533 lines
65 KiB
XML
533 lines
65 KiB
XML
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<mods:title id="6BF4A325F81A23201F4890F104460BA6">New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:name id="C0A1B3FD3CE42AF4D82E4AB009C83EA6" type="personal">
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<mods:namePart id="48960434F189DBFA7C81F95C84C36D90">Fedorenko, D. N.</mods:namePart>
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<mods:title id="80DD430C528815A2847AB749F2B2EF16">Russian Entomological Journal</mods:title>
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<mods:part id="3A27515ABF15792F5F5E55ABC4401C63">
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<mods:date id="2A4A8681D1F8AC6C672A8F029B143A90">2020</mods:date>
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<mods:number id="75CC0DC4418F699CBF3833B4EF3249E4">2020-03-31</mods:number>
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<treatment id="D6009B3AFFF73B43FC247F610C70FC4B" ID-DOI="http://doi.org/10.5281/zenodo.11093715" ID-Zenodo-Dep="11093715" LSID="urn:lsid:plazi:treatment:D6009B3AFFF73B43FC247F610C70FC4B" httpUri="http://treatment.plazi.org/id/D6009B3AFFF73B43FC247F610C70FC4B" lastPageId="6" lastPageNumber="44" pageId="3" pageNumber="41">
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<subSubSection id="16B379A7FFF73B46FC247F610AACFCB9" pageId="3" pageNumber="41" type="nomenclature">
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<paragraph id="5E162A2CFFF73B46FC247F610AACFCB9" blockId="3.[812,1399,784,1181]" pageId="3" pageNumber="41">
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<taxonomicName id="99A951AFFFF73B46FC247F610AEDFCB9" ID-CoL="8MBGZ" authority="Fedorenko" authorityName="Fedorenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="39" phylum="Arthropoda" rank="species" species="securipenis" status="sp. nov." subGenus="Steropanus">
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<heading id="055E9D40FFF73B46FC247F610A9FFCD4" box="[965,1246,784,808]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="41" reason="8">
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<emphasis id="6CDDF63EFFF73B46FC247F610A0AFCD4" box="[965,1099,784,808]" italics="true" pageId="3" pageNumber="41">Pterostichus</emphasis>
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(
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<emphasis id="6CDDF63EFFF73B46FBBC7F610A94FCDB" box="[1117,1237,784,807]" italics="true" pageId="3" pageNumber="41">Steropanus</emphasis>
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)
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</heading>
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||
<emphasis id="6CDDF63EFFF73B46FC547F5F0A6EFCB9" box="[949,1071,814,837]" italics="true" pageId="3" pageNumber="41">securipenis</emphasis>
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Fedorenko
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</taxonomicName>
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,
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<emphasis id="6CDDF63EFFF73B46FB597F5C0AACFCB9" bold="true" box="[1208,1261,813,837]" pageId="3" pageNumber="41">
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<taxonomicNameLabel id="77EE4B45FFF73B46FB597F5C0AACFCB9" box="[1208,1261,813,837]" pageId="3" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="16B379A7FFF73B46FC357F3B0A8EFC9E" box="[980,1231,842,866]" pageId="3" pageNumber="41" type="description">
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<paragraph id="5E162A2CFFF73B46FC357F3B0A8EFC9E" blockId="3.[812,1399,784,1181]" box="[980,1231,842,866]" pageId="3" pageNumber="41">
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<figureCitation id="C69236A9FFF73B46FC357F3B0A66FC9E" box="[980,1063,842,866]" captionStart="Figs 14–16" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[193,1367,1150,1943]" captionTargetId="figure-586@4.[189,1381,1138,1949]" captionTargetPageId="4" captionText="Figs 14–16. Dorsal habitus, holotypes: 14 — Pterostichus (Steropanus) securipenis sp.n; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n. Рис. 14–16. Габитус дорЗально, голотипы: 14 — Pterostichus (Steropanus) securipenis sp.n.; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n." figureDoi="http://doi.org/10.5281/zenodo.10978425" httpUri="https://zenodo.org/record/10978425/files/figure.png" pageId="3" pageNumber="41">Figs 14</figureCitation>
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,
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<figureCitation id="C69236A9FFF73B46FBD27F3B0A38FC9E" box="[1075,1145,842,866]" captionStart="Figs 17–22" captionStartId="5.[226,262,1921,1940]" captionTargetBox="[194,1391,758,1882]" captionTargetId="figure-432@5.[189,1399,755,1886]" captionTargetPageId="5" captionText="Figs 17–22. Aedeagus of Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 17–20 — median lobe; 17, 19, 21 — left lateral aspect; 18, 20, 22 — right lateral aspect. Scale bars: 1 mm. Рис. 17–22. Эдеагус Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 17–20 — среднЯЯ долЯ; 17, 19, 21 — слева; 18, 20, 22 — справа. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978427" httpUri="https://zenodo.org/record/10978427/files/figure.png" pageId="3" pageNumber="41">19–20</figureCitation>
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,
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<figureCitation id="C69236A9FFF73B46FB657F3B0A8AFC9E" box="[1156,1227,842,866]" captionStart="Figs 23–29" captionStartId="6.[226,263,1921,1940]" captionTargetBox="[192,1364,1003,1906]" captionTargetId="figure-533@6.[189,1369,999,1909]" captionTargetPageId="6" captionText="Figs 23–29. Aedeagus of Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (apical margin slightly damaged); 25, 28 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — median lobe, ventral aspect; 26–28 — inside of right paramere. Scale bars: 1 mm. Рис. 23–29. Эдеагус Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (апикальный край слегка обломан); 25, 28 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — среднЯЯ долЯ? вентрально; 26–28 — праваЯ парамера, иЗнутри. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978429" httpUri="https://zenodo.org/record/10978429/files/figure.png" pageId="3" pageNumber="41">24, 27</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="16B379A7FFF73B46FCB37F1E0AD7FC1E" pageId="3" pageNumber="41" type="materials_examined">
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<paragraph id="5E162A2CFFF73B46FCB37F1E0AD7FC1E" blockId="3.[812,1399,784,1181]" pageId="3" pageNumber="41">
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<materialsCitation id="EEC12071FFF73B46FCB37F1E0DA5FC36" collectingDate="2019-04-23" collectingDateMax="2019-05-11" collectingDateMin="2019-04-23" collectionCode="ZMMU" collectorName="D. Fedorenko'" country="Vietnam" county="Prov" elevation="1050" latitude="15.5633335" location="Song Thanh Nat" longLatPrecision="21" longitude="107.38944" municipality="Nam Gian Distr" pageId="3" pageNumber="41" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Quang Nam" typeStatus="holotype">
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MATERIAL.
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<typeStatus id="8112948EFFF73B46FC2A7F1E0A5AFC7E" box="[971,1051,879,898]" pageId="3" pageNumber="41" type="holotype">Holotype</typeStatus>
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<specimenCount id="48AFE1A5FFF73B46FBC17F1C0A75FC7F" box="[1056,1076,877,899]" count="1" pageId="3" pageNumber="41" type="male">♂</specimenCount>
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(
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<collectionCode id="38B8B2E9FFF73B46FBDF7F1E0AC4FC7E" box="[1086,1157,879,898]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="41" type="Museum">ZMMU</collectionCode>
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), labelled: ‘
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<collectingCountry id="26BE6ABCFFF73B46FB047F1E0B74FC7E" box="[1253,1333,879,898]" name="Vietnam" pageId="3" pageNumber="41">Vietnam</collectingCountry>
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,
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||
<collectingRegion id="9C6DE4CEFFF73B46FADC7F1E0D17FC67" country="Vietnam" name="Quang Nam" pageId="3" pageNumber="41">Quang Nam</collectingRegion>
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Prov[ince].,
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<collectingMunicipality id="BE72B056FFF73B46FC2C7FF90A22FC66" box="[973,1123,903,923]" pageId="3" pageNumber="41">Nam Gian Distr</collectingMunicipality>
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[ict].,
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||
<location id="5B767CF7FFF73B46FB747FF60B6EFC66" LSID="urn:lsid:plazi:treatment:D6009B3AFFF73B43FC247F610C70FC4B:5B767CF7FFF73B46FB747FF60B6EFC66" box="[1173,1327,903,922]" country="Vietnam" county="Prov" latitude="15.5633335" longLatPrecision="21" longitude="107.38944" municipality="Nam Gian Distr" name="Song Thanh Nat" pageId="3" pageNumber="41" stateProvince="Quang Nam">Song Thanh Nat</location>
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[io]n[al].
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||
<location id="5B767CF7FFF73B46FCCD7FEE0D17FC4E" LSID="urn:lsid:plazi:treatment:D6009B3AFFF73B43FC247F610C70FC4B:5B767CF7FFF73B46FCCD7FEE0D17FC4E" box="[812,854,927,946]" country="Vietnam" county="Prov" latitude="15.5633335" longLatPrecision="21" longitude="107.38944" municipality="Nam Gian Distr" name="Park" pageId="3" pageNumber="41" stateProvince="Quang Nam">Park</location>
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,
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<geoCoordinate id="3B9D4CEBFFF73B46FC847FEE0D91FC4E" box="[869,976,927,946]" degrees="15" direction="north" minutes="33" orientation="latitude" pageId="3" pageNumber="41" precision="15" seconds="48" value="15.5633335">15°33´48´´N</geoCoordinate>
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/
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<geoCoordinate id="3B9D4CEBFFF73B46FC027FEE0A1AFC4E" box="[995,1115,927,946]" degrees="107" direction="east" minutes="23" orientation="longitude" pageId="3" pageNumber="41" precision="15" seconds="22" value="107.38944">107°23´22´´E</geoCoordinate>
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, h=
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<quantity id="995187C9FFF73B46FB617FEE0A88FC4F" box="[1152,1225,927,947]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="41" unit="m" value="1050.0">
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<elevation id="D584CD1FFFF73B46FB617FEE0A88FC4F" box="[1152,1225,927,947]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="41" unit="m" value="1050.0">1050 m</elevation>
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</quantity>
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,
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<date id="2A170CECFFF73B46FB397FEE0B32FC4E" box="[1240,1395,927,946]" pageId="3" pageNumber="41" value="2019-04-23" valueMax="2019-05-11" valueMin="2019-04-23">
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<collectingDate id="3A53F504FFF73B46FB397FEE0B32FC4E" box="[1240,1395,927,946]" pageId="3" pageNumber="41" value="2019-04-23" valueMax="2019-05-11" valueMin="2019-04-23">23.IV–11.V.2019</collectingDate>
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</date>
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, leg.
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<collectorName id="F35C4FFAFFF73B46FCB97FC90DA1FC36" box="[856,992,951,971]" pageId="3" pageNumber="41">D. Fedorenko’</collectorName>
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.
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</materialsCitation>
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<materialsCitation id="EEC12071FFF73B46FC0F7FC90AD3FC1E" collectingDate="2019-04-23" collectingDateMax="2019-05-11" collectingDateMin="2019-04-23" collectionCode="SIEE" collectorName="D. Fedorenko'" country="Vietnam" county="Prov" elevation="1050" latitude="15.5633335" location="Song Thanh Nat" longLatPrecision="21" longitude="107.38944" municipality="Nam Gian Distr" pageId="3" pageNumber="41" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Quang Nam" typeStatus="paratype">
|
||
<typeStatus id="8112948EFFF73B46FC0F7FC90A7AFC37" box="[1006,1083,952,971]" pageId="3" pageNumber="41" type="paratype">Paratype</typeStatus>
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<specimenCount id="48AFE1A5FFF73B46FBA77FC40A13FC37" box="[1094,1106,949,971]" count="1" pageId="3" pageNumber="41" type="female">♀</specimenCount>
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||
(
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<collectionCode id="38B8B2E9FFF73B46FB857FC60AD7FC36" box="[1124,1174,951,970]" collectionName="SIEE" pageId="3" pageNumber="41">SIEE</collectionCode>
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), same data, except for ‘
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<geoCoordinate id="3B9D4CEBFFF73B46FCD17FBE0DDFFC1E" box="[816,926,975,994]" degrees="15" direction="north" minutes="33" orientation="latitude" pageId="3" pageNumber="41" precision="15" seconds="48" value="15.5633335">15°33´48´´N</geoCoordinate>
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/
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<geoCoordinate id="3B9D4CEBFFF73B46FC4C7FBE0A63FC1E" box="[941,1058,975,994]" degrees="107" direction="east" minutes="23" orientation="longitude" pageId="3" pageNumber="41" precision="15" seconds="22" value="107.38944">107°23´22´´E</geoCoordinate>
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, h=1050 m’
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="16B379A7FFF73B41FCB37F960C45FD8B" lastPageId="4" lastPageNumber="42" pageId="3" pageNumber="41" type="diagnosis">
|
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<paragraph id="5E162A2CFFF73B41FCB37F960C45FD8B" blockId="3.[812,1399,784,1181]" lastBlockId="4.[189,775,315,1111]" lastPageId="4" lastPageNumber="42" pageId="3" pageNumber="41">
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DIAGNOSIS. This species belongs to a species group defined by four derived characters, tarsomere 5 glabrous ventrally, frontal sulci very deep; pronotal lateral groove wide, deep and distinctly punctate, and elytral striae deep and punctate. The new species is distinguished from the other three species of the group,
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<taxonomicName id="99A951AFFFF73B46FBCC781C0A89FB7E" authorityName="Fedorenko" authorityYear="2018" box="[1069,1224,1133,1154]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="sulcatipennis">
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<emphasis id="6CDDF63EFFF73B46FBCC781C0A89FB7E" box="[1069,1224,1133,1154]" italics="true" pageId="3" pageNumber="41">P. sulcatipennis</emphasis>
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</taxonomicName>
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<bibRefCitation id="3A3857DDFFF73B46FB2D781C0B33FB7E" author="Fedorenko D. N." box="[1228,1394,1133,1155]" pageId="3" pageNumber="41" pagination="107 - 121" refId="ref13361" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">Fedorenko, 2018</bibRefCitation>
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||
;
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||
<taxonomicName id="99A951AFFFF73B46FCCD78F90DE3FB60" authorityName="Fedorenko" authorityYear="2018" box="[812,930,1159,1181]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="cavifrons">
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<emphasis id="6CDDF63EFFF73B46FCCD78F90DE3FB60" box="[812,930,1159,1181]" italics="true" pageId="3" pageNumber="41">P. cavifrons</emphasis>
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</taxonomicName>
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<bibRefCitation id="3A3857DDFFF73B46FC4978F60A0EFB60" author="Fedorenko D. N." box="[936,1103,1159,1181]" pageId="3" pageNumber="41" pagination="107 - 121" refId="ref13361" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">Fedorenko, 2018</bibRefCitation>
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||
; and
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||
<taxonomicName id="99A951AFFFF73B46FB6278F90B43FB60" authorityName="Fedorenko" authorityYear="2018" box="[1155,1282,1159,1181]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="alveolatus">
|
||
<emphasis id="6CDDF63EFFF73B46FB6278F90B43FB60" box="[1155,1282,1159,1181]" italics="true" pageId="3" pageNumber="41">P. alveolatus</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="3A3857DDFFF73B41FAE878F60EB1FEAC" author="Fedorenko D. N." lastPageId="4" lastPageNumber="42" pageId="3" pageNumber="41" pagination="107 - 121" refId="ref13361" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">Fedorenko, 2018</bibRefCitation>
|
||
, by the elytral striae very coarsely punctate, two (
|
||
<emphasis id="6CDDF63EFFF03B41FD0D7D4D0D46FEAD" box="[748,775,316,337]" italics="true" pageId="4" pageNumber="42">vs.</emphasis>
|
||
one) apical setae in interval 8, and a distinctive aedeagus, with very wide, securiform apex of median lobe. Besides,
|
||
<taxonomicName id="99A951AFFFF03B41FD137D000F7CFE5C" authorityName="Fedorenko" authorityYear="2018" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="sulcatipennis">
|
||
<emphasis id="6CDDF63EFFF03B41FD137D000F7CFE5C" italics="true" pageId="4" pageNumber="42">P. sulcatipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="99A951AFFFF03B41FE997DFD0FB5FE5C" authorityName="Fedorenko" authorityYear="2018" box="[376,500,395,417]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="cavifrons">
|
||
<emphasis id="6CDDF63EFFF03B41FE997DFD0FB5FE5C" box="[376,500,395,417]" italics="true" pageId="4" pageNumber="42">P. cavifrons</emphasis>
|
||
</taxonomicName>
|
||
are distinguishable in the pronotum not or indistinctly cordate, with sides not or barely sinuate in front of basal angles.
|
||
<taxonomicName id="99A951AFFFF03B41FDEB7DB00CEBFE2A" authorityName="Fedorenko" authorityYear="2018" box="[522,682,449,470]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="sulcatipennis">
|
||
<emphasis id="6CDDF63EFFF03B41FDEB7DB00CEBFE2A" box="[522,682,449,470]" italics="true" pageId="4" pageNumber="42">P. sulcatipennis</emphasis>
|
||
</taxonomicName>
|
||
is larger (
|
||
<collectionCode id="38B8B2E9FFF03B41FF227DAA0EA2FE0D" box="[195,227,475,497]" pageId="4" pageNumber="42">BL</collectionCode>
|
||
>
|
||
<quantity id="995187C9FFF03B41FF107DAA0F08FE0D" box="[241,329,475,497]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="4" pageNumber="42" unit="mm" value="15.8">15.8 mm</quantity>
|
||
), frontal sulci narrower and more shallow, and pronotal lateral groove narrower and finely punctate;
|
||
<taxonomicName id="99A951AFFFF03B41FD137D860F56FDDA" authorityName="Fedorenko" authorityYear="2018" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="cavifrons">
|
||
<emphasis id="6CDDF63EFFF03B41FD137D860F56FDDA" italics="true" pageId="4" pageNumber="42">P. cavifrons</emphasis>
|
||
</taxonomicName>
|
||
has pronotum smooth and mesotibia with a few additional setae just in front of one-row apical setal brush;
|
||
<taxonomicName id="99A951AFFFF03B41FD137E5D0F61FDA7" authorityName="Fedorenko" authorityYear="2018" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="alveolatus">
|
||
<emphasis id="6CDDF63EFFF03B41FD137E5D0F61FDA7" italics="true" pageId="4" pageNumber="42">P. alveolatus</emphasis>
|
||
</taxonomicName>
|
||
is distinctive in having elytral discal seta present (
|
||
<emphasis id="6CDDF63EFFF03B41FF247E100E98FD8A" box="[197,217,609,630]" italics="true" pageId="4" pageNumber="42">vs</emphasis>
|
||
. missing in the other species).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0AD67AA4FFF73B46FF037B200FD1F7ED" ID-DOI="http://doi.org/10.5281/zenodo.10978421" ID-Zenodo-Dep="10978421" httpUri="https://zenodo.org/record/10978421/files/figure.png" pageId="3" pageNumber="41" startId="3.[226,263,1873,1892]" targetBox="[192,1396,1226,1849]" targetPageId="3" targetType="figure">
|
||
<paragraph id="5E162A2CFFF73B46FF037B200F9EF851" blockId="3.[189,1399,1873,2065]" pageId="3" pageNumber="41">
|
||
Figs 10–13.
|
||
<taxonomicName id="99A951AFFFF73B46FEAF7B200C17F898" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[334,598,1873,1892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="hoii" status="sp. nov." subGenus="Tausternus">
|
||
<emphasis id="6CDDF63EFFF73B46FEAF7B200FFBF898" box="[334,442,1873,1892]" italics="true" pageId="3" pageNumber="41">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF73B46FE297B230C66F898" box="[456,551,1874,1892]" italics="true" pageId="3" pageNumber="41">Tausternus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF73B46FDD47B200C17F898" box="[565,598,1873,1892]" italics="true" pageId="3" pageNumber="41">hoii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF73B46FDBD7B200CC6F898" bold="true" box="[604,647,1873,1892]" pageId="3" pageNumber="41">
|
||
<taxonomicNameLabel id="77EE4B45FFF73B46FDBD7B200CC6F898" box="[604,647,1873,1892]" pageId="3" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 10–11 — median lobe of aedeagus; 12 — right paramere; 13 — female genitalia and reproductive tract (spermathecal gland and major part of spermatheca removed), ventral aspect; 10 — left lateral aspect; 11 — dorsal aspect;
|
||
<emphasis id="6CDDF63EFFF73B46FF5C7BF00E90F868" box="[189,209,1921,1940]" italics="true" pageId="3" pageNumber="41">bc</emphasis>
|
||
— bursa copulatrix;
|
||
<emphasis id="6CDDF63EFFF73B46FE6C7BF30FE8F868" box="[397,425,1922,1940]" italics="true" pageId="3" pageNumber="41">gsc</emphasis>
|
||
— gonosubcoxite; LTIX — laterotergite IX;
|
||
<emphasis id="6CDDF63EFFF73B46FCDF7BF30D13F868" box="[830,850,1922,1940]" italics="true" pageId="3" pageNumber="41">ov</emphasis>
|
||
— common oviduct;
|
||
<emphasis id="6CDDF63EFFF73B46FBF57BF00A6EF868" box="[1044,1071,1921,1940]" italics="true" pageId="3" pageNumber="41">sbs</emphasis>
|
||
— spermathecal basal sclerite;
|
||
<emphasis id="6CDDF63EFFF73B46FAA97BF30B1BF868" box="[1352,1370,1922,1940]" italics="true" pageId="3" pageNumber="41">sc</emphasis>
|
||
— seminal canal. Scale bars: 1 mm.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF73B46FF037BC70FD1F7ED" blockId="3.[189,1399,1873,2065]" pageId="3" pageNumber="41">
|
||
Рис. 10–13.
|
||
<taxonomicName id="99A951AFFFF73B46FEB07BC40C19F834" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[337,600,1973,1992]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="hoii" status="sp. nov." subGenus="Tausternus">
|
||
<emphasis id="6CDDF63EFFF73B46FEB07BC40FFDF834" box="[337,444,1973,1992]" italics="true" pageId="3" pageNumber="41">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF73B46FE2B7BC70C68F834" box="[458,553,1974,1992]" italics="true" pageId="3" pageNumber="41">Tausternus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF73B46FDD67BC40C19F834" box="[567,600,1973,1992]" italics="true" pageId="3" pageNumber="41">hoii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF73B46FDBE7BC40CCBF834" bold="true" box="[607,650,1973,1992]" pageId="3" pageNumber="41">
|
||
<taxonomicNameLabel id="77EE4B45FFF73B46FDBE7BC40CCBF834" box="[607,650,1973,1992]" pageId="3" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 10–11 — среднЯЯ долЯ Эдеагуса; 12 — праваЯ парамера; 13 — половой аппарат самки (больШаЯ часть сперматеки и желеЗа сперматеки удалены), вентрально; 10 — слева; 11 — дорЗально;
|
||
<emphasis id="6CDDF63EFFF73B46FB5E7BBC0A92F81C" box="[1215,1235,1997,2016]" italics="true" pageId="3" pageNumber="41">bc</emphasis>
|
||
— копулЯтивнаЯ сумка;
|
||
<emphasis id="6CDDF63EFFF73B46FF1F7B970F5AF804" box="[254,283,2022,2040]" italics="true" pageId="3" pageNumber="41">gsc</emphasis>
|
||
— гоносубкоксит; LTIX — латеротергит IX;
|
||
<emphasis id="6CDDF63EFFF73B46FD5F7B970C93F804" box="[702,722,2022,2040]" italics="true" pageId="3" pageNumber="41">ov</emphasis>
|
||
— обЩий Яйцевод;
|
||
<emphasis id="6CDDF63EFFF73B46FC6B7B940DE4F804" box="[906,933,2021,2040]" italics="true" pageId="3" pageNumber="41">sbs</emphasis>
|
||
— баЗальный склерит сперматеки;
|
||
<emphasis id="6CDDF63EFFF73B46FB0C7B970ABEF804" box="[1261,1279,2022,2040]" italics="true" pageId="3" pageNumber="41">sc</emphasis>
|
||
— семенной канал. МасШтаб: 1 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="16B379A7FFF03B43FF037E0A0FE8FE2A" lastPageId="6" lastPageNumber="44" pageId="4" pageNumber="42" type="description">
|
||
<paragraph id="5E162A2CFFF03B41FF037E0A0CB9FBFB" blockId="4.[189,775,315,1111]" pageId="4" pageNumber="42">
|
||
DESCRIPTION. BL
|
||
<quantity id="995187C9FFF03B41FE4C7E0A0C7AFD6D" box="[429,571,635,657]" metricMagnitude="-2" metricUnit="m" metricValue="1.265" metricValueMax="1.28" metricValueMin="1.25" pageId="4" pageNumber="42" unit="mm" value="12.65" valueMax="12.8" valueMin="12.5">12.5–12.8 mm</quantity>
|
||
. Body (
|
||
<figureCitation id="C69236A9FFF03B41FD637E0A0C86FD6D" box="[642,711,635,657]" captionStart="Figs 14–16" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[193,1367,1150,1943]" captionTargetId="figure-586@4.[189,1381,1138,1949]" captionTargetPageId="4" captionText="Figs 14–16. Dorsal habitus, holotypes: 14 — Pterostichus (Steropanus) securipenis sp.n; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n. Рис. 14–16. Габитус дорЗально, голотипы: 14 — Pterostichus (Steropanus) securipenis sp.n.; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n." figureDoi="http://doi.org/10.5281/zenodo.10978425" httpUri="https://zenodo.org/record/10978425/files/figure.png" pageId="4" pageNumber="42">Fig. 14</figureCitation>
|
||
) shiny black, with pronotal lateral groove and elytral striae dull due to coarse microsculpture. Legs red (
|
||
<typeStatus id="8112948EFFF03B41FDD77EC00CCDFD3B" box="[566,652,689,711]" pageId="4" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
) or reddish brown (
|
||
<typeStatus id="8112948EFFF03B41FEEA7EBA0F24FD1D" box="[267,357,715,737]" pageId="4" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
), antennae reddish brown, paler apically, maxillary and labial palps red. Dorsal microsculpture meshed, superficial isodiametric, more (
|
||
<typeStatus id="8112948EFFF03B41FE127F700C0BFCEB" box="[499,586,769,791]" pageId="4" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
) or less (
|
||
<typeStatus id="8112948EFFF03B41FD4B7F700D41FCEB" box="[682,768,769,791]" pageId="4" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
) distinct on head, distinct and moderately transverse on pronotum; pronotal lateral groove with coarse microsculpture, consisting of slightly longitudinal meshes. Elytral microsculpture superficial, consisting of narrow transverse meshes along middle of elytral interval (costa), turning into increasingly oblique meshes toward striae, and becoming coarse, slightly longitudinal, meshes at bottom of striae, with admixture of isodiametric meshes in strial punctures. Clypeus and pronotum densely or sparsely micropunctate, respectively.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF03B41FF03787A0ADEFE40" blockId="4.[189,775,315,1111]" lastBlockId="4.[812,1399,315,1111]" pageId="4" pageNumber="42">
|
||
Head: Eyes dorsal, small and prominent; gena about a third as long as eye, meeting neck at very obtuse angle. Frontal sulci smooth, diverging basad, in form of extremely deep, slightly oblong pits anteriorly, less deep behind, disappearing just posterior to the level of anterior supra-ocular seta. Supra-ocular groove very deep, straight anteriorly. Labrum subsinuate apically, apical setae equidistant
|
||
<emphasis id="6CDDF63EFFF03B41FAC47DFD0B31FE5D" box="[1317,1392,396,417]" italics="true" pageId="4" pageNumber="42">inter se</emphasis>
|
||
. Antennae just reaching pronotal base.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF03B41FCB37DB00AA9FDBD" blockId="4.[812,1399,315,1111]" pageId="4" pageNumber="42">Penultimate labial palpomere quadrisetose (two fixed setae at inner margin plus two additional latero-apical setae, dorsal and ventral), except for a minute apical seta at outer margin. Terminal labial palpomere triangular, a third as wide at apex as long at inner margin in both sexes.</paragraph>
|
||
<paragraph id="5E162A2CFFF03B41FCB37E370B50FBAB" blockId="4.[812,1399,315,1111]" pageId="4" pageNumber="42">
|
||
Pronotum cordate,
|
||
<collectionCode id="38B8B2E9FFF03B41FBF97E370A7DFDA7" box="[1048,1084,582,603]" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="42">PW</collectionCode>
|
||
/
|
||
<collectionCode id="38B8B2E9FFF03B41FBA57E370A21FDA7" box="[1092,1120,582,603]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="42" type="Museum">PL</collectionCode>
|
||
1.28–1.29,
|
||
<collectionCode id="38B8B2E9FFF03B41FB017E370B44FDA7" box="[1248,1285,582,603]" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="42">PW</collectionCode>
|
||
/HW 1.57– 1.58, broadest 1/3–2/5 from apex (PLw/
|
||
<collectionCode id="38B8B2E9FFF03B41FB387E100AB7FD8A" box="[1241,1270,609,630]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="42" type="Museum">PL</collectionCode>
|
||
0.33–0.39); sides evenly rounded, sinuate just in front of basal angles. Base sinuate medially, oblique towards obtuse and blunt basal angles, slightly wider than apex, PB/PA 1.11–1.13. Apex rather deeply sinuate, barely convex just inside apical angles, without apical bead; apical angles porrect, with blunt apices. Lateral bead entire, fairly thick all along, lateral groove deep, wide, nearly flat, moderately and densely punctate, distinctly wider than lateral bead in medial two quarters, extended inside base into a fine and deep line almost reaching inner basal sulcus. These sulci very deep, running parallel to each other in basal third, obliterate just in front of base; outer basal sulcus missing. Median line very deep, sulcate, crenulate at bottom, reaching base and almost reaching apex. Disc smooth, coarsely punctate at base, confluently within inner basal sulci, densely inside, with punctures not quite reaching median line; 1–2 punctures present outside basal sulcus. Anterolateral seta in lateral groove, posterolateral seta in basal angle.
|
||
</paragraph>
|
||
<caption id="0AD67AA4FFF03B41FF037BC00FF8F7EC" ID-DOI="http://doi.org/10.5281/zenodo.10978425" ID-Zenodo-Dep="10978425" httpUri="https://zenodo.org/record/10978425/files/figure.png" pageId="4" pageNumber="42" startId="4.[226,263,1969,1988]" targetBox="[193,1367,1150,1943]" targetPageId="4" targetType="figure">
|
||
<paragraph id="5E162A2CFFF03B41FF037BC00FC3F820" blockId="4.[189,1398,1969,2064]" pageId="4" pageNumber="42">
|
||
Figs 14–16. Dorsal habitus, holotypes: 14 —
|
||
<taxonomicName id="99A951AFFFF03B41FD927BC00DFCF838" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[627,957,1969,1988]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="securipenis" status="sp. nov." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF03B41FD927BC00C9FF838" box="[627,734,1969,1988]" italics="true" pageId="4" pageNumber="42">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF03B41FD0C7BC00D0FF838" box="[749,846,1969,1988]" italics="true" pageId="4" pageNumber="42">Steropanus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF03B41FCBA7BC30DFCF838" box="[859,957,1970,1988]" italics="true" pageId="4" pageNumber="42">securipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF03B41FC257BC00DA8F838" bold="true" box="[964,1001,1969,1988]" pageId="4" pageNumber="42">
|
||
<taxonomicNameLabel id="77EE4B45FFF03B41FC257BC00DA8F838" box="[964,1001,1969,1988]" pageId="4" pageNumber="42" rank="species">sp.n</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 15 —
|
||
<taxonomicName id="99A951AFFFF03B41FBCF7BC30B47F838" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1070,1286,1969,1988]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="pseudoglymmiger" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF03B41FBCF7BC30A03F838" box="[1070,1090,1970,1988]" italics="true" pageId="4" pageNumber="42">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF03B41FBAE7BC00A21F838" box="[1103,1120,1969,1988]" italics="true" pageId="4" pageNumber="42">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF03B41FB8C7BC00B47F838" box="[1133,1286,1969,1988]" italics="true" pageId="4" pageNumber="42">pseudoglymmiger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF03B41FAEC7BC00B79F838" bold="true" box="[1293,1336,1969,1988]" pageId="4" pageNumber="42">
|
||
<taxonomicNameLabel id="77EE4B45FFF03B41FAEC7BC00B79F838" box="[1293,1336,1969,1988]" pageId="4" pageNumber="42" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 16 —
|
||
<taxonomicName id="99A951AFFFF03B41FF5C7BBB0F11F820" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[189,336,1993,2012]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="fossifrons" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF03B41FF5C7BBB0E91F820" box="[189,208,1994,2012]" italics="true" pageId="4" pageNumber="42">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF03B41FF3F7BB80EAEF820" box="[222,239,1993,2012]" italics="true" pageId="4" pageNumber="42">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF03B41FF1D7BB80F11F820" box="[252,336,1993,2012]" italics="true" pageId="4" pageNumber="42">fossifrons</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF03B41FEB67BB80FC3F820" bold="true" box="[343,386,1993,2012]" pageId="4" pageNumber="42">
|
||
<taxonomicNameLabel id="77EE4B45FFF03B41FEB67BB80FC3F820" box="[343,386,1993,2012]" pageId="4" pageNumber="42" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF03B41FF037B940FF8F7EC" blockId="4.[189,1398,1969,2064]" pageId="4" pageNumber="42">
|
||
Рис. 14–16. Габитус дорЗально, голотипы: 14 —
|
||
<taxonomicName id="99A951AFFFF03B41FD467B940DB0F804" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[679,1009,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="securipenis" status="sp. nov." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF03B41FD467B940D53F804" box="[679,786,2021,2040]" italics="true" pageId="4" pageNumber="42">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF03B41FCC07B940DC3F804" box="[801,898,2021,2040]" italics="true" pageId="4" pageNumber="42">Steropanus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF03B41FC6E7B970DB0F804" box="[911,1009,2022,2040]" italics="true" pageId="4" pageNumber="42">securipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF03B41FC167B940A63F804" bold="true" box="[1015,1058,2021,2040]" pageId="4" pageNumber="42">
|
||
<taxonomicNameLabel id="77EE4B45FFF03B41FC167B940A63F804" box="[1015,1058,2021,2040]" pageId="4" pageNumber="42" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 15 —
|
||
<taxonomicName id="99A951AFFFF03B41FB867B970B7EF804" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1127,1343,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="pseudoglymmiger" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF03B41FB867B970A3AF804" box="[1127,1147,2022,2040]" italics="true" pageId="4" pageNumber="42">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF03B41FB697B940AD8F804" box="[1160,1177,2021,2040]" italics="true" pageId="4" pageNumber="42">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF03B41FB477B940B7EF804" box="[1190,1343,2021,2040]" italics="true" pageId="4" pageNumber="42">pseudoglymmiger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF03B41FAA47B940B31F804" bold="true" box="[1349,1392,2021,2040]" pageId="4" pageNumber="42">
|
||
<taxonomicNameLabel id="77EE4B45FFF03B41FAA47B940B31F804" box="[1349,1392,2021,2040]" pageId="4" pageNumber="42" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 16 —
|
||
<taxonomicName id="99A951AFFFF03B41FF147B8F0FC9F7EC" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[245,392,2045,2064]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="fossifrons" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF03B41FF147B8F0F49F7EC" box="[245,264,2046,2064]" italics="true" pageId="4" pageNumber="42">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF03B41FEF47B8C0F66F7EC" box="[277,295,2045,2064]" italics="true" pageId="4" pageNumber="42">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF03B41FED57B8C0FC9F7EC" box="[308,392,2045,2064]" italics="true" pageId="4" pageNumber="42">fossifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="77EE4B45FFF03B41FE6F7B8C0FF8F7EC" box="[398,441,2045,2064]" pageId="4" pageNumber="42" rank="species">sp.n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5E162A2CFFF13B40FF037D4A0AE1FE97" blockId="5.[189,775,315,737]" lastBlockId="5.[812,1399,315,737]" pageId="5" pageNumber="43">
|
||
Elytra connate, elliptic, broadest behind middle, EW/EL 1.59–1.60, EW/
|
||
<collectionCode id="38B8B2E9FFF13B40FEB87D270F3CFE97" box="[345,381,342,363]" country="Thailand" name="Paleontological Collections" pageId="5" pageNumber="43">PW</collectionCode>
|
||
1.19–1.20, without preapical sinuation or internal plica; apices contiguous, apex otherwise slightly trilobed due to summit of apically confluent costae 2 to 7 projecting slightly apicad. Base narrow and oblique, humeri rounded, humeral tooth blunt and only traceable in posterodorsal view; basal ridge outside stria 3, with a vestige between striae 4 and 5; humeral angle very obtuse. Striae 1–9 deep, very coarsely punctate, in apical third with punctures increasingly deep and large apicad; striae 1 and 7 with apical two punctures in form of large and very deep foveae; striae 1– 6 adjoining basal ridge. Parascutellar striole missing. Intervals costate, 7
|
||
<superScript id="A9DC8764FFF13B40FEB07E0A0F1DFD7B" attach="none" box="[337,348,635,647]" fontSize="5" pageId="5" pageNumber="43">th</superScript>
|
||
, 5
|
||
<superScript id="A9DC8764FFF13B40FE9A7E0A0FC7FD7B" attach="none" box="[379,390,635,647]" fontSize="5" pageId="5" pageNumber="43">th</superScript>
|
||
, 3
|
||
<superScript id="A9DC8764FFF13B40FE447E0A0FF0FD7B" attach="right" box="[421,433,635,647]" fontSize="5" pageId="5" pageNumber="43">rd</superScript>
|
||
, 2
|
||
<superScript id="A9DC8764FFF13B40FE2E7E0A0F9CFD7B" attach="left" box="[463,477,635,647]" fontSize="5" pageId="5" pageNumber="43">nd</superScript>
|
||
and 1st confluent apicad in succession and then merging into a vanished lateral bead; intervals 8 and 9 vanished before apex. Reflexed lateral margin subequally narrow throughout its length. Parascutellar setigerous pore at base of stria 2, discal setae missing, two preapical setae in interval 8; USS: 18.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF13B40FCB37D000A1EFDA7" blockId="5.[812,1399,315,737]" pageId="5" pageNumber="43">Underside. Prosternum with a fairly deep median groove terminating in a very deep pit 2/5 from apical margin. Prosternal process in ventral view truncate, with fairly sharp angles; in lateral view rectangular and blunt; inclination wide, rounded in posterior view, slightly oblique and concave in lateral view. Abdominal lateral bead entire and conspicuous; sternite VII similar in both sexes. Mesepisternum coarsely and rather densely punctate in anterior half; metepisternum smooth or with a few coarse punctures.</paragraph>
|
||
<paragraph id="5E162A2CFFF13B43FCB37E100C49FE90" blockId="5.[812,1399,315,737]" lastBlockId="6.[189,776,315,951]" lastPageId="6" lastPageNumber="44" pageId="5" pageNumber="43">Legs: profemur posteriorly trisetose, protibia barely dilated apicad, with three spinules at posterolateral edge, preapical spinule separate from two apical ones. Tarsi laterally neither carinate nor sulcate; tarsomere 5 glabrous ventrally. Protarsomeres 1–3 dilated and biseriately squamose ventrally in male; strongly dentate at apical angles in female. Mesotibia with a one-row apical setal brush.</paragraph>
|
||
<caption id="0AD67AA4FFF13B40FF037BF00C12F7ED" ID-DOI="http://doi.org/10.5281/zenodo.10978427" ID-Zenodo-Dep="10978427" httpUri="https://zenodo.org/record/10978427/files/figure.png" pageId="5" pageNumber="43" startId="5.[226,262,1921,1940]" targetBox="[194,1391,758,1882]" targetPageId="5" targetType="figure">
|
||
<paragraph id="5E162A2CFFF13B40FF037BF00CC5F839" blockId="5.[188,1399,1921,2065]" pageId="5" pageNumber="43">
|
||
Figs 17–22. Aedeagus of
|
||
<taxonomicName id="99A951AFFFF13B40FE5E7BF00CDDF868" authorityName="Fairmaire" authorityYear="1889" box="[447,668,1921,1941]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="subGenus" subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FE5E7BF00C69F868" box="[447,552,1921,1940]" italics="true" pageId="5" pageNumber="43">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FDD77BF30CD4F869" box="[566,661,1922,1941]" italics="true" pageId="5" pageNumber="43">Steropanus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.: 17–18 —
|
||
<taxonomicName id="99A951AFFFF13B40FCC97BF30DF6F868" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[808,951,1921,1941]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="fossifrons" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FCC97BF30D7AF868" box="[808,827,1922,1940]" italics="true" pageId="5" pageNumber="43">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FCA97BF30D18F869" box="[840,857,1922,1941]" italics="true" pageId="5" pageNumber="43">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF13B40FC847BF00DF6F868" box="[869,951,1921,1940]" italics="true" pageId="5" pageNumber="43">fossifrons</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF13B40FC5F7BF00DA9F868" bold="true" box="[958,1000,1921,1940]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="77EE4B45FFF13B40FC5F7BF00DA9F868" box="[958,1000,1921,1940]" pageId="5" pageNumber="43" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 19–20 —
|
||
<taxonomicName id="99A951AFFFF13B40FBAA7BF30AA8F868" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1099,1257,1921,1941]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="securipenis" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FBAA7BF30A1EF868" box="[1099,1119,1922,1940]" italics="true" pageId="5" pageNumber="43">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FB8A7BF30A3DF869" box="[1131,1148,1922,1941]" italics="true" pageId="5" pageNumber="43">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF13B40FB687BF30AA8F868" box="[1161,1257,1922,1940]" italics="true" pageId="5" pageNumber="43">securipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF13B40FB0E7BF00B58F868" bold="true" box="[1263,1305,1921,1940]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="77EE4B45FFF13B40FB0E7BF00B58F868" box="[1263,1305,1921,1940]" pageId="5" pageNumber="43" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 21–22 —
|
||
<taxonomicName id="99A951AFFFF13B40FF5D7BEB0FD5F850" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[188,404,1945,1965]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="pseudoglymmiger" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FF5D7BEB0E91F850" box="[188,208,1946,1964]" italics="true" pageId="5" pageNumber="43">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FF3C7BEB0EAFF851" box="[221,238,1946,1965]" italics="true" pageId="5" pageNumber="43">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF13B40FF1A7BE80FD5F850" box="[251,404,1945,1964]" italics="true" pageId="5" pageNumber="43">pseudoglymmiger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF13B40FE7B7BE80F84F850" bold="true" box="[410,453,1945,1964]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="77EE4B45FFF13B40FE7B7BE80F84F850" box="[410,453,1945,1964]" pageId="5" pageNumber="43" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, median lobe with everted and inflated internal sac; 17–20 — median lobe; 17, 19, 21 — left lateral aspect; 18, 20, 22 — right lateral aspect. Scale bars: 1 mm.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF13B40FF037BBF0C12F7ED" blockId="5.[188,1399,1921,2065]" pageId="5" pageNumber="43">
|
||
Рис. 17–22. Эдеагус
|
||
<taxonomicName id="99A951AFFFF13B40FE427BBC0CC4F81C" authorityName="Fairmaire" authorityYear="1889" box="[419,645,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="subGenus" subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FE427BBC0C4EF81C" box="[419,527,1997,2016]" italics="true" pageId="5" pageNumber="43">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FDFC7BBC0C3FF81C" box="[541,638,1997,2016]" italics="true" pageId="5" pageNumber="43">Steropanus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.: 17–18 —
|
||
<taxonomicName id="99A951AFFFF13B40FCF77BBF0DEBF81C" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[790,938,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="fossifrons" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FCF77BBF0D68F81C" box="[790,809,1998,2016]" italics="true" pageId="5" pageNumber="43">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FCD67BBC0D09F81C" box="[823,840,1997,2016]" italics="true" pageId="5" pageNumber="43">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF13B40FCB77BBC0DEBF81C" box="[854,938,1997,2016]" italics="true" pageId="5" pageNumber="43">fossifrons</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF13B40FC507BBC0D9DF81C" bold="true" box="[945,988,1997,2016]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="77EE4B45FFF13B40FC507BBC0D9DF81C" box="[945,988,1997,2016]" pageId="5" pageNumber="43" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 19–20 —
|
||
<taxonomicName id="99A951AFFFF13B40FBA37BBF0AA5F81C" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1090,1252,1997,2016]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="securipenis" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FBA37BBF0A17F81C" box="[1090,1110,1998,2016]" italics="true" pageId="5" pageNumber="43">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FB857BBC0A34F81C" box="[1124,1141,1997,2016]" italics="true" pageId="5" pageNumber="43">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF13B40FB637BBF0AA5F81C" box="[1154,1252,1998,2016]" italics="true" pageId="5" pageNumber="43">securipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF13B40FB0A7BBC0B57F81C" bold="true" box="[1259,1302,1997,2016]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="77EE4B45FFF13B40FB0A7BBC0B57F81C" box="[1259,1302,1997,2016]" pageId="5" pageNumber="43" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 21–22 —
|
||
<taxonomicName id="99A951AFFFF13B40FF5D7B970FD5F804" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[188,404,2021,2040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="pseudoglymmiger" status="sp.n." subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF13B40FF5D7B970E91F804" box="[188,208,2022,2040]" italics="true" pageId="5" pageNumber="43">P.</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF13B40FF3D7B940EACF804" box="[220,237,2021,2040]" italics="true" pageId="5" pageNumber="43">S.</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF13B40FF1A7B940FD5F804" box="[251,404,2021,2040]" italics="true" pageId="5" pageNumber="43">pseudoglymmiger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF13B40FE7A7B940F87F804" bold="true" box="[411,454,2021,2040]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="77EE4B45FFF13B40FE7A7B940F87F804" box="[411,454,2021,2040]" pageId="5" pageNumber="43" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 17–20 — среднЯЯ долЯ; 17, 19, 21 — слева; 18, 20, 22 — справа. МасШтаб: 1 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5E162A2CFFF23B43FF037D000FE8FE2A" blockId="6.[189,776,315,951]" pageId="6" pageNumber="44">
|
||
Aedeagus (
|
||
<figureCitation id="C69236A9FFF23B43FEB27D000F89FE7A" box="[339,456,369,391]" captionStart="Figs 17–22" captionStartId="5.[226,262,1921,1940]" captionTargetBox="[194,1391,758,1882]" captionTargetId="figure-432@5.[189,1399,755,1886]" captionTargetPageId="5" captionText="Figs 17–22. Aedeagus of Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 17–20 — median lobe; 17, 19, 21 — left lateral aspect; 18, 20, 22 — right lateral aspect. Scale bars: 1 mm. Рис. 17–22. Эдеагус Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 17–20 — среднЯЯ долЯ; 17, 19, 21 — слева; 18, 20, 22 — справа. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978427" httpUri="https://zenodo.org/record/10978427/files/figure.png" pageId="6" pageNumber="44">Figs 19–20</figureCitation>
|
||
,
|
||
<figureCitation id="C69236A9FFF23B43FE377D000C5DFE7B" box="[470,540,369,391]" captionStart="Figs 23–29" captionStartId="6.[226,263,1921,1940]" captionTargetBox="[192,1364,1003,1906]" captionTargetId="figure-533@6.[189,1369,999,1909]" captionTargetPageId="6" captionText="Figs 23–29. Aedeagus of Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (apical margin slightly damaged); 25, 28 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — median lobe, ventral aspect; 26–28 — inside of right paramere. Scale bars: 1 mm. Рис. 23–29. Эдеагус Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (апикальный край слегка обломан); 25, 28 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — среднЯЯ долЯ? вентрально; 26–28 — праваЯ парамера, иЗнутри. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978429" httpUri="https://zenodo.org/record/10978429/files/figure.png" pageId="6" pageNumber="44">24, 27</figureCitation>
|
||
): Median lobe arcuate. Apex conspicuously bent ventrad, oblique and very wide. Right paramere moderately long, subtriangular, with a narrow parallel-sided apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF23B43FF037DAA0F4AFDDB" pageId="6" pageNumber="44" type="distribution">
|
||
<paragraph id="5E162A2CFFF23B43FF037DAA0CB3FE0D" blockId="6.[189,776,315,951]" box="[226,754,475,497]" pageId="6" pageNumber="44">
|
||
DISTRIBUTION. Known from the
|
||
<typeStatus id="8112948EFFF23B43FDA17DAD0C2BFE0D" box="[576,618,476,497]" pageId="6" pageNumber="44">type</typeStatus>
|
||
locality only.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF23B43FF037D870F4AFDDB" blockId="6.[189,776,315,951]" pageId="6" pageNumber="44">
|
||
<collectionCode id="38B8B2E9FFF23B43FF037D870F6DFDF7" box="[226,300,502,523]" pageId="6" pageNumber="44">NAME</collectionCode>
|
||
. Refers to the securiform apex of aedeagus median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF23B43FF037E5A0F1AFD8B" pageId="6" pageNumber="44" type="biology_ecology">
|
||
<paragraph id="5E162A2CFFF23B43FF037E5A0F1AFD8B" blockId="6.[189,776,315,951]" pageId="6" pageNumber="44">
|
||
HABITATS AND HABITS.
|
||
<typeStatus id="8112948EFFF23B43FE1C7E5A0C16FDBD" box="[509,599,555,577]" pageId="6" pageNumber="44" type="holotype">Holotype</typeStatus>
|
||
was hand collected and
|
||
<typeStatus id="8112948EFFF23B43FEE67E370F1BFDA7" box="[263,346,582,603]" pageId="6" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
taken by pitfall trapping in a broad-leaved monsoon forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF23B43FF037E0A0C70FC4B" pageId="6" pageNumber="44" type="discussion">
|
||
<paragraph id="5E162A2CFFF23B43FF037E0A0C70FC4B" blockId="6.[189,776,315,951]" pageId="6" pageNumber="44">
|
||
COMMENTS. Apical setal brush of the mesotibia has not been mentioned in the redescription of
|
||
<taxonomicName id="99A951AFFFF23B43FD767EE70D46FD57" authorityName="Andrewes" authorityYear="1937" box="[663,775,662,683]" pageId="6" pageNumber="52" rank="subGenus" subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF23B43FD767EE70D46FD57" box="[663,775,662,683]" italics="true" pageId="6" pageNumber="44">Steropanus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<bibRefCitation id="3A3857DDFFF23B43FF237EC00F34FD3A" author="Fedorenko D. N." box="[194,373,688,711]" pageId="6" pageNumber="44" pagination="107 - 121" refId="ref13361" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">
|
||
<taxonomicName id="99A951AFFFF23B43FF237EC00F70FD3B" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[194,305,689,711]" pageId="6" pageNumber="39" rank="subGenus" status="subgen. nov." subGenus="Fedorenko">Fedorenko</taxonomicName>
|
||
, 2018
|
||
</bibRefCitation>
|
||
]. It is certain that the originally transverse and one-row setal brush (
|
||
<taxonomicName id="99A951AFFFF23B43FDEE7EBD0CE0FD1D" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[527,673,716,737]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="securipenis" status="sp.n.">
|
||
<emphasis id="6CDDF63EFFF23B43FDEE7EBD0CE0FD1D" box="[527,673,716,737]" italics="true" pageId="6" pageNumber="44">P. securipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF23B43FD4C7EBA0C9EFD1C" bold="true" box="[685,735,715,736]" pageId="6" pageNumber="44">
|
||
<taxonomicNameLabel id="77EE4B45FFF23B43FD4C7EBA0C9EFD1C" box="[685,735,715,736]" pageId="6" pageNumber="44" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="99A951AFFFF23B43FD107EBD0F7EFD07" authorityName="Fedorenko" authorityYear="2018" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="sulcatipennis">
|
||
<emphasis id="6CDDF63EFFF23B43FD107EBD0F7EFD07" italics="true" pageId="6" pageNumber="44">P. sulcatipennis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="99A951AFFFF23B43FEAC7E960C4DFD07" authorityName="Fedorenko" authorityYear="2018" box="[333,524,742,764]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="pseudoviolaceus">
|
||
<emphasis id="6CDDF63EFFF23B43FEAC7E960C4DFD07" box="[333,524,742,764]" italics="true" pageId="6" pageNumber="44">P. pseudoviolaceus</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="3A3857DDFFF23B43FDF57E970CFEFD07" author="Fedorenko D. N." box="[532,703,742,764]" pageId="6" pageNumber="44" pagination="107 - 121" refId="ref13361" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">Fedorenko, 2018</bibRefCitation>
|
||
,
|
||
<taxonomicName id="99A951AFFFF23B43FD287E960F4EFCEA" authorityName="Fedorenko" authorityYear="2018" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="obliteratus">
|
||
<emphasis id="6CDDF63EFFF23B43FD287E960F4EFCEA" italics="true" pageId="6" pageNumber="44">P. obliteratus</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="3A3857DDFFF23B43FEF67F700F85FCEA" author="Fedorenko D. N." box="[279,452,769,791]" pageId="6" pageNumber="44" pagination="107 - 121" refId="ref13361" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">Fedorenko, 2018</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="99A951AFFFF23B43FE1D7F700C38FCEA" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[508,633,769,790]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="fossifrons" status="sp.n.">
|
||
<emphasis id="6CDDF63EFFF23B43FE1D7F700C38FCEA" box="[508,633,769,790]" italics="true" pageId="6" pageNumber="44">P. fossifrons</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF23B43FD637F700CF5FCEA" bold="true" box="[642,692,769,790]" pageId="6" pageNumber="44">
|
||
<taxonomicNameLabel id="77EE4B45FFF23B43FD637F700CF5FCEA" box="[642,692,769,790]" pageId="6" pageNumber="44" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) at first evolved into that with a few additional setae appeared just above it in
|
||
<taxonomicName id="99A951AFFFF23B43FEC97F460C24FCB0" authority="(Andrewes, 1937)" baseAuthorityName="Andrewes" baseAuthorityYear="1937" box="[296,613,822,844]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="glymmiger">
|
||
<emphasis id="6CDDF63EFFF23B43FEC97F460FEDFCB7" box="[296,428,822,844]" italics="true" pageId="6" pageNumber="44">P. glymmiger</emphasis>
|
||
(
|
||
<bibRefCitation id="3A3857DDFFF23B43FE597F470C1CFCB0" author="Andrewes H. E." box="[440,605,822,844]" pageId="6" pageNumber="44" pagination="489 - 495" refId="ref13093" refString="Andrewes H. E. 1937. Papers on Oriental Carabidae. - XXXI // Ann. Mag. Nat. Hist. Ser. 9. Vol. 19. P. 489 - 495." type="journal article" year="1937">Andrewes, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="99A951AFFFF23B43FD8F7F460EB7FC9A" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="pseudoglymmiger" status="sp.n.">
|
||
<emphasis id="6CDDF63EFFF23B43FD8F7F460EB7FC9A" italics="true" pageId="6" pageNumber="44">P. pseudoglymmiger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF23B43FF1F7F200F6EFC9A" bold="true" box="[254,303,849,870]" pageId="6" pageNumber="44">
|
||
<taxonomicNameLabel id="77EE4B45FFF23B43FF1F7F200F6EFC9A" box="[254,303,849,870]" pageId="6" pageNumber="44" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="99A951AFFFF23B43FE887F200FA3FC9A" authorityName="Fedorenko" authorityYear="2018" box="[361,482,849,870]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="cavifrons">
|
||
<emphasis id="6CDDF63EFFF23B43FE887F200FA3FC9A" box="[361,482,849,870]" italics="true" pageId="6" pageNumber="44">P. cavifrons</emphasis>
|
||
</taxonomicName>
|
||
, and then into triangular and plurisetose setal brush in
|
||
<taxonomicName id="99A951AFFFF23B43FE247F1D0CA5FC7D" authority="(Andrewes 1937)" baseAuthorityName="Andrewes" baseAuthorityYear="1937" box="[453,740,875,897]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="aequus">
|
||
<emphasis id="6CDDF63EFFF23B43FE247F1D0C68FC7D" box="[453,553,876,897]" italics="true" pageId="6" pageNumber="44">P. aequus</emphasis>
|
||
(
|
||
<bibRefCitation id="3A3857DDFFF23B43FDD87F1A0C9DFC7D" author="Andrewes H. E." box="[569,732,875,897]" pageId="6" pageNumber="44" pagination="489 - 495" refId="ref13093" refString="Andrewes H. E. 1937. Papers on Oriental Carabidae. - XXXI // Ann. Mag. Nat. Hist. Ser. 9. Vol. 19. P. 489 - 495." type="journal article" year="1937">Andrewes 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="99A951AFFFF23B43FD107F1D0F5BFC67" authorityName="Fedorenko" authorityYear="2018" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="asulcatus">
|
||
<emphasis id="6CDDF63EFFF23B43FD107F1D0F5BFC67" italics="true" pageId="6" pageNumber="44">P. asulcatus</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="3A3857DDFFF23B43FEC07FF70F8CFC67" author="Fedorenko D. N." box="[289,461,902,924]" pageId="6" pageNumber="44" pagination="107 - 121" refId="ref13361" refString="Fedorenko D. N. 2018. Notes on Pterostichus subgenus Steropanus (Coleoptera: Carabidae: Pterostichini), with descriptions of new species // Russian Entomol. J. Vol. 27. No. 2. P. 107 - 121. doi: 10.15298 / rusentj. 27.2.01." type="journal article" year="2018">Fedorenko, 2018</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="99A951AFFFF23B43FDE27FF60CDBFC67" authorityName="Fedorenko" authorityYear="2020" box="[515,666,902,924]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="boriskataevi" status="nom.n.">
|
||
<emphasis id="6CDDF63EFFF23B43FDE27FF60CDBFC67" box="[515,666,902,924]" italics="true" pageId="6" pageNumber="44">P. boriskataevi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF23B43FD437FF70CABFC67" bold="true" box="[674,746,902,923]" pageId="6" pageNumber="44">
|
||
<taxonomicNameLabel id="77EE4B45FFF23B43FD437FF70CABFC67" box="[674,746,902,923]" pageId="6" pageNumber="44">nom.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
as the additional setae became multiple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |