,193,868]" captionTargetId="figure@7.[151,1436,193,868]" captionTargetPageId="7" captionText="FIGURE 8. Neoelmis guarani, female lateral habitus; length 2.3 mm." httpUri="https://zenodo.org/record/1053876/files/figure.png" pageId="7" pageNumber="425">Fig. 8</figureCitation>
). Mesoventrite and abdominal ventrite 2 lacking modifications. All legs (
<figureCitation id="CAEE2A7FFFC2F313848BFDBEB7A69BF9" box="[420,499,1712,1737]" captionStart="FIGURE 7" captionStartId="6.[151,250,1948,1970]" captionTargetBox="[151,1436,193,1927]" captionTargetId="figure@6.[151,1436,193,1927]" captionTargetPageId="6" captionText="FIGURE 7. Neoelmis guarani, female dorsal habitus; length 2.3 mm. Arrow indicates paired elytral protuberances." httpUri="https://zenodo.org/record/1053874/files/figure.png" pageId="7" pageNumber="425">Figs. 7</figureCitation>
, 
<figureCitation id="CAEE2A7FFFC2F31384D1FDBFB4599BF8" box="[510,524,1713,1737]" captionStart="FIGURE 8" captionStartId="7.[151,250,890,912]" captionTargetBox="[151,1436,193,868]" captionTargetId="figure@7.[151,1436,193,868]" captionTargetPageId="7" captionText="FIGURE 8. Neoelmis guarani, female lateral habitus; length 2.3 mm." httpUri="https://zenodo.org/record/1053876/files/figure.png" pageId="7" pageNumber="425">8</figureCitation>
) without special modifications, similar to each other; tibiae with cleaning fringes. Ovipositor very elongate; valvifers more than 10 times longer than wide, thin basally then widest apically by coxites; coxites twice as long as wide, rectangular; styli 2-segmented, second segment very short.
</paragraph>
</subSubSection>
<caption id="06AA6672FFCDF31C85B8FC50B5749A45" box="[151,801,1886,1908]" httpUri="https://zenodo.org/record/1053878/files/figure.png" pageId="8" pageNumber="426" targetBox="[151,1436,151,1864]" targetPageId="8">
<paragraph id="526A36FAFFCDF31C85B8FC50B5749A45" blockId="8.[151,801,1886,1908]" box="[151,801,1886,1908]" pageId="8" pageNumber="426">
<emphasis id="60A1EAE8FFCDF31C85B8FC50B7479A42" bold="true" box="[151,274,1886,1908]" pageId="8" pageNumber="426">FIGURE 9.</emphasis>
Arroyo Naranjo, type locality of 
<taxonomicName id="95D54D79FFCDF31C8749FC50B54E9A45" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[614,795,1886,1908]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="426" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCDF31C8749FC50B54E9A45" box="[614,795,1886,1908]" italics="true" pageId="8" pageNumber="426">Neoelmis guarani</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="06AA6672FFCCF31D85B8FF75B57899A0" box="[151,813,1147,1169]" httpUri="https://zenodo.org/record/1053880/files/figure.png" pageId="9" pageNumber="427" targetBox="[151,1436,161,1125]" targetPageId="9">
<paragraph id="526A36FAFFCCF31D85B8FF75B57899A0" blockId="9.[151,813,1147,1169]" box="[151,813,1147,1169]" pageId="9" pageNumber="427">
<emphasis id="60A1EAE8FFCCF31D85B8FF75B74A99A1" bold="true" box="[151,287,1147,1169]" pageId="9" pageNumber="427">FIGURE 10.</emphasis>
Arroyo Naranjo, type locality of 
<taxonomicName id="95D54D79FFCCF31D875CFF75B57299A0" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[627,807,1147,1169]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCCF31D875CFF75B57299A0" box="[627,807,1147,1169]" italics="true" pageId="9" pageNumber="427">Neoelmis guarani</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="1ACF6571FFCCF31D85E8FFB3B4D2989F" pageId="9" pageNumber="427" type="description">
<paragraph id="526A36FAFFCCF31D85E8FFB3B4D2989F" blockId="9.[151,1437,1213,2030]" pageId="9" pageNumber="427">
<emphasis id="60A1EAE8FFCCF31D85E8FFB3B76A99E7" bold="true" box="[199,319,1213,1238]" pageId="9" pageNumber="427">Variation.</emphasis>
Although seven specimens (three males and four females) is a small sample with which to examine variation, some differences were noted. Most striking of these are the different external modifications of males and females, detailed in the descriptions and Diagnosis. In addition, the females (
<quantity id="952D9B1FFFCCF31D86D3FE08B2D0982F" box="[1020,1157,1286,1310]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="9" pageNumber="427" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.2–2.3 mm</quantity>
long, 
<quantity id="952D9B1FFFCCF31D81E2FE08B300982F" box="[1229,1365,1286,1310]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="9" pageNumber="427" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
wide) are slightly larger than the males (
<quantity id="952D9B1FFFCCF31D870CFE24B4FB9870" box="[547,686,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="9" pageNumber="427" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.1–2.2 mm</quantity>
long, 
<quantity id="952D9B1FFFCCF31D87D5FE24B5079870" box="[762,850,1322,1346]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="427" unit="mm" value="0.7">0.7 mm</quantity>
wide). In some specimens mineral deposits on the dorsum obscure details of the surface, causing semi-glabrous, shiny areas to appear granulate. Specimens also exhibit varying degrees of surface abrasion which can lessen the number of setae present in the abraded area compared to that of an unabraded specimen.
</paragraph>
</subSubSection>
<subSubSection id="1ACF6571FFCCF31D85E8FEB7B2D09A27" pageId="9" pageNumber="427" type="diagnosis">
<paragraph id="526A36FAFFCCF31D85E8FEB7B7D69B53" blockId="9.[151,1437,1213,2030]" pageId="9" pageNumber="427">
<emphasis id="60A1EAE8FFCCF31D85E8FEB7B76F98E3" bold="true" box="[199,314,1465,1490]" pageId="9" pageNumber="427">Diagnosis</emphasis>
. The distinctive secondary sexual characters of 
<taxonomicName id="95D54D79FFCCF31D867DFEB4B59F98E0" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[850,970,1466,1489]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCCF31D867DFEB4B59F98E0" box="[850,970,1466,1489]" italics="true" pageId="9" pageNumber="427">N. guarani</emphasis>
</taxonomicName>
, present in both males and females, serve to distinguish this species from all other known species of 
<taxonomicName id="95D54D79FFCCF31D861FFED0B5CD98C7" authorityName="" authorityYear="1935" box="[816,920,1502,1526]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="60A1EAE8FFCCF31D861FFED0B5CD98C7" box="[816,920,1502,1526]" italics="true" pageId="9" pageNumber="427">Neoelmis</emphasis>
</taxonomicName>
. Males (
<figureCitation id="CAEE2A7FFFCCF31D86D3FED3B23398C7" box="[1020,1126,1501,1526]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 3 – 5" captionStartId-0="2.[151,250,1973,1995]" captionStartId-1="3.[151,250,1973,1995]" captionStartId-2="4.[151,264,1934,1957]" captionTargetBox-0="[181,1334,264,1941]" captionTargetBox-1="[247,1333,193,1916]" captionTargetBox-2="[257,1336,276,1839]" captionTargetId-0="figure@2.[171,1415,193,1943]" captionTargetId-1="figure@3.[247,1334,193,1916]" captionTargetId-2="figure@4.[180,1406,193,2007]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Neoelmis guarani, male dorsal habitus; length 2.1 mm." captionText-1="FIGURE 2. Neoelmis guarani, male ventral habitus; length 2.2 mm. Arrows indicate paired processes on the mesothorax and abdominal ventrite 2." captionText-2="FIGURES 3 – 5. Neoelmis guarani, male. 3. Left prothoracic leg, femur, dorsal view. 4. Left prothoracic leg, posteroventral view. 5. Left mesothoracic leg, anteroventral view." httpUri-0="https://zenodo.org/record/1053866/files/figure.png" httpUri-1="https://zenodo.org/record/1053868/files/figure.png" httpUri-2="https://zenodo.org/record/1053870/files/figure.png" pageId="9" pageNumber="427">Figs. 1–5</figureCitation>
) have strong modifications of the profemora, protibiae, mesofemora, mesotibiae, and metatibiae, and bear a pair of ventral processes on both the mesoventrite and second abdominal ventrite. Females (
<figureCitation id="CAEE2A7FFFCCF31D8617FD2BB5DC9B0F" box="[824,905,1573,1598]" captionStart="FIGURE 7" captionStartId="6.[151,250,1948,1970]" captionTargetBox="[151,1436,193,1927]" captionTargetId="figure@6.[151,1436,193,1927]" captionTargetPageId="6" captionText="FIGURE 7. Neoelmis guarani, female dorsal habitus; length 2.3 mm. Arrow indicates paired elytral protuberances." httpUri="https://zenodo.org/record/1053874/files/figure.png" pageId="9" pageNumber="427">Figs. 7</figureCitation>
, 
<figureCitation id="CAEE2A7FFFCCF31D86B7FD28B5F39B0F" box="[920,934,1574,1598]" captionStart="FIGURE 8" captionStartId="7.[151,250,890,912]" captionTargetBox="[151,1436,193,868]" captionTargetId="figure@7.[151,1436,193,868]" captionTargetPageId="7" captionText="FIGURE 8. Neoelmis guarani, female lateral habitus; length 2.3 mm." httpUri="https://zenodo.org/record/1053876/files/figure.png" pageId="9" pageNumber="427">8</figureCitation>
) have a dorsal pair of elytral protuberances and unmodified legs.
</paragraph>
<paragraph id="526A36FAFFCCF31D85E8FD63B2D09A27" blockId="9.[151,1437,1213,2030]" pageId="9" pageNumber="427">
The aedeagus of 
<taxonomicName id="95D54D79FFCCF31D84A5FD61B4509BB7" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[394,517,1647,1670]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCCF31D84A5FD61B4509BB7" box="[394,517,1647,1670]" italics="true" pageId="9" pageNumber="427">N. guarani</emphasis>
</taxonomicName>
most closely resembles that of 
<taxonomicName id="95D54D79FFCCF31D8641FD61B2339BB7" authority="(Grouvelle)" authorityName="Grouvelle" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" box="[878,1126,1645,1670]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="60A1EAE8FFCCF31D8641FD61B58E9BB7" box="[878,987,1647,1670]" italics="true" pageId="9" pageNumber="427">N. simoni</emphasis>
(Grouvelle)
</taxonomicName>
which is known only from 
<collectingCountry id="2AC2766AFFCCF31D85B8FD9FB75F9B9B" box="[151,266,1681,1706]" name="Venezuela" pageId="9" pageNumber="427">Venezuela</collectingCountry>
(
<bibRefCitation id="36444B0BFFCCF31D8437FD9FB7FF9B9B" author="Hinton, H. E." box="[280,426,1681,1706]" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="9" pageNumber="427" pagination="228 - 234" part="75" refString="Hinton, H. E. (1939) On some new and little known South American Neoelmis Musgrave (Coleoptera, Elmidae). The Entomologist's Monthly Magazine, 75, 228 - 234." title="On some new and little known South American Neoelmis Musgrave (Coleoptera, Elmidae)" type="journal article" year="1939">Hinton 1939</bibRefCitation>
). However, in 
<taxonomicName id="95D54D79FFCCF31D8762FD9CB4939B98" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[589,710,1682,1705]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCCF31D8762FD9CB4939B98" box="[589,710,1682,1705]" italics="true" pageId="9" pageNumber="427">N. guarani</emphasis>
</taxonomicName>
the phallobase is open only in the basal half and the penis barely projects beyond the tips of the parameres, whereas in 
<taxonomicName id="95D54D79FFCCF31D863AFDB9B5D09BFF" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" box="[789,901,1719,1742]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="60A1EAE8FFCCF31D863AFDB9B5D09BFF" box="[789,901,1719,1742]" italics="true" pageId="9" pageNumber="427">N. simoni</emphasis>
</taxonomicName>
the phallobase is open entirely and the penis projects well beyond the tips of the parameres. The aedeagus of 
<taxonomicName id="95D54D79FFCCF31D8652FDD4B5AD9BC0" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[893,1016,1754,1777]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCCF31D8652FDD4B5AD9BC0" box="[893,1016,1754,1777]" italics="true" pageId="9" pageNumber="427">N. guarani</emphasis>
</taxonomicName>
bears no resemblance to those of 
<taxonomicName id="95D54D79FFCCF31D80ADFDD4B6AA9A27" authorityName="Hinton" authorityYear="1940" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="60A1EAE8FFCCF31D80ADFDD4B6AA9A27" italics="true" pageId="9" pageNumber="427">N. maculata</emphasis>
</taxonomicName>
, 
<taxonomicName id="95D54D79FFCCF31D8422FDF1B7089A27" authorityName="Hinton" authorityYear="1972" box="[269,349,1790,1814]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="nelo">
<emphasis id="60A1EAE8FFCCF31D8422FDF1B7089A27" box="[269,349,1790,1814]" italics="true" pageId="9" pageNumber="427">N. nelo</emphasis>
</taxonomicName>
, nor 
<taxonomicName id="95D54D79FFCCF31D84B7FDF1B7B29A27" authorityName="Hinton (Shepard &amp; Aguilar Julio" authorityYear="2010" box="[408,487,1791,1814]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="opis">
<emphasis id="60A1EAE8FFCCF31D84B7FDF1B7B29A27" box="[408,487,1791,1814]" italics="true" pageId="9" pageNumber="427">N. opis</emphasis>
</taxonomicName>
, other species occurring in 
<collectingRegion id="9011F818FFCCF31D8632FDF1B5D09A26" box="[797,901,1791,1815]" country="Paraguay" name="Asuncion" pageId="9" pageNumber="427">
<collectingCountry id="2AC2766AFFCCF31D8632FDF1B5D09A26" box="[797,901,1791,1815]" name="Paraguay" pageId="9" pageNumber="427">Paraguay</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="36444B0BFFCCF31D86BCFDF3B2649A27" author="Hinton, H. E." box="[915,1073,1789,1814]" journalOrPublisher="The Annals and Magazine of Natural History, Series 11" pageId="9" pageNumber="427" pagination="129 - 153" part="5" refString="Hinton, H. E. (1940 a) A synopsis of the Brazilian species of Neoelmis Musgrave (Coleoptera, Elmidae). The Annals and Magazine of Natural History, Series 11, 5 (26), 129 - 153." title="A synopsis of the Brazilian species of Neoelmis Musgrave (Coleoptera, Elmidae)" type="journal article" year="1940" yearSuffix="a">Hinton 1940a</bibRefCitation>
, 
<bibRefCitation id="36444B0BFFCCF31D8112FDF0B22C9A27" author="Hinton, H. E." box="[1085,1145,1790,1814]" journalOrPublisher="Papeis Avulsos de Zoologia, 26" pageId="9" pageNumber="427" pagination="117 - 135" part="9" refString="Hinton, H. E. (1972) New species of Neoelmis from South America (Coleoptera, Elmidae). Papeis Avulsos de Zoologia, 26 (9), 117 - 135, pls. I - II." title="New species of Neoelmis from South America (Coleoptera, Elmidae)" type="journal article" year="1972">1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1ACF6571FFCCF31F85E8FC2FB5069B05" lastPageId="11" lastPageNumber="429" pageId="9" pageNumber="427" type="materials_examined">
<paragraph id="526A36FAFFCCF31D85E8FC2FB69C9ADF" blockId="9.[151,1437,1213,2030]" pageId="9" pageNumber="427">
<emphasis id="60A1EAE8FFCCF31D85E8FC2FB7239A0B" bold="true" box="[199,374,1825,1850]" pageId="9" pageNumber="427">
<typeStatus id="8D6E8858FFCCF31D85E8FC2FB7579A0B" box="[199,258,1825,1850]" pageId="9" pageNumber="427">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="E2BD3CA7FFCCF31D8452FC2CB2AB9A6F" ID-GBIF-Occurrence="1701204106" collectingDate="2011-02-17" collectorName="C. B. Barr" elevation="214" latitude="-25.533766" location="Arroyo Naranjo at Balneario Salto Cristal 7.5 km S Piribebuy" longLatPrecision="1" longitude="-57.028618" municipality="Dpto" pageId="9" pageNumber="427" specimenCount="1" specimenCount-male="1" stateProvince="PARAGUAY" typeStatus="holotype">
<typeStatus id="8D6E8858FFCCF31D8452FC2CB7B19A0B" box="[381,484,1826,1850]" pageId="9" pageNumber="427" type="holotype">
<emphasis id="60A1EAE8FFCCF31D8452FC2CB7B19A0B" box="[381,484,1826,1850]" italics="true" pageId="9" pageNumber="427">Holotype</emphasis>
</typeStatus>
(
<specimenCount id="44D3FD73FFCCF31D84DBFC2FB4649A0B" box="[500,561,1825,1850]" pageId="9" pageNumber="427" type="male">male</specimenCount>
): 
<emphasis id="60A1EAE8FFCCF31D876AFC2FB4BD9A0B" bold="true" box="[581,744,1825,1850]" pageId="9" pageNumber="427">
<collectingRegion id="9011F818FFCCF31D876AFC2FB4B69A0B" box="[581,739,1825,1850]" country="Paraguay" name="Asuncion" pageId="9" pageNumber="427">PARAGUAY</collectingRegion>
:
</emphasis>
<collectingRegion id="9011F818FFCCF31D87C0FC2CB50F9A0B" box="[751,858,1826,1850]" country="Paraguay" name="Paraguari" pageId="9" pageNumber="427">Paraguarí</collectingRegion>
[
<collectingMunicipality id="B20EAC80FFCCF31D8645FC2CB5F99A08" box="[874,940,1826,1849]" pageId="9" pageNumber="427">Dpto</collectingMunicipality>
.], 
<location id="570A6021FFCCF31D86EBFC2CB72E9A6F" LSID="urn:lsid:plazi:treatment:DA7C87ECFFC4F31885B8F8FAB7269DE4:570A6021FFCCF31D86EBFC2CB72E9A6F" latitude="-25.533766" longLatPrecision="1" longitude="-57.028618" municipality="Dpto" name="Arroyo Naranjo at Balneario Salto Cristal 7.5 km S Piribebuy" pageId="9" pageNumber="427" stateProvince="PARAGUAY">
Arroyo Naranjo at Balneario Salto Cristal 
<quantity id="952D9B1FFFCCF31D85B8FC4BB6BD9A6F" box="[151,232,1861,1886]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="427" unit="km" value="7.5">7.5 km</quantity>
S Piribebuy
</location>
, 
<collectingDate id="362FE9D2FFCCF31D84A5FC48B4729A6F" box="[394,551,1861,1886]" pageId="9" pageNumber="427" value="2011-02-17">
<date id="266B103AFFCCF31D84A5FC48B4729A6F" box="[394,551,1861,1886]" pageId="9" pageNumber="427" value="2011-02-17">17 Feb. 2011</date>
</collectingDate>
, 
<collectorName id="FF20532CFFCCF31D8719FC48B4E49A6F" box="[566,689,1862,1886]" pageId="9" pageNumber="427">C. B. Barr</collectorName>
// 
<geoCoordinate id="37E1503DFFCCF31D87FAFC4BB53E9A6F" box="[725,875,1861,1886]" direction="south" orientation="latitude" pageId="9" pageNumber="427" precision="1" value="-25.533766">25º32.026’ S</geoCoordinate>
<geoCoordinate id="37E1503DFFCCF31D8659FC4BB24D9A6F" box="[886,1048,1861,1886]" direction="west" orientation="longitude" pageId="9" pageNumber="427" precision="1" value="-57.028618">57º01.717’ W</geoCoordinate>
; elevation 
<elevation id="D9F8D1C9FFCCF31D81B6FC48B2B69A6F" box="[1177,1251,1862,1886]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="9" pageNumber="427" unit="m" value="214.0">
<quantity id="952D9B1FFFCCF31D81B6FC48B2B69A6F" box="[1177,1251,1862,1886]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="9" pageNumber="427" unit="m" value="214.0">214 m</quantity>
</elevation>
//
</materialsCitation>
<typeStatus id="8D6E8858FFCCF31D8027FC48B3C99A6F" box="[1288,1436,1862,1886]" pageId="9" pageNumber="427" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="95D54D79FFCCF31D85B8FC64B4429AB0" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[151,535,1897,1922]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCCF31D85B8FC64B70B9AB0" box="[151,350,1898,1922]" italics="true" pageId="9" pageNumber="427">Neoelmis guarani</emphasis>
Shepard &amp; Barr
</taxonomicName>
[red label, handwritten]. Deposited in 
<collectionCode id="34C4AE3FFFCCF31D86E6FC64B24F9AB3" box="[969,1050,1898,1922]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="9" pageNumber="427">EMEC</collectionCode>
. A 
<emphasis id="60A1EAE8FFCCF31D8116FC64B2D39AB3" box="[1081,1158,1898,1922]" italics="true" pageId="9" pageNumber="427">llotype</emphasis>
(female): 
<emphasis id="60A1EAE8FFCCF31D81D5FC67B3C99AB3" bold="true" box="[1274,1436,1897,1922]" pageId="9" pageNumber="427">
<collectingRegion id="9011F818FFCCF31D81D5FC67B3CD9AB3" box="[1274,1432,1897,1922]" country="Paraguay" name="Asuncion" pageId="9" pageNumber="427">
<collectingCountry id="2AC2766AFFCCF31D81D5FC67B3CD9AB3" box="[1274,1432,1897,1922]" name="Paraguay" pageId="9" pageNumber="427">PARAGUAY</collectingCountry>
</collectingRegion>
:
</emphasis>
<collectingRegion id="9011F818FFCCF31D85B8FC83B75C9A97" box="[151,265,1933,1958]" country="Paraguay" name="Cordillera" pageId="9" pageNumber="427">Cordillera</collectingRegion>
[Dpto.], Piribebuy-Barrio Santa Ana, unnamed stream, 
<quantity id="952D9B1FFFCCF31D86A7FC80B5999A97" box="[904,972,1933,1958]" metricMagnitude="2" metricUnit="m" metricValue="2.49936" pageId="9" pageNumber="427" unit="ft" value="820.0">820 ft</quantity>
[
<quantity id="952D9B1FFFCCF31D86F4FC83B27E9A97" box="[987,1067,1933,1958]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="427" unit="m" value="250.0">250 m</quantity>
], 
<geoCoordinate id="37E1503DFFCCF31D816FFC83B2E99A97" box="[1088,1212,1933,1958]" direction="south" orientation="latitude" pageId="9" pageNumber="427" precision="9" value="-25.465834">25º27.95’S</geoCoordinate>
<geoCoordinate id="37E1503DFFCCF31D81EAFC83B3189A97" box="[1221,1357,1933,1958]" direction="west" orientation="longitude" pageId="9" pageNumber="427" precision="9" value="-57.033165">57º01.99’W</geoCoordinate>
, 
<date id="266B103AFFCCF31D8075FC80B6879AFB" pageId="9" pageNumber="427" value="2006-06-18">18 VI 2006</date>
, [WDS-A-1687, on underside of label] // W. D. Shepard, leg. // 
<typeStatus id="8D6E8858FFCCF31D869AFCBCB2139AFB" box="[949,1094,1970,1994]" pageId="9" pageNumber="427" type="allotype">ALLOTYPE</typeStatus>
<taxonomicName id="95D54D79FFCCF31D8161FCBCB69C9ADF" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" authorityYear="2016" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="427" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCCF31D8161FCBCB34C9AF8" box="[1102,1305,1970,1994]" italics="true" pageId="9" pageNumber="427">Neoelmis guarani</emphasis>
Shepard &amp; Barr
</taxonomicName>
</paragraph>
<caption id="06AA6672FFCFF31E85B8FC86B5729AAF" box="[151,807,1928,1950]" httpUri="https://zenodo.org/record/1053882/files/figure.png" pageId="10" pageNumber="428" targetBox="[151,1436,193,1906]" targetPageId="10">
<paragraph id="526A36FAFFCFF31E85B8FC86B5729AAF" blockId="10.[151,807,1928,1950]" box="[151,807,1928,1950]" pageId="10" pageNumber="428">
<emphasis id="60A1EAE8FFCFF31E85B8FC86B74B9AAC" bold="true" box="[151,286,1928,1950]" pageId="10" pageNumber="428">FIGURE 11.</emphasis>
Arroyo Mborebí, 
<taxonomicName id="95D54D79FFCFF31E84F8FC86B4D99AAF" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[471,652,1928,1950]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="428" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCFF31E84F8FC86B4D99AAF" box="[471,652,1928,1950]" italics="true" pageId="10" pageNumber="428">Neoelmis guarani</emphasis>
</taxonomicName>
collection site.
</paragraph>
</caption>
<caption id="06AA6672FFCEF31F85B8FF94B4AF9981" box="[151,762,1178,1200]" httpUri="https://zenodo.org/record/1053884/files/figure.png" pageId="11" pageNumber="429" targetBox="[151,1436,193,1157]" targetPageId="11">
<paragraph id="526A36FAFFCEF31F85B8FF94B4AF9981" blockId="11.[151,762,1178,1200]" box="[151,762,1178,1200]" pageId="11" pageNumber="429">
<emphasis id="60A1EAE8FFCEF31F85B8FF94B74A999E" bold="true" box="[151,287,1178,1200]" pageId="11" pageNumber="429">FIGURE 12.</emphasis>
Capilla Cue, 
<taxonomicName id="95D54D79FFCEF31F8485FF94B40A9981" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[426,607,1178,1200]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCEF31F8485FF94B40A9981" box="[426,607,1178,1200]" italics="true" pageId="11" pageNumber="429">Neoelmis guarani</emphasis>
</taxonomicName>
collection site.
</paragraph>
</caption>
<paragraph id="526A36FAFFCEF31F85B8FFD9B5069B05" blockId="11.[151,1437,1239,2021]" pageId="11" pageNumber="429">
[red label, handwritten]. Deposited in 
<collectionCode id="34C4AE3FFFCEF31F875BFFD6B49099C1" box="[628,709,1240,1264]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="11" pageNumber="429">EMEC</collectionCode>
. 
<typeStatus id="8D6E8858FFCEF31F87F5FFD7B51999C1" box="[730,844,1241,1264]" pageId="11" pageNumber="429" type="paratype">
<emphasis id="60A1EAE8FFCEF31F87F5FFD7B51999C1" box="[730,844,1241,1264]" italics="true" pageId="11" pageNumber="429">Paratypes</emphasis>
</typeStatus>
(2MM, 3 
<collectionCode id="34C4AE3FFFCEF31F86E4FFD6B5A599C1" box="[971,1008,1240,1264]" pageId="11" pageNumber="429">FF</collectionCode>
): 
<emphasis id="60A1EAE8FFCEF31F8122FFD9B2E599C1" bold="true" box="[1037,1200,1239,1264]" pageId="11" pageNumber="429">
<collectingRegion id="9011F818FFCEF31F8122FFD9B2FE99C1" box="[1037,1195,1239,1264]" country="Paraguay" name="Asuncion" pageId="11" pageNumber="429">
<collectingCountry id="2AC2766AFFCEF31F8122FFD9B2FE99C1" box="[1037,1195,1239,1264]" name="Paraguay" pageId="11" pageNumber="429">PARAGUAY</collectingCountry>
</collectingRegion>
:
</emphasis>
<collectingRegion id="9011F818FFCEF31F81EEFFD9B36699C1" box="[1217,1331,1239,1264]" country="Paraguay" name="Cordillera" pageId="11" pageNumber="429">Cordillera</collectingRegion>
[Dpto.], Piribebuy-Barrio Santa Ana, unnamed stream, 
<quantity id="952D9B1FFFCEF31F878CFFF2B4B09825" box="[675,741,1275,1300]" metricMagnitude="2" metricUnit="m" metricValue="2.49936" pageId="11" pageNumber="429" unit="ft" value="820.0">820 ft</quantity>
[
<quantity id="952D9B1FFFCEF31F87DDFFF5B5159822" box="[754,832,1275,1300]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="429" unit="m" value="250.0">250 m</quantity>
], 
<geoCoordinate id="37E1503DFFCEF31F867CFFF5B59A9825" box="[851,975,1275,1300]" direction="south" orientation="latitude" pageId="11" pageNumber="429" precision="9" value="-25.465834">25º27.95’S</geoCoordinate>
<geoCoordinate id="37E1503DFFCEF31F86F9FFF5B2089825" box="[982,1117,1275,1300]" direction="west" orientation="longitude" pageId="11" pageNumber="429" precision="9" value="-57.033165">57º01.99’W</geoCoordinate>
, 
<date id="266B103AFFCEF31F8146FFF2B2BD9825" box="[1129,1256,1275,1300]" pageId="11" pageNumber="429" value="2006-06-18">18 VI 2006</date>
[WDS-A-1687, on underside of label] // W. D. Shepard, leg. // 
<typeStatus id="8D6E8858FFCEF31F8789FE2EB5649809" box="[678,817,1312,1336]" pageId="11" pageNumber="429" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="95D54D79FFCEF31F8617FE2EB2EF9806" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[824,1210,1311,1336]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCEF31F8617FE2EB2559806" box="[824,1024,1312,1336]" italics="true" pageId="11" pageNumber="429">Neoelmis guarani</emphasis>
Shepard &amp; Barr
</taxonomicName>
[yellow label] (1M, 
<collectionCode id="34C4AE3FFFCEF31F85B8FE4AB6BF986D" box="[151,234,1348,1372]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="11" pageNumber="429">EMEC</collectionCode>
); 
<collectingRegion id="9011F818FFCEF31F85D3FE4DB73B986D" box="[252,366,1347,1372]" country="Paraguay" name="Cordillera" pageId="11" pageNumber="429">Cordillera</collectingRegion>
[Dpto.], Piribebuy–B. Sta. Ana, 
<date id="266B103AFFCEF31F87F3FE4AB504986D" box="[732,849,1348,1372]" pageId="11" pageNumber="429" value="2011-02-17">
17 II 
<quantity id="952D9B1FFFCEF31F8638FE4AB504986D" box="[791,849,1348,1372]" metricMagnitude="6" metricUnit="m" metricValue="2.01123" pageId="11" pageNumber="429" unit="m" value="2011230.0">2011</quantity>
</date>
, 230 m, Arroyo Mborebi, 
<geoCoordinate id="37E1503DFFCEF31F8159FE4DB355986D" box="[1142,1280,1347,1372]" direction="south" orientation="latitude" pageId="11" pageNumber="429" precision="1" value="-25.467916">S25º28.075’</geoCoordinate>
<geoCoordinate id="37E1503DFFCEF31F8028FE4DB3C9986D" box="[1287,1436,1347,1372]" direction="west" orientation="longitude" pageId="11" pageNumber="429" precision="1" value="-57.038834">W57º02.330’</geoCoordinate>
// W. D. Shepard, leg. [WDS-A- 1830, on underside of label] // 
<typeStatus id="8D6E8858FFCEF31F8652FE66B25D98B1" box="[893,1032,1384,1408]" pageId="11" pageNumber="429" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="95D54D79FFCEF31F813EFE66B3C9984E" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[1041,1436,1383,1408]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCEF31F813EFE66B28E984E" box="[1041,1243,1384,1408]" italics="true" pageId="11" pageNumber="429">Neoelmis guarani</emphasis>
Shepard &amp; Barr
</taxonomicName>
[yellow label] (1M, 
<collectionCode id="34C4AE3FFFCEF31F8451FE82B7969895" box="[382,451,1420,1444]" country="Paraguay" httpUri="http://grbio.org/cool/nb6b-jwsh" name="Inventorio Biologico Nacional [Museo Nacional de Historia Natural del Paraguay]" pageId="11" pageNumber="429">INBP</collectionCode>
); 
<collectingRegion id="9011F818FFCEF31F84F7FE82B4169895" box="[472,579,1420,1444]" country="Paraguay" name="Paraguari" pageId="11" pageNumber="429">Paraguarí</collectingRegion>
[Dpto.], Arroyo Naranjo at Balneario Salto Cristal 
<quantity id="952D9B1FFFCEF31F81BBFE85B2B19895" box="[1172,1252,1419,1444]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="429" unit="km" value="7.5">7.5 km</quantity>
S Piribebuy, 
<date id="266B103AFFCEF31F80AFFE82B75C98F9" pageId="11" pageNumber="429" value="2011-02-17">17 Feb. 2011</date>
, C.B. Barr // 
<geoCoordinate id="37E1503DFFCEF31F848CFEA1B46098F9" box="[419,565,1455,1480]" direction="south" orientation="latitude" pageId="11" pageNumber="429" precision="1" value="-25.533766">25º32.026’ S</geoCoordinate>
<geoCoordinate id="37E1503DFFCEF31F8711FEA1B48998F6" box="[574,732,1455,1480]" direction="west" orientation="longitude" pageId="11" pageNumber="429" precision="1" value="-57.028618">57º01.717’ W</geoCoordinate>
, elevation 
<quantity id="952D9B1FFFCEF31F8678FEBEB5CA98F6" box="[855,927,1456,1480]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="11" pageNumber="429" unit="m" value="214.0">214 m</quantity>
// 
<typeStatus id="8D6E8858FFCEF31F8691FEBEB21C98F9" box="[958,1097,1456,1480]" pageId="11" pageNumber="429" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="95D54D79FFCEF31F817EFEBEB69C98DA" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" authorityYear="2016" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCEF31F817EFEBEB34F98F6" box="[1105,1306,1456,1480]" italics="true" pageId="11" pageNumber="429">Neoelmis guarani</emphasis>
Shepard &amp; Barr
</taxonomicName>
[yellow label] (2 
<collectionCode id="34C4AE3FFFCEF31F84B5FEDAB7E898DA" box="[410,445,1492,1515]" pageId="11" pageNumber="429">FF</collectionCode>
; 
<collectionCode id="34C4AE3FFFCEF31F84E5FEDAB44E98DD" box="[458,539,1492,1516]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="11" pageNumber="429">EMEC</collectionCode>
, 
<collectionCode id="34C4AE3FFFCEF31F8707FEDAB43898DD" box="[552,621,1492,1516]" country="Paraguay" httpUri="http://grbio.org/cool/nb6b-jwsh" name="Inventorio Biologico Nacional [Museo Nacional de Historia Natural del Paraguay]" pageId="11" pageNumber="429">INBP</collectionCode>
); 
<collectingRegion id="9011F818FFCEF31F87ADFEDDB4B898DD" box="[642,749,1491,1516]" country="Paraguay" name="Paraguari" pageId="11" pageNumber="429">Paraguari</collectingRegion>
[Dpto.], 
<quantity id="952D9B1FFFCEF31F8677FEDDB5E298DD" box="[856,951,1491,1516]" metricMagnitude="4" metricUnit="m" metricValue="1.53" pageId="11" pageNumber="429" unit="km" value="15.3">15.3 km</quantity>
NE 
<collectingRegion id="9011F818FFCEF31F86C1FEDDB20998DD" box="[1006,1116,1491,1516]" country="Paraguay" name="Paraguari" pageId="11" pageNumber="429">Paraguari</collectingRegion>
, 
<date id="266B103AFFCEF31F8145FEDAB2B098DD" box="[1130,1253,1492,1516]" pageId="11" pageNumber="429" value="2011-02-18">
18 II 
<quantity id="952D9B1FFFCEF31F8184FEDAB2B098DD" box="[1195,1253,1492,1516]" metricMagnitude="6" metricUnit="m" metricValue="2.011282" pageId="11" pageNumber="429" unit="m" value="2011282.0">2011</quantity>
</date>
, 282 m, Capilla Cue, 
<geoCoordinate id="37E1503DFFCEF31F85F7FEF9B7379B21" box="[216,354,1527,1552]" direction="south" orientation="latitude" pageId="11" pageNumber="429" precision="1" value="-25.5535">S25º33.210’</geoCoordinate>
<geoCoordinate id="37E1503DFFCEF31F8440FEF9B4519B21" box="[367,516,1527,1552]" direction="west" orientation="longitude" pageId="11" pageNumber="429" precision="1" value="-57.04715">W57º02.829’</geoCoordinate>
// W. D. Shepard, leg. [WDS-A-1834, on underside of label] // 
<typeStatus id="8D6E8858FFCEF31F803EFEF6B3C99B21" box="[1297,1436,1528,1552]" pageId="11" pageNumber="429" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="95D54D79FFCEF31F85B8FD12B4429B02" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[151,535,1563,1588]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCEF31F85B8FD12B70B9B02" box="[151,350,1564,1588]" italics="true" pageId="11" pageNumber="429">Neoelmis guarani</emphasis>
Shepard &amp; Barr
</taxonomicName>
[yellow label] (1F, 
<collectionCode id="34C4AE3FFFCEF31F87DBFD12B5129B05" box="[756,839,1564,1588]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="11" pageNumber="429">EMEC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="1ACF6571FFCEF31F85E8FD31B5319B4D" pageId="11" pageNumber="429" type="etymology">
<paragraph id="526A36FAFFCEF31F85E8FD31B5319B4D" blockId="11.[151,1437,1239,2021]" pageId="11" pageNumber="429">
<emphasis id="60A1EAE8FFCEF31F85E8FD31B7139B69" bold="true" box="[199,326,1599,1624]" pageId="11" pageNumber="429">Etymology</emphasis>
. 
<emphasis id="60A1EAE8FFCEF31F847BFD4EB7EC9B66" box="[340,441,1600,1623]" italics="true" pageId="11" pageNumber="429">Guarani,</emphasis>
a noun in apposition, was chosen to honor the indigenous Guaraní people who are native to the region and whose language is widely spoken in 
<collectingRegion id="9011F818FFCEF31F87D9FD6AB5359B4D" box="[758,864,1636,1660]" country="Paraguay" name="Asuncion" pageId="11" pageNumber="429">
<collectingCountry id="2AC2766AFFCEF31F87D9FD6AB5359B4D" box="[758,864,1636,1660]" name="Paraguay" pageId="11" pageNumber="429">Paraguay</collectingCountry>
</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="1ACF6571FFCEF31885E8FD89B7269DE4" lastPageId="12" lastPageNumber="430" pageId="11" pageNumber="429" type="distribution">
<paragraph id="526A36FAFFCEF31F85E8FD89B5579A01" blockId="11.[151,1437,1239,2021]" pageId="11" pageNumber="429">
<emphasis id="60A1EAE8FFCEF31F85E8FD89B7359B91" bold="true" box="[199,352,1671,1696]" pageId="11" pageNumber="429">Distribution.</emphasis>
<taxonomicName id="95D54D79FFCEF31F8445FD86B4619BAE" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[362,564,1672,1696]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCEF31F8445FD86B4619BAE" box="[362,564,1672,1696]" italics="true" pageId="11" pageNumber="429">Neoelmis guarani</emphasis>
</taxonomicName>
is known from four localities in the highlands area of the 
<collectingRegion id="9011F818FFCEF31F81F3FD89B31B9B91" box="[1244,1358,1671,1696]" country="Paraguay" name="Cordillera" pageId="11" pageNumber="429">Cordillera</collectingRegion>
de los Altos southeast of 
<collectingRegion id="9011F818FFCEF31F8458FDA5B7B79BF5" box="[375,482,1707,1732]" country="Paraguay" name="Asuncion" pageId="11" pageNumber="429">Asunción</collectingRegion>
in the departments of 
<collectingRegion id="9011F818FFCEF31F87D9FDA5B53D9BF5" box="[758,872,1707,1732]" country="Paraguay" name="Cordillera" pageId="11" pageNumber="429">Cordillera</collectingRegion>
and 
<collectingRegion id="9011F818FFCEF31F8686FDA2B24D9BF5" box="[937,1048,1708,1732]" country="Paraguay" name="Paraguari" pageId="11" pageNumber="429">Paraguarí</collectingRegion>
. Although 42 streams have been sampled during our survey of the aquatic Byrrhoidea of 
<collectingRegion id="9011F818FFCEF31F860FFDDEB5DC9BD9" box="[800,905,1744,1768]" country="Paraguay" name="Asuncion" pageId="11" pageNumber="429">
<collectingCountry id="2AC2766AFFCEF31F860FFDDEB5DC9BD9" box="[800,905,1744,1768]" name="Paraguay" pageId="11" pageNumber="429">Paraguay</collectingCountry>
</collectingRegion>
, the species has been found only in this small area. The elevations at the collection sites range from 
<quantity id="952D9B1FFFCEF31F87DCFDFAB5C49A3D" box="[755,913,1779,1804]" metricMagnitude="2" metricUnit="m" metricValue="2.48" metricValueMax="2.82" metricValueMin="2.1399999999999997" pageId="11" pageNumber="429" unit="m" value="248.0" valueMax="282.0" valueMin="214.0">214–282 masl</quantity>
and they are all less than 
<quantity id="952D9B1FFFCEF31F819CFDFAB2AC9A3D" box="[1203,1273,1779,1804]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="429" unit="km" value="10.0">10 km</quantity>
apart (straight line distance) near the towns of Piribebuy and Chololó.
</paragraph>
<paragraph id="526A36FAFFCEF31885E8FC35B7269DE4" blockId="11.[151,1437,1239,2021]" lastBlockId="12.[151,1436,151,213]" lastPageId="12" lastPageNumber="430" pageId="11" pageNumber="429">
<emphasis id="60A1EAE8FFCEF31F85E8FC35B77F9A65" bold="true" box="[199,298,1851,1876]" pageId="11" pageNumber="429">Habitat.</emphasis>
The 
<typeStatus id="8D6E8858FFCEF31F844DFC32B7C79A65" box="[354,402,1852,1876]" pageId="11" pageNumber="429">type</typeStatus>
locality, Arroyo Naranjo (
<figureCitation id="CAEE2A7FFFCEF31F8797FC35B5529A65" box="[696,775,1851,1876]" captionStart="FIGURE 9" captionStartId="8.[151,250,1886,1908]" captionTargetBox="[151,1436,151,1864]" captionTargetId="figure@8.[151,1436,151,1864]" captionTargetPageId="8" captionText="FIGURE 9. Arroyo Naranjo, type locality of Neoelmis guarani." httpUri="https://zenodo.org/record/1053878/files/figure.png" pageId="11" pageNumber="429">Figs. 9</figureCitation>
, 
<figureCitation id="CAEE2A7FFFCEF31F863DFC32B5649A65" box="[786,817,1852,1876]" captionStart="FIGURE 10" captionStartId="9.[151,250,1147,1169]" captionTargetBox="[151,1436,161,1125]" captionTargetId="figure@9.[151,1436,161,1125]" captionTargetPageId="9" captionText="FIGURE 10. Arroyo Naranjo, type locality of Neoelmis guarani." httpUri="https://zenodo.org/record/1053880/files/figure.png" pageId="11" pageNumber="429">10</figureCitation>
), is a small to medium-sized stream with slightly turbid water and a substrate of orange sand with sparse gravel/cobbles, numerous bedrock outcrops and ledges, and small waterfalls. The three other streams are similar in size and substrate, except that Arroyo Mborebí (
<figureCitation id="CAEE2A7FFFCEF31F81D1FC8DB3059AAD" box="[1278,1360,1923,1948]" captionStart="FIGURE 11" captionStartId="10.[151,250,1928,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@10.[151,1436,193,1906]" captionTargetPageId="10" captionText="FIGURE 11. Arroyo Mborebí, Neoelmis guarani collection site." httpUri="https://zenodo.org/record/1053882/files/figure.png" pageId="11" pageNumber="429">Fig. 11</figureCitation>
) lacks extensive bedrock outcrops, at least at the collection site (
<figureCitation id="CAEE2A7FFFCEF31F860BFCA9B5D49AF1" box="[804,897,1959,1984]" captionStart="FIGURE 11" captionStartId="10.[151,250,1928,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@10.[151,1436,193,1906]" captionTargetPageId="10" captionText="FIGURE 11. Arroyo Mborebí, Neoelmis guarani collection site." httpUri="https://zenodo.org/record/1053882/files/figure.png" pageId="11" pageNumber="429">Figs. 11</figureCitation>
, 
<figureCitation id="CAEE2A7FFFCEF31F86A1FCA6B5F89AF1" box="[910,941,1960,1984]" captionStart="FIGURE 12" captionStartId="11.[151,250,1178,1200]" captionTargetBox="[151,1436,193,1157]" captionTargetId="figure@11.[151,1436,193,1157]" captionTargetPageId="11" captionText="FIGURE 12. Capilla Cue, Neoelmis guarani collection site." httpUri="https://zenodo.org/record/1053884/files/figure.png" pageId="11" pageNumber="429">12</figureCitation>
). Other elmids collected in association with 
<taxonomicName id="95D54D79FFCEF31F85B8FCC3B7469AD5" authorityName="Shepard &amp; Barr" authorityYear="2016" box="[151,275,1997,2020]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="60A1EAE8FFCEF31F85B8FCC3B7469AD5" box="[151,275,1997,2020]" italics="true" pageId="11" pageNumber="429">N. guarani</emphasis>
</taxonomicName>
include 
<taxonomicName id="95D54D79FFCEF31F8458FCC2B7A49AD5" authorityName="Sharp" authorityYear="1882" box="[375,497,1996,2020]" class="Insecta" family="Elmidae" genus="Heterelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="60A1EAE8FFCEF31F8458FCC2B7A49AD5" box="[375,497,1996,2020]" italics="true" pageId="11" pageNumber="429">Heterelmis</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="95D54D79FFCEF31F8705FCC2B49A9AD5" authorityName="Hinton" authorityYear="1940" box="[554,719,1996,2020]" class="Insecta" family="Elmidae" genus="Hexacylloepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="60A1EAE8FFCEF31F8705FCC2B49A9AD5" box="[554,719,1996,2020]" italics="true" pageId="11" pageNumber="429">Hexacylloepus</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="95D54D79FFCEF31F8627FCC2B5C89AD5" authorityName="Sharp" authorityYear="1882" box="[776,925,1996,2020]" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="60A1EAE8FFCEF31F8627FCC2B5C89AD5" box="[776,925,1996,2020]" italics="true" pageId="11" pageNumber="429">Hexanchorus</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="95D54D79FFCEF31F86F8FCC2B21B9AD5" box="[983,1102,1996,2020]" class="Insecta" family="Elmidae" genus="Macrelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="genus">
<emphasis id="60A1EAE8FFCEF31F86F8FCC2B21B9AD5" box="[983,1102,1996,2020]" italics="true" pageId="11" pageNumber="429">Macrelmis</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="95D54D79FFCEF31881A8FCC2B7499D81" authority="(Grouvelle)" authorityName="Grouvelle" baseAuthorityName="Grouvelle" baseAuthorityYear="1888" class="Insecta" family="Elmidae" genus="Microcylloepus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="12" lastPageNumber="430" order="Coleoptera" pageId="11" pageNumber="429" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="60A1EAE8FFCEF31F81A8FCC2B3C99AD5" box="[1159,1436,1996,2020]" italics="true" pageId="11" pageNumber="429">Microcylloepus longipes</emphasis>
(Grouvelle)
</taxonomicName>
, 
<taxonomicName id="95D54D79FFC9F3188402FB97B4649D81" authority="(Sharp)" authorityName="Sharp" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[301,561,151,176]" class="Insecta" family="Elmidae" genus="Microcylloepus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="430" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="60A1EAE8FFC9F3188402FB97B79B9D81" box="[301,462,152,176]" italics="true" pageId="12" pageNumber="430">M. inaequalis</emphasis>
(Sharp)
</taxonomicName>
, 
<taxonomicName id="95D54D79FFC9F318876CFB96B5139D81" authority="Hinton" authorityName="Hinton" authorityYear="1972" box="[579,838,151,176]" class="Insecta" family="Elmidae" genus="Neoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="430" phylum="Arthropoda" rank="species" species="nelo">
<emphasis id="60A1EAE8FFC9F318876CFB96B4BD9D81" box="[579,744,152,176]" italics="true" pageId="12" pageNumber="430">Neoelmis nelo</emphasis>
Hinton
</taxonomicName>
, 
<taxonomicName id="95D54D79FFC9F3188678FB96B5889D81" authorityName="Sharp" authorityYear="1882" box="[855,989,152,176]" class="Insecta" family="Elmidae" genus="Phanocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="60A1EAE8FFC9F3188678FB96B5889D81" box="[855,989,152,176]" italics="true" pageId="12" pageNumber="430">Phanocerus</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="95D54D79FFC9F3188131FB96B2EA9D81" authorityName="Grouvelle" authorityYear="1908" box="[1054,1215,152,176]" class="Insecta" family="Elmidae" genus="Stenhelmoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="60A1EAE8FFC9F3188131FB96B2EA9D81" box="[1054,1215,152,176]" italics="true" pageId="12" pageNumber="430">Stenhelmoides</emphasis>
</taxonomicName>
sp., and 
<taxonomicName id="95D54D79FFC9F318801AFB96B73A9DE4" authority="(Grouvelle)" authorityName="Grouvelle" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" class="Insecta" family="Elmidae" genus="Xenelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="430" phylum="Arthropoda" rank="species" species="micros">
<emphasis id="60A1EAE8FFC9F318801AFB96B6B79DE5" italics="true" pageId="12" pageNumber="430">Xenelmis micros</emphasis>
(Grouvelle)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>