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<mods:title id="93225494409F08FCDE711EEFB4D16597">First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995</mods:title>
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<mods:namePart id="121B9AF18E5FE66E024B306C358E107E">Rivera-Quiroz, Francisco Andres</mods:namePart>
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<mods:affiliation id="99C7E39EB43B218E000B7D8BA5E05B0C">Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands &amp; Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier id="1D7C76280854F5A0051A44CBFF637AF3" type="email">andres.riveraquiroz@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="8569AE3BD07BB08518FC555770A1A889">Petcharad, Booppa</mods:namePart>
<mods:affiliation id="4F01236CCB82CFCFBCEF5279C2F9871C">Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, the Netherlands</mods:affiliation>
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<mods:namePart id="03ADBE5B29D2F1ECAD4FFDDD82C27E59">Miller, Jeremy A.</mods:namePart>
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<mods:affiliation id="49D5B3C074A2B96A7CC499B43B5A76E9">Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands</mods:affiliation>
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<treatment id="D1FCD9D1AC07541DB881DFE02A1C1BD7" ID-GBIF-Taxon="177132022" LSID="urn:lsid:plazi:treatment:D1FCD9D1AC07541DB881DFE02A1C1BD7" httpUri="http://treatment.plazi.org/id/D1FCD9D1AC07541DB881DFE02A1C1BD7" lastPageNumber="21" pageId="0" pageNumber="21" scope_class="Arachnida" scope_family="Symphytognathidae" scope_order="Araneae">
<subSubSection id="8CF5F39997C7023E53F222BB58624DA6" pageId="0" pageNumber="21" type="nomenclature">
<paragraph id="EE1CC0986FC7AFF75CB76D9825EFC9D3" pageId="0" pageNumber="21">
<taxonomicName id="21B71287D00228534DD10AEB53E91A12" ID-CoL="6TTRS" LSID="D1FCD9D1-AC07-541D-B881-DFE02A1C1BD7" authority="Lin &amp; Li, 2009" authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">Patu shiluensis Lin &amp; Li, 2009</taxonomicName>
<figureCitation id="03FA58BC96411DFF7CF275BB998A06A3" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">Figures 14</figureCitation>
<figureCitation id="0F58C5AC9BDF28FED0E46B40ECA76E0C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">, 15a, c</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0913A900B842A51943FFA547DAF61545" pageId="0" pageNumber="21" type="reference_group">
<paragraph id="931F32283E45A54670DA2AD4E7E9FB2E" pageId="0" pageNumber="21">
<taxonomicName id="28DF1CBF0E25081A8CF2302CB9CABC4A" authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">Patu shiluensis</taxonomicName>
Lin &amp; Li, 2009: 59, figs 11A, B, 12A, B, 13A-D.
</paragraph>
</subSubSection>
<subSubSection id="93347AF5435458A13E71E0ECF1FC282F" pageId="0" pageNumber="21" type="materials_examined">
<paragraph id="A911F5BE0DAA16CC21429E368D5EF12C" pageId="0" pageNumber="21">Collected material.</paragraph>
<paragraph id="0CA63BDAD1E1ABE1773FB95687BF339D" pageId="0" pageNumber="21">
Thailand • 4♀; Phuket Province, Siray Island. Mixed tropical forest;
<geoCoordinate id="464B9B99DCFE66193604A904581B21D5" degrees="7" direction="north" minutes="53.355" orientation="latitude" precision="1" value="7.88925">7°53.355'N</geoCoordinate>
,
<geoCoordinate id="E6C60A8C681B4BFEE86E2431327EC476" degrees="98" direction="east" minutes="26.083" orientation="longitude" precision="1" value="98.434715">98°26.083'E</geoCoordinate>
. 132 m; August 02-06, 2018; Booppa Petcharad, Jeremy Miller, F. Andres Rivera-Quiroz leg.; Winkler extractor; RMNH.5106642.
</paragraph>
</subSubSection>
<subSubSection id="26D6A76DFF717CEAB0674BB658B2B6B2" pageId="0" pageNumber="21" type="distribution">
<paragraph id="7C21DD642B8B38149C161E024377568D" pageId="0" pageNumber="21">Distribution.</paragraph>
<paragraph id="8A9B7C516CEB9FF1497A61ECF419E640" pageId="0" pageNumber="21">Known only from its type locality, Shilu Town, Hainan Province, China and the specimens collected for the present work.</paragraph>
</subSubSection>
<subSubSection id="8C235D60DF462C20D2175FF4DE34653E" pageId="0" pageNumber="21" type="morphological remarks">
<paragraph id="3A69AC305B9DB2F124B48A19D5460633" pageId="0" pageNumber="21">Morphological remarks.</paragraph>
<paragraph id="CC11FC4CA17C72BB66DEA5D5D148BDC6" pageId="0" pageNumber="21">
Carapace pale yellow with black margin, smooth texture (Fig.
<figureCitation id="C7C7B8059CC6EC4662B1020F1D9B89FE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14b</figureCitation>
). Legs black and semi-transparent. Abdomen oval, longer than wide (Fig.
<figureCitation id="282B45F06251BBE62D34775AA718B5D5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14a, b</figureCitation>
). Ventrally same color as carapace, dorsally, darker with pale yellow patches.
</paragraph>
<caption id="56CFC6F10B313996B15F7E2957B31DA7" doi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21" start="Figure 14" startId="F14">
<paragraph id="2869F66EBCDB0CEFA2E7A91011806EF4" pageId="0" pageNumber="21">
<emphasis id="37D44CFCBF77A777A60010C5FD4B1179" bold="true" pageId="0" pageNumber="21">Figure 14.</emphasis>
<taxonomicName id="329D259201052C7F6F010680671EFE22" authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">
<emphasis id="D946A70106FCF74AE34263DA43993F0D" italics="true" pageId="0" pageNumber="21">Patu shiluensis</emphasis>
</taxonomicName>
Lin &amp; Li, 2009 female: Habitus:
<emphasis id="AFF77C5669A4DA0320C78849432C9FB1" bold="true" pageId="0" pageNumber="21">a</emphasis>
ventral view
<emphasis id="205985FBA88B89A7114B85049C39EAF2" bold="true" pageId="0" pageNumber="21">b</emphasis>
dorsal view. Epigynum:
<emphasis id="A3E581702803460E71E6D09559366D41" bold="true" pageId="0" pageNumber="21">c</emphasis>
ventral view
<emphasis id="30F0D35CE62B6D3F5A69EDBF2EE1E996" bold="true" pageId="0" pageNumber="21">d</emphasis>
dorsal view, cleared. Scale bars: 0.2 mm (
<emphasis id="AA40764FEF1C23579A3EB13A15FAC6D7" bold="true" pageId="0" pageNumber="21">a, b</emphasis>
); 0.06 mm (
<emphasis id="81B2CCAAAD4762D7533410C31CB644C8" bold="true" pageId="0" pageNumber="21">c</emphasis>
); 0.03 mm (
<emphasis id="F900282AE9ED78EA05092F38196D145F" bold="true" pageId="0" pageNumber="21">d</emphasis>
).
</paragraph>
</caption>
<paragraph id="1A397BF3B4D058F600D637C79C29F0C6" pageId="0" pageNumber="21">
<emphasis id="769A9A5F814198E144B40A8FC80C60C4" bold="true" italics="true" pageId="0" pageNumber="21">Vulva</emphasis>
: Epigynum weakly sclerotized, semi-transparent (Fig.
<figureCitation id="D45F4942D64DB7FC6354B29472CB1CB4" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14c</figureCitation>
). Atrium semi-circular slightly wider than inner distance between S (Figs
<figureCitation id="674C058CB325C7CDBFB9B38B03363715" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14c</figureCitation>
,
<figureCitation id="3BED0478587CD252635D461099775CCC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">15c</figureCitation>
). Spermathecae spherical slightly more sclerotized than epigynum, separated by 0.5
<normalizedToken id="54A87038658C6F1E9B07AE7A33FBB2F7" originalValue="×">x</normalizedToken>
their diameter (Fig.
<figureCitation id="E1BF1935D875A22DC1768435AFCB59BC" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14d</figureCitation>
). Copulatory ducts spring-like, spiraling 3
<normalizedToken id="75E812C867E96E86137FE43879C06F7D" originalValue="×">x</normalizedToken>
over themselves. Fertilization ducts as long as S width, projecting posteriorly (Figs
<figureCitation id="7CB349C9AB766E37ADBB1825CC8C3422" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14d</figureCitation>
,
<figureCitation id="5613157B44827A597466C8B5ACE14B38" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">15a, c</figureCitation>
).
</paragraph>
<caption id="4F79BB8254AD1358100C8D4EB1CFA9B0" doi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21" start="Figure 15" startId="F15">
<paragraph id="E68E498452500E326A3372F5F3830C37" pageId="0" pageNumber="21">
<emphasis id="3E73FD5BD4FB6329FF249108FC901B4E" bold="true" pageId="0" pageNumber="21">Figure 15.</emphasis>
<emphasis id="A3E92283DFCB0A7CD3B0AF39C1208E6C" bold="true" pageId="0" pageNumber="21">a, c</emphasis>
<taxonomicName id="1851BB5A423A959AEF742716BBE37B9D" authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu shiluensis" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="shiluensis">
<emphasis id="7A35FAD054E28FA4358F06803AD65A20" italics="true" pageId="0" pageNumber="21">Patu shiluensis</emphasis>
</taxonomicName>
Lin &amp; Li, 2009
<emphasis id="E8FBCA833F84052616BF958AFE9F9C59" bold="true" pageId="0" pageNumber="21">b, d</emphasis>
<taxonomicName id="BCD66D841ED82CF105E29AAF21364C8D" authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2021" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha seedam" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="seedam">
<emphasis id="3793BB5B36C37294BE9A48B9EB0C16C1" italics="true" pageId="0" pageNumber="21">Crassignatha seedam</emphasis>
</taxonomicName>
sp. nov. Epigynum, cleared:
<emphasis id="8DAA81EF7B15375F23AAB4DB9B945C55" bold="true" pageId="0" pageNumber="21">a, b</emphasis>
dorsal view
<emphasis id="7099429823E631309EA2DE60B1E4CA2C" bold="true" pageId="0" pageNumber="21">c, d</emphasis>
ventral view. Scale bars: 0.03 mm (
<emphasis id="0405FCA15D098CC77364E170D4B5A6B6" bold="true" pageId="0" pageNumber="21">a, c</emphasis>
); 0.05 mm (
<emphasis id="74F5739DA66246ACC752C7E9B2F37ED7" bold="true" pageId="0" pageNumber="21">b, d</emphasis>
).
</paragraph>
</caption>
<paragraph id="A65D7C029CE6438BBBEBB5660D7304AE" pageId="0" pageNumber="21">
<emphasis id="8DFAE898FBCEE47C126DF25FC342A464" bold="true" pageId="0" pageNumber="21">Female</emphasis>
: Total length 0.52, carapace 0.21 long, 0.2 wide; clypeus 0.04; Chelicera 0.07 long, 0.05 wide; Leg I: femur 0.15, patella 0.07, tibia 0.1, metatarsus 0.07, tarsus 0.1; leg formula I-II-IV-III; abdomen 0.34 long, 0.28 wide.
</paragraph>
</subSubSection>
<subSubSection id="75A9CA705996875059B075DAA2FBC1E6" pageId="0" pageNumber="21" type="notes">
<paragraph id="F2FD6B8DE9FEA34A9996670C54FE963F" pageId="0" pageNumber="21">Notes.</paragraph>
<paragraph id="240380B4ADA10122C098286485BA155B" pageId="0" pageNumber="21">
Small somatic variations can be seen between the specimen we collected in Thailand and the ones previously described from China (compare Fig.
<figureCitation id="E57F6B113EF1E33E51625DE97276BF66" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14b</figureCitation>
to
<bibRefCitation id="440506AEDB95D16FEF8608ED1E3BC32A" author="Lin, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="47 - 68" refId="B23" refString="Lin, Y, Li, S, 2009. First described Patu spiders (Araneae, Symphytognathidae) from Asia. Zootaxa 2154: 47 - 68" title="First described Patu spiders (Araneae, Symphytognathidae) from Asia." volume="2154" year="2009">Lin and Li 2009</bibRefCitation>
: fig. 11). However, we did not find any objective differences in the female genitalia.
</paragraph>
<paragraph id="5EBE6ED9607D21B173A84372BFD36102" pageId="0" pageNumber="21">
Secretory ampullae (Figs
<figureCitation id="4FC25607AE3145AC20BD5DC86CFE104A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Patu shiluensis Lin &amp; Li, 2009 female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.2 mm (a, b); 0.06 mm (c); 0.03 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure14" httpUri="https://binary.pensoft.net/fig/501622" pageId="0" pageNumber="21">14d</figureCitation>
,
<figureCitation id="734E0ACA52265B016D312144A24B8375" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. a, c Patu shiluensis Lin &amp; Li, 2009 b, d Crassignatha seedam sp. nov. Epigynum, cleared: a, b dorsal view c, d ventral view. Scale bars: 0.03 mm (a, c); 0.05 mm (b, d)." figureDoi="10.3897/zookeys.1012.57047.figure15" httpUri="https://binary.pensoft.net/fig/501623" pageId="0" pageNumber="21">15a</figureCitation>
) were very evident in our specimens; these glandular structures might be homologous to the accessory glands in
<bibRefCitation id="944907E2612C7A999417C3100A41491C" DOI="https://doi.org/10.1111/zoj.12199" author="Lopardo, L" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="21" pagination="527 - 786" refId="B27" refString="Lopardo, L, Hormiga, G, 2015. Out of the twilight zone: Phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea). Zoological Journal of the Linnean Society 173 (3): 527 - 786, DOI: https://doi.org/10.1111/zoj.12199" title="Out of the twilight zone: Phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea)." url="https://doi.org/10.1111/zoj.12199" volume="173" year="2015">Lopardo and Hormiga (2015)</bibRefCitation>
. These structures were found in one anapid (
<taxonomicName id="4EC77696A7CB5C2E900B4646EABF5BEB" authorityName="Kobayashi" authorityYear="1940" class="Trilobita" family="Hystricuridae" genus="Tasmanaspis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tasmanaspis" order="Aulacopleurida" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="19CFF98F262BC988CBB5A27BD1C072BC" italics="true" pageId="0" pageNumber="21">Tasmanaspis</emphasis>
</taxonomicName>
) and several mysmenids, but scored as absent or unknown for all the symphytognathids.
</paragraph>
<paragraph id="882DFC65A610B426F68DEB360E5D4849" pageId="0" pageNumber="21">
The authors of this species mentioned it to be close to
<taxonomicName id="7C809843F1B738300AB973227DD56451" authorityName="Saaristo" authorityYear="1996" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu silho" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="silho">
<emphasis id="6705B84084B9335F5180C78E4F973C0E" italics="true" pageId="0" pageNumber="21">Patu silho</emphasis>
</taxonomicName>
Saaristo, 1996 from Seychelles. The possibility of
<taxonomicName id="69D873EE241A51B84E58670311D8F551" lsidName="P. silho" pageId="0" pageNumber="21" rank="species" species="silho">
<emphasis id="E4651376BECB2E71CDF884A99ED80A7C" italics="true" pageId="0" pageNumber="21">P. silho</emphasis>
</taxonomicName>
not being a true
<taxonomicName id="016D4E1F37C2AA54C7B0B7C6118B4DA9" authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="8A275C2168866F62D7D77BF0DCDCA150" italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
was discussed by its author (
<bibRefCitation id="B4BA70851E07D92BD8A77CDA4514C11D" author="Saaristo, MI" journalOrPublisher="Phelsuma" pageId="0" pageNumber="21" pagination="53 - 56" refId="B43" refString="Saaristo, MI, 1996. Symphytognathidae (Arachnida, Araneae), a new spider family for the granitic islands of Seychelles. Phelsuma 4: 53 - 56" title="Symphytognathidae (Arachnida, Araneae), a new spider family for the granitic islands of Seychelles." volume="4" year="1996">Saaristo 1996</bibRefCitation>
;
<bibRefCitation id="AEA8190CCD10C806DD379244D2FCEB7E" author="Saaristo, MI" journalOrPublisher="Press Manchester, UK" pageId="0" pageNumber="21" refId="B44" refString="Saaristo, MI, 2010. Araneae. In: Gerlach J, Marusik Y (Eds) Arachnida and Myriapoda of the Seychelles islands. Press Manchester, UK" title="Araneae. In: Gerlach J, Marusik Y (Eds) Arachnida and Myriapoda of the Seychelles islands." year="2010">2010</bibRefCitation>
) mentioning evident differences on somatic and sexual characters between
<taxonomicName id="EB1A02038F28B7B8D11EE868438CECD4" lsidName="P. silho" pageId="0" pageNumber="21" rank="species" species="silho">
<emphasis id="50CDC0C152ED7B17759C814FF6456B1E" italics="true" pageId="0" pageNumber="21">P. silho</emphasis>
</taxonomicName>
and other
<taxonomicName id="67DAE9241E2B809EA6F173632BA6021D" authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="76B145CEC3150B9D40DEA959D45D0FA2" italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
species. Nevertheless, the author deemed appropriate to place it in this genus. We also consider this species might be misplaced in
<taxonomicName id="36330303A3B9320B48FE744D6492FF72" authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="DF12CFBD429BA726546A03A42D02E643" italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
but would need further and more detailed analysis out of the scope of this work to clarify it (see discussion on
<taxonomicName id="D84D6B5FC4E9D700D3C72B352B23D1FA" authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="75C0AFC9D34922109F7550D78C3B2E40" italics="true" pageId="0" pageNumber="21">Patu</emphasis>
</taxonomicName>
relationships below).
</paragraph>
</subSubSection>
</treatment>
</document>