treatments-xml/data/CF/BE/FA/CFBEFA1D9678E75A54D66E9AB3CF3401.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.78.800" ID-GBIF-Dataset="824a277f-3e29-43cb-b3b4-797aa5895c03" ID-PMC="PMC3088061" ID-Pensoft-Pub="1313-2970-78-15" ID-PubMed="21594154" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-78-15" ModsDocOrigin="ZooKeys 78" ModsDocTitle="On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)" checkinTime="1451250817608" checkinUser="pensoft" docAuthor="Martin, Fikacek, Hajek, Jiri &amp; Schmied, Heiko" docDate="2011" docId="CFBEFA1D9678E75A54D66E9AB3CF3401" docLanguage="en" docName="ZooKeys 78: 15-25" docOrigin="ZooKeys 78" docSource="http://dx.doi.org/10.3897/zookeys.78.800" docTitle="Escheria crassipunctata Foerster 1891" docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="0E71FFE5A5468A341831690CFF9AFFD7" masterDocTitle="On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)" masterLastPageNumber="25" masterPageNumber="15" pageNumber="19" updateTime="1668166156320" updateUser="ExternalLinkService">
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<mods:title>On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Martin, Fikacek</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hajek, Jiri</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schmied, Heiko</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>78</mods:number>
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<mods:start>15</mods:start>
<mods:end>25</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.78.800</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.78.800</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-78-15</mods:identifier>
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<treatment ID-GBIF-Taxon="159361187" LSID="urn:lsid:plazi:treatment:CFBEFA1D9678E75A54D66E9AB3CF3401" httpUri="http://treatment.plazi.org/id/CFBEFA1D9678E75A54D66E9AB3CF3401" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<subSubSection pageId="4" pageNumber="19" type="nomenclature">
<paragraph pageId="4" pageNumber="19">
<taxonomicName authority="Foerster, 1891" authorityName="Foerster" authorityYear="1891" class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">
Escheria crassipunctata
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1891
</taxonomicName>
Fig. 7
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="19" type="reference_group">
<paragraph pageId="4" pageNumber="19">
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des &quot; Plattigen Steinmergels &quot; von Brunnstatt" volume="5" year="1891">
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
1891
</bibRefCitation>
: 364, plate XI, Fig. 11 (original description from Brunstatt);
<bibRefCitation author="Handlirsch, A" journalOrPublisher="Wilhelm Engelmann, Leipzig" pageId="8" pageNumber="23" title="Die fossilen Insekten und die Phylogenie der rezenten Formen" volumeTitle="Ein Handbuch fuer Palaeontologen und Zoologen" year="1908">Handlirsch 1908</bibRefCitation>
: 767 (catalogue).
</paragraph>
<paragraph pageId="4" pageNumber="19">
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius crassipunctatus" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctatus">Hydrobius crassipunctatus</taxonomicName>
:
<bibRefCitation author="Theobald, N" journalOrPublisher="G. Thomas, Nancy" pageId="8" pageNumber="23" title="Les insectes fossiles des terrains oligocenes de France" year="1937">
<normalizedToken originalValue="Théobald">Theobald</normalizedToken>
1937
</bibRefCitation>
: 169, plate II, Fig. 28 (transferred to
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Hydrobius</taxonomicName>
, referred from Kleinkems);
<bibRefCitation author="Hansen, M" journalOrPublisher="Apollo Books, Stenstrup" pageId="8" pageNumber="23" title="World Catalogue of Insects. Volume 2. Hydrophiloidea (s. str.) (Coleoptera)" year="1999">Hansen 1999</bibRefCitation>
: 319 (catalogue).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="taxonomic notes">
<paragraph pageId="4" pageNumber="19">Taxonomic notes.</paragraph>
<paragraph pageId="4" pageNumber="19">
As in the case of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus convexus" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="convexus">Copelatus convexus</taxonomicName>
,
<bibRefCitation author="Theobald, N" journalOrPublisher="G. Thomas, Nancy" pageId="8" pageNumber="23" title="Les insectes fossiles des terrains oligocenes de France" year="1937">
<normalizedToken originalValue="Théobald">Theobald</normalizedToken>
(1937)
</bibRefCitation>
transferred
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
to the hydrophilid genus
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Hydrobius</taxonomicName>
and assigned fossil specimen no. R 707 from the locality of Kleinkems (deposited in NHMB) to this species. We have examined the specimen from Kleinkems for this study (Figs 10-11) but we cannot confirm that it is conspecific with
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
for the following reasons: (i) the elytra are slightly constricted sub-basally in the specimen from Kleinkems, but evenly rounded laterally in
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
; (ii) the body outline is distinctly interrupted between the pronotum and the elytra, but it is uninterrupted in
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
, (iii) eyes are large and globular in the specimen from Kleinkems, but relatively small in
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
. A more detailed comparison is impossible as the holotype of
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
is lost and was moreover preserved in dorsal view based on the drawing by
<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des &quot; Plattigen Steinmergels &quot; von Brunnstatt" volume="5" year="1891">
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
(1891)
</bibRefCitation>
, whereas the specimen from Kleinkems is preserved in ventral view.
</paragraph>
<paragraph pageId="4" pageNumber="19">
Based on the original drawing by
<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des &quot; Plattigen Steinmergels &quot; von Brunnstatt" volume="5" year="1891">
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
(1891)
</bibRefCitation>
,
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
does not bear any synapomorphy of the
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
. For this reason, the species is removed from the fossil record of the
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
and is placed in
<taxonomicName class="Insecta" family="Corydiidae" genus="Polyphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphaga" order="Blattodea" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Polyphaga</taxonomicName>
incertae sedis.
</paragraph>
<paragraph pageId="5" pageNumber="20">
<pageBreakToken pageId="5" pageNumber="20" start="start">Specimen</pageBreakToken>
no. R707 from Kleinkems does not bear any synapomorphy of the
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
, and moreover bears a combination of characters which excludes its assignment to the
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
: (i) prosternal process wide, (ii) mesocoxal cavities rather wide apart, and (iii) eyes large and globular. The preserved characters of this specimen do not allow an unambiguous family assignment (see
<bibRefCitation author="Lawrence, JF" journalOrPublisher="CSIRO Publishing, Melbourne" pageId="8" pageNumber="23" title="Beetles of the world. A key and information system for families and subfamilies. CD-ROM, Version 1.0 for MS-Windows" year="1999">Lawrence et al. 1999</bibRefCitation>
).
</paragraph>
<caption pageId="5" pageNumber="20">
<paragraph pageId="5" pageNumber="20">
Figures 7-12. 7
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1891, original illustration of the holotype 8
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria dimidiata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="dimidiata">Escheria dimidiata</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1891, original illustration of the holotype 9
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria punctulata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="punctulata">Escheria punctulata</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1891, original illustration of the holotype 10-12 specimen NHMB R707 (10 photograph 11 drawing 12 original drawing by
<bibRefCitation author="Theobald, N" journalOrPublisher="G. Thomas, Nancy" pageId="8" pageNumber="23" title="Les insectes fossiles des terrains oligocenes de France" year="1937">
<normalizedToken originalValue="Théobald">Theobald</normalizedToken>
(1937)
</bibRefCitation>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>