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<mods:title id="35E64BF4B49CFF55D1D818DB6064BA27">Revision of the Genus Paracloeodes (Ephemeroptera: Baetidae) in South America</mods:title>
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<paragraph id="5943A451966D7B0DFE90AFD7FE438E4D" blockId="4.[264,483,758,784]" box="[264,483,758,784]" pageId="4" pageNumber="5">
<heading id="020B133D966D7B0DFE90AFD7FE438E4D" bold="true" box="[264,483,758,784]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis id="6B887843966D7B0DFE90AFD7FE438E4D" bold="true" box="[264,483,758,784]" pageId="4" pageNumber="5">
<emphasis id="6B887843966D7B0DFE90AFD7FE098E4D" bold="true" box="[264,425,758,784]" italics="true" pageId="4" pageNumber="5">Paracloeodes</emphasis>
Day
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<taxonomicName id="9EFCDFD2966D7B0DFE90AE63FDE78E05" ID-CoL="63J5X" authority="Day, 1955: 121" authorityName="Day" authorityPageNumber="121" authorityYear="1955" box="[264,583,833,857]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966D7B0DFE90AE63FE3B8E04" box="[264,411,834,857]" italics="true" pageId="4" pageNumber="5">Paracloeodes</emphasis>
<treatmentCitation id="D85D8240966D7B0DFE39AE60FDE78E05" author="Day" box="[417,583,833,856]" page="121" pageId="4" pageNumber="5" year="1955">
<bibRefCitation id="3D6DD9A0966D7B0DFE39AE60FDB28E05" author="Day" box="[417,530,833,856]" pageId="4" pageNumber="16" refString="Day, W. C. (1955) New genera of mayflies from California. Pan-Pacific Entomologist. 31, 121 - 137." type="journal article" year="1955">Day, 1955</bibRefCitation>
: 121
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;
<treatmentCitation id="D85D8240966D7B0DFDCAAE60FC428E05" author="Lugo-Ortiz" box="[594,994,833,856]" page="162" pageId="4" pageNumber="5" year="1996">
<bibRefCitation id="3D6DD9A0966D7B0DFDCAAE60FC0D8E05" author="Lugo-Ortiz" box="[594,941,833,856]" pageId="4" pageNumber="16" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1996) The genus Paracloeodes (Insecta: Ephemeroptera: Baetidae) and its presence in South America. Annales Limnologie. 32 (3), 161 - 169." type="journal article" year="1996">Lugo­Ortiz &amp; McCafferty, 1996</bibRefCitation>
: 162
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.
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</paragraph>
<paragraph id="5943A451966D7B0DFE90AE43FC9F8E25" blockId="4.[264,1311,833,921]" box="[264,831,865,889]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="79EC837F966D7B0DFE90AE43FC9F8E25" box="[264,831,865,889]" pageId="4" pageNumber="5">
<taxonomicName id="9EFCDFD2966D7B0DFE90AE43FD148E25" authority="Salles &amp; Lugo-Ortiz, 2003: 202" authorityName="Salles &amp; Lugo-Ortiz" authorityPageNumber="202" authorityYear="2003" box="[264,692,865,889]" class="Insecta" family="Baetidae" genus="Iguaira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966D7B0DFE90AE43FEF88E24" box="[264,344,866,889]" italics="true" pageId="4" pageNumber="5">Iguaira</emphasis>
<treatmentCitation id="D85D8240966D7B0DFEC7AE40FD148E25" author="Salles" box="[351,692,865,888]" page="202" pageId="4" pageNumber="5" year="2003">
<bibRefCitation id="3D6DD9A0966D7B0DFEC7AE40FDDF8E25" author="Salles" box="[351,639,865,888]" pageId="4" pageNumber="16" refString="Salles, F. F. &amp; Lugo-Ortiz, C. R. (2003) Um novo genero e especie de Baetidae (Ephemeroptera) do Estado de Minas Gerais, Sudeste do Brasil. Iheringia, Serie Zoologia, Porto Alegre. 93 (2), 201 - 206." type="journal article" year="2003">Salles &amp; Lugo­Ortiz, 2003</bibRefCitation>
: 202
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. SYN. NOV.
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<typeStatus id="86471AF3966D7B0DFE90AEA0FE9D8EC5" box="[264,317,897,920]" pageId="4" pageNumber="5">Type</typeStatus>
species:
<taxonomicName id="9EFCDFD2966D7B0DFE07AEA3FD558EC5" authority="Day 1955" authorityName="Day" authorityYear="1955" box="[415,757,897,921]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="abditus">
<emphasis id="6B887843966D7B0DFE07AEA3FD268EC4" box="[415,646,898,921]" italics="true" pageId="4" pageNumber="5">Paracloeodes abditus</emphasis>
<bibRefCitation id="3D6DD9A0966D7B0DFD15AEA0FD558EC5" author="Day" box="[653,757,897,920]" pageId="4" pageNumber="16" refString="Day, W. C. (1955) New genera of mayflies from California. Pan-Pacific Entomologist. 31, 121 - 137." type="journal article" year="1955">Day 1955</bibRefCitation>
</taxonomicName>
[synonym of
<taxonomicName id="9EFCDFD2966D7B0DFC12AEA3FAB98EC5" authority="(Daggy) 1945" authorityName="Daggy" baseAuthorityName="Daggy" baseAuthorityYear="1945" box="[906,1305,897,921]" class="Insecta" family="Baetidae" genus="Pseudocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="6B887843966D7B0DFC12AEA3FBDA8EC4" box="[906,1146,898,921]" italics="true" pageId="4" pageNumber="5">Pseudocloeon minutus</emphasis>
(Daggy) 1945
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]
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<subSubSection id="11E6F7DA966D7B0DFE90AEE0FC298B8E" pageId="4" pageNumber="5" type="description">
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<emphasis id="6B887843966D7B0DFE90AEE0FE088E86" bold="true" box="[264,424,961,987]" pageId="4" pageNumber="5">Male Imago.</emphasis>
Turbinate eyes oval, height of stalks half of eye diameter. Forewings with paired marginal intercalaries.
<collectingCountry id="21EBE4C1966D7B0DFDF0AEC8FD03895E" box="[616,675,1001,1027]" name="India" pageId="4" pageNumber="5">Hind</collectingCountry>
wings present or absent, if present with 2 longitudinal veins and with quadrangular costal process on basal third of anterior margin. Genitalia with forceps three­segmented, segment III elongate, bases of forceps widely separated.
</paragraph>
<paragraph id="5943A451966D7B0DFEA0A940FC298B8E" blockId="4.[264,1325,961,1867]" pageId="4" pageNumber="5">
<emphasis id="6B887843966D7B0DFEA0A940FE3C8926" bold="true" box="[312,412,1121,1147]" pageId="4" pageNumber="5">Nymph.</emphasis>
Head as long as wide. Frontal keel present or absent. Antennae 1.54 times the head capsule. Mouthparts: labrum (
<figureCitation id="C1C7B8D4966D7B0DFD46A9A8FC8289FE" box="[734,802,1161,1187]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
a) wider than long, cleft of anterior margin with a small central lobe, dorsal surface with bifid setae on anterior margin (
<figureCitation id="C1C7B8D4966D7B0DFB2BA990FAAD8996" box="[1203,1293,1201,1227]" captionStart="FIGURE 28 39" captionStartId="26.[264,368,1734,1758]" captionTargetBox="[300,1255,324,1733]" captionTargetId="figure@26.[293,1294,308,1738]" captionTargetPageId="26" captionText="FIGURE 28 39. Paracloeodes eurybranchus. Nymph. Mouthparts (Figs. 28 33): 28 a, labrum, left d. v., right v. v; 28 b, labrum: apical setae; 29, left mandible v. v.; 30, right mandible v. v.; 31, hypopharynx v. v.; 32, maxilla v. v.; 33, labium, left d. v., right v. v. 34, leg I; 35, tarsal claw I. 36, posterior margin of tergum IV. 37, gill IV. 38, paraproct. 39, caudal filament detail: left cercus, right terminal filament." httpUri="https://zenodo.org/record/173672/files/figure.png" pageId="4" pageNumber="5">Fig. 28</figureCitation>
b). Mandibles (
<figureCitation id="C1C7B8D4966D7B0DFE0DA9F8FE4889AE" box="[405,488,1241,1267]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Figs. 2</figureCitation>
a, 3) without a row of setae on anterior margin between prostheca and mola, incisors cleft in two sets, inner set at right angle with outer set. Left mandible with molars with constrictions (
<figureCitation id="C1C7B8D4966D7B0DFDD1A808FD33881E" box="[585,659,1321,1347]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
b), prostheca with 23 denticles (
<figureCitation id="C1C7B8D4966D7B0DFBBEA808FBCF881E" box="[1062,1135,1321,1347]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
a). Prostheca of right mandible slender and bifid (
<figureCitation id="C1C7B8D4966D7B0DFD09A870FD788836" box="[657,728,1361,1387]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
). Hypopharynx as in
<figureCitation id="C1C7B8D4966D7B0DFC4EA870FBBE8836" box="[982,1054,1361,1387]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
. Maxillae (
<figureCitation id="C1C7B8D4966D7B0DFB39A870FB488836" box="[1185,1256,1361,1387]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
) with palpi two­segmented, longer than apex of galea­lacinia. Labium (
<figureCitation id="C1C7B8D4966D7B0DFBCDA858FB0588CE" box="[1109,1189,1401,1427]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
a): glossae subequal to paraglossae, rounded apically and with short spines on external margins, paraglossae with a row of long spine­like setae; segment II of palpi with a distomedial projection, segment III conical, both segments with long setae (
<figureCitation id="C1C7B8D4966D7B0DFC61A8D0FBED8B56" box="[1017,1101,1521,1547]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Figs. 6</figureCitation>
a, b). Legs (
<figureCitation id="C1C7B8D4966D7B0DFB43A8D0FA828B56" box="[1243,1314,1521,1547]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
) with femora with a dorsal row of short spines, tibiae subequal to tarsi. Tarsal claws elongate, with two rows of denticles (
<figureCitation id="C1C7B8D4966D7B0DFD5EAB60FCAE8B06" box="[710,782,1601,1627]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
).
<collectingCountry id="21EBE4C1966D7B0DFCBEAB60FCC18B06" box="[806,865,1601,1627]" name="India" pageId="4" pageNumber="5">Hind</collectingCountry>
wing pads present or absent. Posterior margins of abdominal terga with spines (
<figureCitation id="C1C7B8D4966D7B0DFC9BAB48FCEF8BDE" box="[771,847,1641,1667]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
). Gills (
<figureCitation id="C1C7B8D4966D7B0DFC20AB48FBB18BDE" box="[952,1041,1641,1667]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
) present on abdominal segments IVII, elongate, more than two times length of tergum. Paraprocts with spines apically (
<figureCitation id="C1C7B8D4966D7B0DFEEDAB98FE6B8B8E" box="[373,459,1721,1747]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
). Terminal filament subequal to cerci.
</paragraph>
</subSubSection>
<subSubSection id="11E6F7DA966D7B0CFEA0ABC0FC1B8E4B" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="5943A451966D7B0CFEA0ABC0FE4D8C8B" blockId="4.[264,1325,961,1867]" lastBlockId="5.[264,1324,284,1110]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="6B887843966D7B0DFEA0ABC0FE638BA6" bold="true" box="[312,451,1761,1787]" pageId="4" pageNumber="5">Discussion.</emphasis>
The description of new species and the revision of known species showed new evidence for the genus
<taxonomicName id="9EFCDFD2966D7B0DFDC0AA28FD598A7F" box="[600,761,1801,1826]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966D7B0DFDC0AA28FD598A7F" box="[600,761,1801,1826]" italics="true" pageId="4" pageNumber="5">Paracloeodes</emphasis>
</taxonomicName>
.
<bibRefCitation id="3D6DD9A0966D7B0DFC91AA28FB348A7E" author="Lugo-Ortiz" box="[777,1172,1801,1827]" pageId="4" pageNumber="16" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1996) The genus Paracloeodes (Insecta: Ephemeroptera: Baetidae) and its presence in South America. Annales Limnologie. 32 (3), 161 - 169." type="journal article" year="1996">Lugo­Ortiz &amp; McCafferty (1996)</bibRefCitation>
proposed as diagnostic characteristics the small body size of the nymphs, distinct shape of labial palpi, elongate tarsal claws with poorly defined denticulation, numerous minute ridges on the abdominal terga, relatively elongate gills and adults with small body size and a characteristic hind wing, when present. The cladistic analysis presented here isolated seven synapomorphies which change upon the diagnosis presented by
<bibRefCitation id="3D6DD9A0966C7B0CFB1AACB5FE498C8B" author="Lugo-Ortiz" pageId="5" pageNumber="16" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1996) The genus Paracloeodes (Insecta: Ephemeroptera: Baetidae) and its presence in South America. Annales Limnologie. 32 (3), 161 - 169." type="journal article" year="1996">Lugo­Ortiz &amp; McCafferty (1996)</bibRefCitation>
.
</paragraph>
<paragraph id="5943A451966C7B0CFEA0ACC5FC1B8E4B" blockId="5.[264,1324,284,1110]" pageId="5" pageNumber="6">
<taxonomicName id="9EFCDFD2966C7B0CFEA0ACC5FE308CA0" box="[312,400,484,509]" class="Insecta" family="Baetidae" genus="Iguaira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966C7B0CFEA0ACC5FE308CA0" box="[312,400,484,509]" italics="true" pageId="5" pageNumber="6">Iguaira</emphasis>
</taxonomicName>
was established by presenting a single synapomorphy: the segment I of labial palpi transverse to segments II and III. Only one specimen was studied at this moment. New material collected showed segment I of palpi orientated in the same way as the others two segments, subparallel to glossae and paraglossae. Our examination of material has shown that all characteristics present in
<taxonomicName id="9EFCDFD2966C7B0CFD7DAFA5FC098FC0" box="[741,937,644,669]" class="Insecta" family="Baetidae" genus="Iguaira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="poranga">
<emphasis id="6B887843966C7B0CFD7DAFA5FC098FC0" box="[741,937,644,669]" italics="true" pageId="5" pageNumber="6">Iguaira poranga</emphasis>
</taxonomicName>
are shared with other species of
<taxonomicName id="9EFCDFD2966C7B0CFE90AF8DFE098F98" box="[264,425,684,709]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966C7B0CFE90AF8DFE098F98" box="[264,425,684,709]" italics="true" pageId="5" pageNumber="6">Paracloeodes</emphasis>
</taxonomicName>
. Results from the phylogenetic analysis clearly indicate that
<taxonomicName id="9EFCDFD2966C7B0CFB4BAF8DFECB8FB0" class="Insecta" family="Baetidae" genus="Iguaira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="poranga">
<emphasis id="6B887843966C7B0CFB4BAF8DFECB8FB0" italics="true" pageId="5" pageNumber="6">Iguaira poranga</emphasis>
</taxonomicName>
is a species of
<taxonomicName id="9EFCDFD2966C7B0CFD87AFF5FD608FB0" box="[543,704,724,749]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966C7B0CFD87AFF5FD608FB0" box="[543,704,724,749]" italics="true" pageId="5" pageNumber="6">Paracloeodes</emphasis>
</taxonomicName>
, so
<taxonomicName id="9EFCDFD2966C7B0CFD68AFF5FCE88FB0" box="[752,840,724,749]" class="Insecta" family="Baetidae" genus="Iguaira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966C7B0CFD68AFF5FCE88FB0" box="[752,840,724,749]" italics="true" pageId="5" pageNumber="6">Iguaira</emphasis>
</taxonomicName>
is placed as a junior subjective synonym of
<taxonomicName id="9EFCDFD2966C7B0CFEB0AFDDFE698E48" box="[296,457,764,789]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966C7B0CFEB0AFDDFE698E48" box="[296,457,764,789]" italics="true" pageId="5" pageNumber="6">Paracloeodes</emphasis>
</taxonomicName>
and
<taxonomicName id="9EFCDFD2966C7B0CFD9AAFDDFDDD8E48" box="[514,637,764,789]" class="Insecta" family="Baetidae" genus="Iguaira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="poranga">
<emphasis id="6B887843966C7B0CFD9AAFDDFDDD8E48" box="[514,637,764,789]" italics="true" pageId="5" pageNumber="6">I. poranga</emphasis>
</taxonomicName>
is transferred to the genus.
</paragraph>
</subSubSection>
<subSubSection id="11E6F7DA966C7B0CFEA0AE05FD50890B" pageId="5" pageNumber="6" type="description">
<paragraph id="5943A451966C7B0CFEA0AE05FD50890B" blockId="5.[264,1324,284,1110]" pageId="5" pageNumber="6">
<taxonomicName id="9EFCDFD2966C7B0CFEA0AE05FE798E60" box="[312,473,804,829]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="6B887843966C7B0CFEA0AE05FE798E60" box="[312,473,804,829]" italics="true" pageId="5" pageNumber="6">Paracloeodes</emphasis>
</taxonomicName>
can be distinguished from the other genera of the family by the following combinations of characters. In the male imago: 1) hind wings, if present, with 2 longitudinal veins and with a quadrangular costal process; 2) forceps three­segmented, segment III elongate. In the nymphs: 1) mandibles (
<figureCitation id="C1C7B8D4966C7B0CFCF3AEBDFC1E8EEB" box="[875,958,924,950]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
a, 3) with inner sets of incisors at right angle to outer set; 2) left mandible with molars with constrictions (
<figureCitation id="C1C7B8D4966C7B0CFB04AEE5FB468E83" box="[1180,1254,964,990]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
b); 3) prostheca of right mandible bifid; 4) labium (
<figureCitation id="C1C7B8D4966C7B0CFCDCAECDFC30895B" box="[836,912,1004,1030]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
a) with glossae rounded apically, segment II of palpi with a distomedial projection, segment III conical (
<figureCitation id="C1C7B8D4966C7B0CFBCFA935FB3F8973" box="[1111,1183,1044,1070]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
b); 5) tarsal claws with two rows of denticles (
<figureCitation id="C1C7B8D4966C7B0CFD01A91DFD40890B" box="[665,736,1084,1110]" captionStart="FIGURE 1 11" captionStartId="23.[264,368,1775,1799]" captionTargetBox="[321,1300,335,1722]" captionTargetId="figure@23.[250,1468,326,1770]" captionTargetPageId="23" captionText="FIGURE 1 11. Paracloeodes assu sp. n. Nymph. Mouthparts (Figs. 1 6): 1 a, labrum d. v.; 1 b, labrum: apical setae; 2 a, left mandible v. v.; 2 b, detail of teeth of molars; 3, right mandible v. v.; 4, hypopharynx v. v.; 5, maxilla v. v.; 6 a, labium, left d. v., right v. v.; 6 b, palp: segments II III. 7, leg I; 8, tarsal claw I. 9, posterior margin of tergum IV. 10, gill IV. 11, paraproct." httpUri="https://zenodo.org/record/173669/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>