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<document id="22BD243382A3DBFC66B137094066E743" ID-CLB-Dataset="32713" ID-DOI="10.11646/zootaxa.4287.1.1" ID-GBIF-Dataset="91e8f5fd-1e15-4c4f-872f-3d4c50d0c8a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="829051" ID-ZooBank="13C4F702-EF00-4F04-B38E-3F0AA6CAF718" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110702684" checkinUser="plazi" docAuthor="ROBERTO CALDARA, CHARLES W. O’BRIEN &amp; MASSIMO MEREGALLI" docDate="2017" docId="D17787854F5D1F2B0DD76BAE13C3FEBC" docLanguage="en" docName="zootaxa.4287.1.1.pdf.imf" docOrigin="Zootaxa 4287 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Memptorrhynchus Iablokoff-Khnzorian 1960" docType="treatment" docVersion="8" lastPageNumber="34" masterDocId="2D4EFFFD4F7D1F0A0D406F35171FFFD9" masterDocTitle="A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults" masterLastPageNumber="63" masterPageNumber="1" pageNumber="33" updateTime="1698465391549" updateUser="ExternalLinkService">
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<mods:title id="817B55DD95EC37BF813C65501B19BA1F">A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults</mods:title>
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<mods:namePart id="305DAD094724971E9A3CAA777E402D56">ROBERTO CALDARA</mods:namePart>
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<mods:namePart id="039B117F6DC739631D99BFF3E4FE7101">CHARLES W. O’BRIEN</mods:namePart>
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<mods:namePart id="AE754C3623F4ABD00AFB31DE83376FB0">MASSIMO MEREGALLI</mods:namePart>
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<treatment id="D17787854F5D1F2B0DD76BAE13C3FEBC" ID-DOI="http://doi.org/10.5281/zenodo.4386628" ID-GBIF-Taxon="132549331" ID-Zenodo-Dep="4386628" LSID="urn:lsid:plazi:treatment:D17787854F5D1F2B0DD76BAE13C3FEBC" httpUri="http://treatment.plazi.org/id/D17787854F5D1F2B0DD76BAE13C3FEBC" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="11C465184F5D1F2A0DD76BAE1319FB6C" box="[151,1030,1178,1205]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="596136934F5D1F2A0DD76BAE1319FB6C" blockId="32.[151,1030,1178,1205]" box="[151,1030,1178,1205]" pageId="32" pageNumber="33">
<heading id="022981FF4F5D1F2A0DD76BAE1319FB6C" bold="true" box="[151,1030,1178,1205]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="6BAAEA814F5D1F2A0DD76BAE1319FB6C" bold="true" box="[151,1030,1178,1205]" pageId="32" pageNumber="33">
Genus
<taxonomicName id="9EDE4D104F5D1F2A0DAE6BAF143BFB6C" ID-CoL="8HM6X" authority="Iablokoff-Khnzorian, 1960" authorityName="Iablokoff-Khnzorian" authorityYear="1960" box="[238,804,1178,1205]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5D1F2A0DAE6BAF16D0FB6D" bold="true" box="[238,463,1178,1204]" italics="true" pageId="32" pageNumber="33">Memptorrhynchus</emphasis>
Iablokoff-Khnzorian, 1960
</taxonomicName>
resurrected name
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="11C465184F5D1F2A0DD76BD41289FACF" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="596136934F5D1F2A0DD76BD41289FACF" blockId="32.[151,1435,1249,1302]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="79CE11BD4F5D1F2A0DD76BD41289FACF" pageId="32" pageNumber="33">
<taxonomicName id="9EDE4D104F5D1F2A0DD76BD415B9FB2E" ID-CoL="8HM6X" authority="Iablokoff-Khnzorian, 1960: 253" authorityName="Iablokoff-Khnzorian" authorityPageNumber="253" authorityYear="1960" box="[151,678,1249,1271]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5D1F2A0DD76BD4164FFB2E" box="[151,336,1249,1271]" italics="true" pageId="32" pageNumber="33">Memptorrhynchus</emphasis>
Iablokoff-Khnzorian, 1960: 253
</taxonomicName>
[type species:
<taxonomicName id="9EDE4D104F5D1F2A0E096BD41264FB2E" ID-CoL="3ZPBZ" authority="Iablokoff-Khnzorian, 1960" authorityName="Iablokoff-Khnzorian" authorityYear="1960" box="[841,1403,1249,1271]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ripicola">
<emphasis id="6BAAEA814F5D1F2A0E096BD41343FB2E" box="[841,1116,1249,1271]" italics="true" pageId="32" pageNumber="33">Memptorrhynchus ripicola</emphasis>
Iablokoff-Khnzorian, 1960
</taxonomicName>
(=
<taxonomicName id="9EDE4D104F5D1F2A0D876A341502FACF" ID-CoL="KDN8" authority="Kirsch, 1878" authorityName="Kirsch" authorityYear="1878" box="[199,541,1280,1302]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="6BAAEA814F5D1F2A0D876A341691FACF" box="[199,398,1280,1302]" italics="true" pageId="32" pageNumber="33">Bagous brevipennis</emphasis>
Kirsch, 1878
</taxonomicName>
)].
<treatmentCitation id="D87F10824F5D1F2A0F756A35147DFACF" author="Caldara" box="[565,866,1280,1302]" page="137" pageId="32" pageNumber="33" year="1998">
<bibRefCitation id="3D4F4B624F5D1F2A0F756A351430FACF" author="Caldara" box="[565,815,1280,1302]" pageId="32" pageNumber="33" refString="Caldara, R. &amp; O'Brien, C. W. (1998) Systematics and evolution of weevils of the genus Bagous. VI. Taxonomic treatment of the species of the western Palaearctic Region (Coleoptera Curculionidae). Memorie della Societa Entomologica Italiana, 76 [1997], 131 - 347." type="journal article" year="1998">Caldara &amp; O'Brien 1998</bibRefCitation>
: 137
</treatmentCitation>
.
<treatmentCitation id="D87F10824F5D1F2A0E2C6A3513D5FACF" author="Alonso-Zarazaga" box="[876,1226,1280,1302]" page="88" pageId="32" pageNumber="33" year="1999">
<bibRefCitation id="3D4F4B624F5D1F2A0E2C6A3513BBFACF" author="Alonso-Zarazaga" box="[876,1188,1280,1302]" pageId="32" pageNumber="33" refString="Alonso-Zarazaga, M. A. &amp; Lyal, C. H. C. (1999) A World Catalogue of Families and Genera of Curculionoidea (Insecta: Coleoptera) (Excepting Scolytidae and Platypodidae). Entomopraxis S. C. P. Edition, Barcelona, 315 pp." type="book" year="1999">Alonso-Zarazaga &amp; Lyal 1999</bibRefCitation>
: 88
</treatmentCitation>
.
<treatmentCitation id="D87F10824F5D1F2A09946A35128DFACF" author="Caldara" box="[1236,1426,1280,1302]" page="172" pageId="32" pageNumber="33" year="2013">
<bibRefCitation id="3D4F4B624F5D1F2A09946A351240FACF" author="Caldara" box="[1236,1375,1280,1302]" pageId="32" pageNumber="33" refString="Caldara, R. (2013) Curculionidae: Bagoinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 8. Leiden, Brill, pp. 56 - 58, 172 - 176" type="journal article" year="2013">Caldara 2013</bibRefCitation>
: 172
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="11C465184F5D1F2A0DD76A7716BDF93E" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="596136934F5D1F2A0DD76A7716BDF93E" blockId="32.[151,1437,1346,2019]" pageId="32" pageNumber="33">
<emphasis id="6BAAEA814F5D1F2A0DD76A77160EFA82" bold="true" box="[151,273,1346,1371]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
Body broad-oval and robust. Rostrum unusually flattened at apex (char. 8.1), with lateral plica (char. 9.1); curvature of ventral margin in lateral view strong (char. 7.1), at basal 2/3 with row of sparse, regularly spaced setae along lateral margin close to ventral margin (char. 13.1). Forehead between eyes not sulcate nor foveate (char. 24.0). Pronotal disc moderately to strongly transversely convex. Postocular lobe ventrally indistinct, not raised, not forming discontinuity with anterior prosternal margin (char. 30.0). Prosternal canal weak (char. 31.0). Side margins of prosternum not clearly raised (char. 33.0). Elytra with setae on odd-numbered intervals surrounded by rosette of scales of same color; declivital callus absent (char. 36.0). Outer margin of tibiae, especially on hind tibiae, straight or nearly so toward apex (in lateral view) (char. 48.0); tibiae denticulate along inner margin, sometimes with denticles stouter and more numerous on midtibiae; tarsi short, tarsomere 3 cordate. Penis with long apodemes (char. 110.0). Endophallus with basal sclerite complex formed by elongate curved sclerites (char. 90.2). Apodemes of spiculum ventrale fully bifurcated, arising laterally on plate of sternite, apparently forming two straight rods with arms (char. 114.2).
</paragraph>
</subSubSection>
<subSubSection id="11C465184F5D1F2B0D8769C713C3FEBC" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="discussion">
<paragraph id="596136934F5D1F2A0D8769C713B1F8D2" blockId="32.[151,1437,1346,2019]" box="[199,1198,1778,1803]" pageId="32" pageNumber="33">
<emphasis id="6BAAEA814F5D1F2A0D8769C71624F8D2" bold="true" box="[199,315,1778,1803]" pageId="32" pageNumber="33">Remarks.</emphasis>
Monophyly of the species referred to this genus was supported in all analyses.
</paragraph>
<paragraph id="596136934F5D1F2B0D876823135CFF0C" blockId="32.[151,1437,1346,2019]" lastBlockId="33.[151,1436,151,357]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<taxonomicName id="9EDE4D104F5D1F2A0D876823168BF8F6" box="[199,404,1814,1839]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5D1F2A0D876823168BF8F6" box="[199,404,1814,1839]" italics="true" pageId="32" pageNumber="33">Memptorrhynchus</emphasis>
</taxonomicName>
is characterized by the poorly developed, barely distinguishable prosternal canal, the tibiae almost straight to apex along the outer margin, the tarsi short with tarsomere 3 broadly cordate, and the male genitalia which present certain similarities with those of
<taxonomicName id="9EDE4D104F5D1F2A0E5C686B14B3F8AE" box="[796,940,1886,1911]" class="Insecta" family="Curculionidae" genus="Hydronomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="6BAAEA814F5D1F2A0E5C686B14B3F8AE" box="[796,940,1886,1911]" italics="true" pageId="32" pageNumber="33">Hydronomus</emphasis>
</taxonomicName>
(
<emphasis id="6BAAEA814F5D1F2A0EFE686A14FFF8AF" box="[958,992,1887,1910]" italics="true" pageId="32" pageNumber="33">i.e.</emphasis>
, the features of the sclerite complex at the base of the endophallus). This is a homogeneous group and its species show six synapomorphies (char. 7.1, 8.1, 9.1, 13.1, 90.2, 114.2) in nongenital characters (rostrum strongly curved and with flat apex with lateral carinae, and with row of lateral setae) as well as genital morphology (shape of the penis and especially of the spiculum ventrale with apodemes strongly divergent and arising laterally on plate). The bulk of this group is present in the Afrotropical region with three extralimital species (two Indian and one Palaearctic).
</paragraph>
<paragraph id="596136934F5C1F2B0D876FEA13C3FEBC" blockId="33.[151,1436,151,357]" pageId="33" pageNumber="34">
Species included:
<emphasis id="6BAAEA814F5C1F2B0CDB6FEA16C1FF21" bold="true" box="[411,478,223,248]" pageId="33" pageNumber="34">AFR:</emphasis>
¹
<taxonomicName id="9EDE4D104F5C1F2B0CB26FEA131FFF21" authority="Marshall, 1953" authorityName="Marshall" authorityYear="1953" box="[498,1024,223,248]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="bergensis" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B0CB26FEA142BFF21" box="[498,820,223,248]" italics="true" pageId="33" pageNumber="34">Memptorrhynchus bergensis</emphasis>
(Marshall, 1953)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B094B6FEA136AFF21" box="[1035,1141,223,248]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B094B6FEA136AFF21" bold="true" box="[1035,1141,223,248]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
, ¹
<taxonomicName id="9EDE4D104F5C1F2B09D06FD417C7FEC4" authority="Hoffmann, 1968" authorityName="Hoffmann" authorityYear="1968" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="crassus" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B09D06FD41211FF21" box="[1168,1294,225,248]" italics="true" pageId="33" pageNumber="34">M. crassus</emphasis>
(Hoffmann, 1968)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B0DA16E311657FEC4" box="[225,328,260,285]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B0DA16E311657FEC4" bold="true" box="[225,328,260,285]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
, *¹
<taxonomicName id="9EDE4D104F5C1F2B0C2E6E30159AFEC5" authority="Faust, 1899" authorityName="Faust" authorityYear="1899" box="[366,645,260,284]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="crispus" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B0C2E6E3016FAFEC5" box="[366,485,261,284]" italics="true" pageId="33" pageNumber="34">M. crispus</emphasis>
(Faust, 1899)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B0FCD6E3115EAFEC4" box="[653,757,260,285]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B0FCD6E3115EAFEC4" bold="true" box="[653,757,260,285]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
, ¹
<taxonomicName id="9EDE4D104F5C1F2B0E4D6E30137FFEC4" authority="Hustache, 1936" authorityName="Hustache" authorityYear="1936" box="[781,1120,260,285]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="gaillardi" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B0E4D6E30148AFEC4" box="[781,917,260,285]" italics="true" pageId="33" pageNumber="34">M. gaillardi</emphasis>
(Hustache, 1936)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B09286E3113D0FEC4" box="[1128,1231,260,285]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B09286E3113D0FEC4" bold="true" box="[1128,1231,260,285]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
, ¹
<taxonomicName id="9EDE4D104F5C1F2B09A76E301678FE99" authority="Hoffmann, 1968" authorityName="Hoffmann" authorityYear="1968" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="polysignatus" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B09A76E301284FEC4" box="[1255,1435,260,285]" italics="true" pageId="33" pageNumber="34">M. polysignatus</emphasis>
(Hoffmann, 1968)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B0C336E1216C2FE99" box="[371,477,295,320]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B0C336E1216C2FE99" bold="true" box="[371,477,295,320]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
, ¹
<taxonomicName id="9EDE4D104F5C1F2B0CB86E1C1436FE99" authority="Marshall, 1958" authorityName="Marshall" authorityYear="1958" box="[504,809,295,320]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vafer" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B0CB86E1C1543FE99" box="[504,604,296,320]" italics="true" pageId="33" pageNumber="34">M. vafer</emphasis>
(Marshall, 1958)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B0E746E121480FE99" box="[820,927,295,320]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B0E746E121480FE99" bold="true" box="[820,927,295,320]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
;
<emphasis id="6BAAEA814F5C1F2B0EF36E1214E5FE99" bold="true" box="[947,1018,295,320]" pageId="33" pageNumber="34">ORR:</emphasis>
¹
<taxonomicName id="9EDE4D104F5C1F2B094E6E1C1265FE99" authority="Hustache, 1926" authorityName="Hustache" authorityYear="1926" box="[1038,1402,295,320]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="luteitarsis" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B094E6E1C13B7FE99" box="[1038,1192,295,320]" italics="true" pageId="33" pageNumber="34">M. luteitarsis</emphasis>
(Hustache, 1926)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B08C56E1217C0FEBC" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B08C56E1217C0FEBC" bold="true" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
, ¹
<taxonomicName id="9EDE4D104F5C1F2B0DB56E781559FEBC" authority="Hustache, 1926" authorityName="Hustache" authorityYear="1926" box="[245,582,332,357]" class="Insecta" family="Curculionidae" genus="Memptorrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="ovoideus" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B0DB56E781660FEBC" box="[245,383,332,357]" italics="true" pageId="33" pageNumber="34">M. ovoideus</emphasis>
(Hustache, 1926)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B0F0D6E7915ACFEBC" box="[589,691,332,357]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B0F0D6E7915ACFEBC" bold="true" box="[589,691,332,357]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
;
<emphasis id="6BAAEA814F5C1F2B0F826E7915E0FEBC" bold="true" box="[706,767,332,357]" pageId="33" pageNumber="34">PAL:</emphasis>
*²
<taxonomicName id="9EDE4D104F5C1F2B0E5E6E78136FFEBD" authority="Kirsch, 1878" authorityName="Kirsch" authorityYear="1878" box="[798,1136,332,357]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="brevipennis" status="comb. nov.">
<emphasis id="6BAAEA814F5C1F2B0E5E6E7814D9FEBC" box="[798,966,332,357]" italics="true" pageId="33" pageNumber="34">M. brevipennis</emphasis>
(Kirsch, 1878)
</taxonomicName>
<taxonomicNameLabel id="709957FA4F5C1F2B09376E7913C3FEBC" box="[1143,1244,332,357]" pageId="33" pageNumber="34">
<emphasis id="6BAAEA814F5C1F2B09376E7913C3FEBC" bold="true" box="[1143,1244,332,357]" pageId="33" pageNumber="34">n. comb.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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