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<document ID-DOI="http://dx.doi.org/10.3897/nl.47.114772" ID-Pensoft-Pub="2367-5365-47-57" ID-Pensoft-UUID="4A77F35768A45842A1B27D54D5B50724" ID-ZooBank="18BDAA9F5AE5479ABC3D1998C8193791" ModsDocID="2367-5365-47-57" checkinTime="1710546606512" checkinUser="pensoft" docAuthor="Shovkoon, Dmitry F. &amp; Trofimova, Tatyana A." docDate="2024" docId="D08316C4D7135606AAB3BFC4B9237282" docLanguage="en" docName="NotaLepidopt 47: 57-79" docOrigin="Nota Lepidopterologica 47" docPubDate="2024-03-15" docSource="http://dx.doi.org/10.3897/nl.47.114772" docTitle="Carriola zolotuhini Shovkoon &amp; Trofimova 2024, sp. nov." docType="treatment" docUuid="6673DB22-97EC-4FE2-ACAD-43BE54AD73B2" docUuidSource="ZooBank" docVersion="1" id="4A77F35768A45842A1B27D54D5B50724" lastPageNumber="57" masterDocId="4A77F35768A45842A1B27D54D5B50724" masterDocTitle="Review of the genus Carriola Swinhoe, 1922 (Lepidoptera, Erebidae, Lymantriinae), with descriptions of four new species" masterLastPageNumber="79" masterPageNumber="57" pageNumber="57" updateTime="1710546606512" updateUser="pensoft">
<mods:mods id="6A02F905109BBC7B2F45EC69F2F0704E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0994C87A0CC4E09A29697E739A2B81ED">
<mods:title id="B952313BA6FCFE6D2B20DBDC822D5BF2">Review of the genus Carriola Swinhoe, 1922 (Lepidoptera, Erebidae, Lymantriinae), with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name id="AA80F37682B9194729C21449BD3BFAE6" type="personal">
<mods:role id="5117110CDBD0D832A92796180AC386DC">
<mods:roleTerm id="F3D0F637A25AEBE1991BD05BCB0B7D9D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFC4AB821BD4F1FB8FB3BB961457146A">Shovkoon, Dmitry F.</mods:namePart>
<mods:nameIdentifier id="A64E06C82C648993D06391A3FA83A8D8" type="ORCID">https://orcid.org/0000-0001-7175-3489</mods:nameIdentifier>
<mods:affiliation id="7033004D098F2DF6F7D7E0755C93CD7D">Faculty of Science, University of South Bohemia, Branisovska 31, 370 05 Ceske Budejovice, Czech Republic; shovkoon. d @ googlemail. com &amp; Institute of Entomology, Biology Centre CAS, Branisovska 1760, 370 05 Ceske Budejovice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="9D28283D3DE5361B54EF2E94D5CF9C6E" type="email">shovkoon.d@googlemail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7B44B81293600856EF699B83C2447EC0" type="personal">
<mods:role id="9DF3E2B9B0782B28FD08417DA0DB8584">
<mods:roleTerm id="207ED56FF2345D400E53A12F7F64C60C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6C3052B053B0480D8C350B588729E854">Trofimova, Tatyana A.</mods:namePart>
<mods:nameIdentifier id="1EB9EFB501E5B17D381BCF7C33063BE1" type="ORCID">https://orcid.org/0000-0003-4998-3266</mods:nameIdentifier>
<mods:affiliation id="0955C45790A84ABA4050F214320D348E">Laboratory of Animal Systematic and Faunistic Samara State University, ul. Ac. Pavlova 1, Samara, 443011, Russia; apamea @ mail. ru</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E5CACC53A4335B0A22BF4FE17C2525EB">text</mods:typeOfResource>
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<mods:titleInfo id="9A306BF51F68267B7BCC341C26531FA2">
<mods:title id="2BD32E68000D91C95E21048A982C0DA0">Nota Lepidopterologica</mods:title>
</mods:titleInfo>
<mods:part id="A3CAE05DA99E562AC4B732FF295565FC">
<mods:date id="8D72B3E1A627B2EFFB87C9C26CCD22A2">2024</mods:date>
<mods:detail id="908C7FE591AB18463F816A29FC9CB332" type="pubDate">
<mods:number id="3E3631788B69F6BCE4BEEB90D002B42D">2024-03-15</mods:number>
</mods:detail>
<mods:detail id="543E4BC1B6A9D0AC849C49BEFFA5D79C" type="volume">
<mods:number id="81FD26F856A2BFF6801D0DB4797DBD1C">47</mods:number>
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<mods:extent id="2E389DEC3F30D3979B100F02F83A7C9A" unit="page">
<mods:start id="79A780E00A63C1FCB4D647106C671424">57</mods:start>
<mods:end id="BE2973CCC1266E29ABEA72FFE79BBE9D">79</mods:end>
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<mods:url id="10DAEA8D4B3D59C1A9EE7205B7878B34">http://dx.doi.org/10.3897/nl.47.114772</mods:url>
</mods:location>
<mods:classification id="839DD4D059D0280DFA9A1CD672AD43AD">journal article</mods:classification>
<mods:identifier id="AE00C22B010DF728868A5138802FF03F" type="DOI">http://dx.doi.org/10.3897/nl.47.114772</mods:identifier>
<mods:identifier id="291E652DEBDB923504E95B0E8D59BC85" type="Pensoft-Pub">2367-5365-47-57</mods:identifier>
<mods:identifier id="F006FC333ADAEE90CD0B46FC6804C926" type="ZooBank">18BDAA9F5AE5479ABC3D1998C8193791</mods:identifier>
<mods:identifier id="BACF195C5EF0F8A717C9AE3F0FF9C9E0" type="Pensoft-UUID">4A77F35768A45842A1B27D54D5B50724</mods:identifier>
</mods:mods>
<subSection id="EB15BED08D553724D0311A703C90E1FE" pageId="0" pageNumber="57" type="taxonomy">
<treatment id="D08316C4D7135606AAB3BFC4B9237282" LSID="urn:lsid:zoobank.org:act:6673DB22-97EC-4FE2-ACAD-43BE54AD73B2" httpUri="http://treatment.plazi.org/id/D08316C4D7135606AAB3BFC4B9237282" lastPageNumber="57" pageId="0" pageNumber="57">
<subSubSection id="B4E158CBAA7BA0C6C7689CFF0C5AE81A" pageId="0" pageNumber="57" type="nomenclature">
<paragraph id="013F2FFDD47E867141D2D70C012032A1" pageId="0" pageNumber="57">
<taxonomicName id="2A028646D57306FCA9245FF24E48B771" LSID="https://zoobank.org/6673DB22-97EC-4FE2-ACAD-43BE54AD73B2" authority="Shovkoon &amp; Trofimova, 2024" authorityName="Shovkoon &amp; Trofimova" authorityYear="2024" class="Insecta" family="Erebidae" genus="Carriola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carriola zolotuhini" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="zolotuhini" status="sp. nov.">Carriola zolotuhini</taxonomicName>
<taxonomicNameLabel id="AB477DDDEDC259C3E94186124DC52FA1" pageId="0" pageNumber="57">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="AB923B322AFBAFB3B33B1254E166970A" pageId="0" pageNumber="57" type="description">
<paragraph id="8D16D1AD5B10B4AF82F114563F98BBA7" pageId="0" pageNumber="57">
<figureCitation id="4751EF0818792B8F418F0B2893B84E8F" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">Figs 18</figureCitation>
<figureCitation id="D45B6C7734DDC891B10A09D6A2090185" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">, 27</figureCitation>
<figureCitation id="CF5B0F1AD3D4DD728C41A026A96102BA" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">, 36</figureCitation>
<figureCitation id="7F4D91FF9ED07E5CF05F0F4174800A05" captionStart="Figures 45–53" captionStartId="F7" captionText="Figures 45 - 53. Distribution maps of Carriola species. 45. C. ecnomoda (Swinhoe, 1907); 46. C. thyridophora (Hampson, [1893]); 47. C. witti sp. nov.; 48. C. seminsula (Strand, 1914); 49. C. saturnioides (Snellen, 1879); 50. C. polyakovi sp. nov.; 51. C. shorokhovi sp. nov.; 52. C. fenestrata (Hampson, [1893]); 53. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures45-53" httpUri="https://binary.pensoft.net/fig/1003326" pageId="0" pageNumber="57">, 53</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="847A4D733E024A0C9D78E58E262C3483" pageId="0" pageNumber="57" type="materials_examined">
<paragraph id="2E351CEAA585A0C0E94947F4D0584E5A" pageId="0" pageNumber="57">Material examined.</paragraph>
<paragraph id="C40E46A355C9B5EAB0A807314FE0C2F4" pageId="0" pageNumber="57">
<materialsCitation id="AF931D53069F2DCB7464E2F0AC9DC666" collectingDate="2008-03-30" collectingDateMax="2008-04-01" collectingDateMin="2008-03-30" collectorName="SE Mindanao, JH Lourens" country="Philippines" elevation="90" latitude="7.727783" location="Aliwagwag Primary forest" longLatPrecision="1" longitude="126.2884" municipality="Mindanao Island" specimenCount="1" specimenCount-male="1" stateProvince="Davao Oriental" typeStatus="Holotype">
<emphasis id="28429ED7D9CD67ADF1204643B3F76F00" bold="true" pageId="0" pageNumber="57">
<typeStatus id="82D8B8CA677FE6AB0F3D72AA7008A1C5" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="99EC600E9A4B3C67CF3135106DD3B876" name="Philippines">Philippines</collectingCountry>
:
<collectingMunicipality id="DA82C2C21CEA1664CA90237F3D775B94">Mindanao Island</collectingMunicipality>
•
<specimenCount id="313DE6178EAF5253EDA684EFFCFB1307" count="1" type="male">1 ♂</specimenCount>
;
<collectingCountry id="5518C264013BA844B6647D6AD1327E9D" name="Philippines">Philippines</collectingCountry>
,
<collectorName id="290D08342218CE53C0904A8436715EFB">SE Mindanao</collectorName>
,
<collectingRegion id="D743C09114B54578F15645FC890F2E83" country="Philippines" name="Davao Oriental">Davao Oriental</collectingRegion>
<location id="B46F5CE3D5ECC66A6883D0A9B71CC333" LSID="urn:lsid:plazi:treatment:D08316C4D7135606AAB3BFC4B9237282:B46F5CE3D5ECC66A6883D0A9B71CC333" country="Philippines" latitude="7.727783" longLatPrecision="1" longitude="126.2884" municipality="Mindanao Island" name="Aliwagwag Primary forest" stateProvince="Davao Oriental">Aliwagwag Primary forest</location>
,
<quantity id="6BB07E37377DB1A5B05254EF37D47542" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
<elevation id="964ECCB924365C35506411C9CE1CF21A" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
</quantity>
,
<geoCoordinate id="988CD4AC795138D1FCAA468DBD4AFA06" degrees="07" direction="north" minutes="43.667" orientation="latitude" precision="1" value="7.727783">07°43.667'N</geoCoordinate>
,
<geoCoordinate id="929A5E701A0D3591915111B5FFFA8DFB" degrees="126" direction="east" minutes="17.304" orientation="longitude" precision="1" value="126.2884">126°17.304'E</geoCoordinate>
,
<collectingDate id="29388143F8A33CB31608AE5CE2B7F198" value="2008-03-30" valueMax="2008-04-01" valueMin="2008-03-30">30.iii.-01.iv.2008</collectingDate>
, leg.
<collectorName id="B73C491FBFFBE6E89F15523C5CAAFEB7">JH Lourens</collectorName>
(GU 28.069); ZSM.
</materialsCitation>
</paragraph>
<paragraph id="6BA0E5FB1B92DA3CFCB7066F51F72904" pageId="0" pageNumber="57">
<emphasis id="63164467BC076D895A42BE642C778B0C" bold="true" pageId="0" pageNumber="57">
<typeStatus id="A2E6CA9F3A7F6BEEB62A2AB4E92CCFE7" type="paratype">Paratypes</typeStatus>
</emphasis>
[
<specimenCount id="DE8C4E55ED813C05788376002A016ABB" count="4" type="male">4 ♂</specimenCount>
]:
<materialsCitation id="92989700B0C572A671C83D9BDA6A7886" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Philippines" location="Mindanao Island" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="894036C9FE4A4205B2E509AEE12A9DBB" name="Philippines">Philippines</collectingCountry>
:
<location id="A255FD0A6869BDEC23F6DCAABD4463BD" LSID="urn:lsid:plazi:treatment:D08316C4D7135606AAB3BFC4B9237282:A255FD0A6869BDEC23F6DCAABD4463BD" country="Philippines" name="Mindanao Island">Mindanao Island</location>
•
<specimenCount id="04F2D4DB00E869850E597A2C7316E1FD" count="1" type="male">1 ♂</specimenCount>
</materialsCitation>
;
<materialsCitation id="F32127C382A465FAC2915BE485D03277" collectingDate="2008-03-30" collectingDateMax="2008-04-01" collectingDateMin="2008-03-30" collectorName="SE Mindanao, JH Lourens" country="Philippines" elevation="90" latitude="7.727783" location="Aliwagwag Primary forest" longLatPrecision="1" longitude="126.2884" specimenCount="3" specimenCount-male="3" stateProvince="Davao Oriental" typeStatus="paratype">
<collectingCountry id="59A0386D9BB3BBF33A3BF0C78791CF9C" name="Philippines">Philippines</collectingCountry>
,
<collectorName id="B88E2385E267E7C831E0D35CA9C6ACA1">SE Mindanao</collectorName>
,
<collectingRegion id="F2B68F6AF3C6E560CFFDC4BA61EBDF06" country="Philippines" name="Davao Oriental">Davao Oriental</collectingRegion>
<location id="343927A08032E4A4962BBCE902470373" LSID="urn:lsid:plazi:treatment:D08316C4D7135606AAB3BFC4B9237282:343927A08032E4A4962BBCE902470373" country="Philippines" latitude="7.727783" longLatPrecision="1" longitude="126.2884" name="Aliwagwag Primary forest" stateProvince="Davao Oriental">Aliwagwag Primary forest</location>
,
<quantity id="9FF479F2FA8B698961C6370AEAC8A509" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
<elevation id="77147595F1FE255EF73BEC515BA35B65" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
</quantity>
,
<geoCoordinate id="30F39A9E8E67FDDFAA0FD187BF2557D7" degrees="07" direction="north" minutes="43.667" orientation="latitude" precision="1" value="7.727783">07°43.667'N</geoCoordinate>
,
<geoCoordinate id="5D36C56015BB2123D67EC9A78C495D5B" degrees="126" direction="east" minutes="17.304" orientation="longitude" precision="1" value="126.2884">126°17.304'E</geoCoordinate>
,
<collectingDate id="CF40C4605F64D7242E12CD363762F70E" value="2008-03-30" valueMax="2008-04-01" valueMin="2008-03-30">30.iii.-01.iv.2008</collectingDate>
, leg.
<collectorName id="DE8065006F2DA21A97543F85092319F3">JH Lourens</collectorName>
(GU 28.075); ZSM •
<specimenCount id="E7046FCE718E3D8B42F2179187921584" count="3" type="male">3 ♂</specimenCount>
</materialsCitation>
;
<materialsCitation id="C8171F32E61BBE808A00AEB7BDC67810" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="N. Mindanao, S. Gundorov" country="Philippines" elevation="300" location="Mt. Malasag" municipality="Misamis prov." specimenCount="1" stateProvince="Davao Oriental" typeStatus="paratype">
<collectorName id="68097AA8ECCB0D94EB1B8956E660AA7A">N. Mindanao</collectorName>
, N.
<collectingMunicipality id="0E6509D427CF93802C8464148ABEC1BF">Misamis prov.</collectingMunicipality>
<location id="E589B2D48D69E593C98D846844BAAD13" LSID="urn:lsid:plazi:treatment:D08316C4D7135606AAB3BFC4B9237282:E589B2D48D69E593C98D846844BAAD13" country="Philippines" municipality="Misamis prov." name="Mt. Malasag" stateProvince="Davao Oriental">Mt. Malasag</location>
,
<quantity id="FEDE920F3F0EBAE2B7035FA78168ED2F" metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">
<elevation id="F2E34254067246F8CB57F1010D856516" metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">300 m</elevation>
</quantity>
, 8-15.02 [ii].1996, leg.
<collectorName id="25F73FF6D31930FC59A156694679F074">S. Gundorov</collectorName>
; ZSM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9106C90BEE0449017EBDB30342C8B33B" pageId="0" pageNumber="57" type="diagnosis">
<paragraph id="47AE3091AFC3CD7D5A2302999ED460E6" pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph id="001FDC228BB1E4BD554A3C2C29AE89B5" pageId="0" pageNumber="57">
Externally, the male of
<taxonomicName id="E8B8CDA668DFB29D066B8DA8BF44DDB6" lsidName="C. zolotuhini" pageId="0" pageNumber="57" rank="species" species="zolotuhini">
<emphasis id="AA80D8E06AA9BD9274F21D4468388790" italics="true" pageId="0" pageNumber="57">C. zolotuhini</emphasis>
</taxonomicName>
sp. nov. is easily distinguished from other
<taxonomicName id="320ACAB7B852F1E2DAE3070B03FF62F6" authorityName="Swinhoe" authorityYear="1922" class="Insecta" family="Erebidae" genus="Carriola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carriola" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="EB9E8A2EE5340434928F3484B757A309" italics="true" pageId="0" pageNumber="57">Carriola</emphasis>
</taxonomicName>
by its dark grey colour and the absence of a hyaline window on the hindwings (Figs
<figureCitation id="60DBAB67AB0271B6FC863801280529EA" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">18</figureCitation>
,
<figureCitation id="00D7CAE2D492495B1752B3BE3D0F6850" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">27</figureCitation>
). The male genitalia differ from those of the related species by the broad superuncus, the width of which is in a ratio of 2:1 to the length (Fig.
<figureCitation id="98611BE9C399A20AAF1703C19052E4C8" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">36c</figureCitation>
). The valva is rectangular and has the characteristic folds: a curved saccular fold with a downwardly curved process and a broad costal fold. The width of the basal part of the valva has a ratio of 1:2.4 to the length of the valva (Fig.
<figureCitation id="7674A11C056A936B5D37BDC2FD2CBAAA" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">36a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="416C28D7E95A05DC8F9E129401341D00" pageId="0" pageNumber="57" type="description">
<paragraph id="5C8E7E5AE18E01AB5892E0D6A80D98B9" pageId="0" pageNumber="57">Description.</paragraph>
<paragraph id="A2728673AAFEB5B9BA72CE47716978B8" pageId="0" pageNumber="57">
<emphasis id="0DB26EF7A68B8A19045972340FDC08A0" bold="true" italics="true" pageId="0" pageNumber="57">External appearance. Head.</emphasis>
Frons and vertex dark grey and covered with dense, drooping scales, with small tufts at the base of antennae. Eyes large, round. Labial palpi obliquely curved upwards, densely hairy, on underside of the same grey colour as abdomen; dorsally dark brown. Male antennae bipectinate with long branches.
<emphasis id="A4874367FDBB8752CD6102555885480E" bold="true" italics="true" pageId="0" pageNumber="57">Male forewings</emphasis>
(Figs
<figureCitation id="0F5CBF7742B2DE1DA58370093787DFD3" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">18</figureCitation>
,
<figureCitation id="372DE3014EF59CAFFDC3EDFF75D57019" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">27a, b</figureCitation>
) triangular with pointed tip. Ground colour dark grey, with central hyaline window. Length is 13-13.5 mm (13.2 mm in the holotype). The ratio between the length of forewing margin and costa is 1.2:1. Central hyaline window occupies central cell and covers up to half the length of the cells between veins M2-M3, M3-CuA1, CuA1-CuA2 and CuA2-1+2A. Hyaline windows covered by sparse black scales, which make them appear blurred and without clear boundaries, and bordered by faintly visible antemedial and subterminal lines of black scales; discal spot chevron-shaped. Underside of forewing with dark grey, monochrome background (Fig.
<figureCitation id="52FD685F7B1BE968674B42A1087B0015" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">27b</figureCitation>
).
<emphasis id="7A6151174D1A440E957E5BFED80AAD5E" bold="true" italics="true" pageId="0" pageNumber="57">Male hindwings</emphasis>
(Figs
<figureCitation id="FF0B01A306821AB70E23D0467F5CC567" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">18</figureCitation>
,
<figureCitation id="ECC0503240CD1454D00A1429610DCD0D" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">27a, b</figureCitation>
) same coloured as forewing without hyaline window, costal margin brownish.
<emphasis id="07794E48ABED5BB27C2CC0BF6CFDF7B4" bold="true" italics="true" pageId="0" pageNumber="57">Male genitalia</emphasis>
(Figs
<figureCitation id="5F5A9DEF18DF724BFB8FC5FF6DBCFED8" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">36a, b, c</figureCitation>
). Superuncus very broad, its width in a ratio of 2:1 to length, slightly divided into two lobes at apex (Fig.
<figureCitation id="4C4CC8759CB5D3290C40969BFCBEB334" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">36c</figureCitation>
). Uncus (Fig.
<figureCitation id="AA751AA02BEF2F20C2EC4858EB98E5E9" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">36a</figureCitation>
) short, with slightly constricted base and apically rounded. Valva rectangular with sclerotised curved fold originating from base of sacculus and ending in downward curved sclerotised pointed process extending to distal ⅓ of valva. Costal margin of valva with additional broad sclerotised pocket-like fold. The ratio of the width of the basal part of the valva to the length of the valva is approximately 1:2.4. Juxta A-shaped, triangular, ratio length to width as 1.5:1. Aedeagus short, strongly bevelled at its anterior and posterior end. Everted vesica round, without cornuti, covered with very small spinules (Fig.
<figureCitation id="447ACE48E4C19BC1151DD4622143DA47" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">36b</figureCitation>
).
<emphasis id="0A31D7A303FDB126A6D6DD7A7F836C20" bold="true" italics="true" pageId="0" pageNumber="57">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="76F400E8476DC37F3A2A7F89B854C79B" pageId="0" pageNumber="57" type="biology_ecology">
<paragraph id="0851019B21AC8D9090DCBEAFC6F7AD6D" pageId="0" pageNumber="57">Biology.</paragraph>
<paragraph id="6A4EECADDA43B145F88006C3CBF2D83B" pageId="0" pageNumber="57">Unknown. The flight period is from February to April. The species occurs in wooded lowlands up to an elevation of 300 m.a.s.l.</paragraph>
</subSubSection>
<subSubSection id="C8AC516C8588899D6B99F557273B5404" pageId="0" pageNumber="57" type="distribution">
<paragraph id="B7042867A843D02B27A98EA527B81BB3" pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph id="86D5D4FCBB3D87E4F633D9A5783D996B" pageId="0" pageNumber="57">
(Fig.
<figureCitation id="ABE7D9FC67FA430A5D48AF9514FFF3B0" captionStart="Figures 45–53" captionStartId="F7" captionText="Figures 45 - 53. Distribution maps of Carriola species. 45. C. ecnomoda (Swinhoe, 1907); 46. C. thyridophora (Hampson, [1893]); 47. C. witti sp. nov.; 48. C. seminsula (Strand, 1914); 49. C. saturnioides (Snellen, 1879); 50. C. polyakovi sp. nov.; 51. C. shorokhovi sp. nov.; 52. C. fenestrata (Hampson, [1893]); 53. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures45-53" httpUri="https://binary.pensoft.net/fig/1003326" pageId="0" pageNumber="57">53</figureCitation>
). Philippines (Mindanao).
</paragraph>
</subSubSection>
<subSubSection id="10C44740A50E5B449D210FF3AC9DCEAD" pageId="0" pageNumber="57" type="etymology">
<paragraph id="DE7E0E2E7BC1B554D579B60DE6EC0F27" pageId="0" pageNumber="57">Etymology.</paragraph>
<paragraph id="C3FE5B920BD5B40C8AA261A54B2D9AC6" pageId="0" pageNumber="57">This species is dedicated to the memory of Dr. Vadim V. Zolotuhin, a famous lepidopterist, whose kind help and support was always invaluable to the authors.</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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