161 lines
14 KiB
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161 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.78.800" ID-GBIF-Dataset="824a277f-3e29-43cb-b3b4-797aa5895c03" ID-PMC="PMC3088061" ID-Pensoft-Pub="1313-2970-78-15" ID-PubMed="21594154" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-78-15" ModsDocOrigin="ZooKeys 78" ModsDocTitle="On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)" checkinTime="1451250817608" checkinUser="pensoft" docAuthor="Martin, Fikacek, Hajek, Jiri & Schmied, Heiko" docDate="2011" docId="CFBEFA1D9678E75A54D66E9AB3CF3401" docLanguage="en" docName="ZooKeys 78: 15-25" docOrigin="ZooKeys 78" docSource="http://dx.doi.org/10.3897/zookeys.78.800" docTitle="Escheria crassipunctata Foerster 1891" docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="0E71FFE5A5468A341831690CFF9AFFD7" masterDocTitle="On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)" masterLastPageNumber="25" masterPageNumber="15" pageNumber="19" updateTime="1668166156320" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martin, Fikacek</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hajek, Jiri</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schmied, Heiko</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>78</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>25</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.78.800</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.78.800</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-78-15</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159361187" LSID="urn:lsid:plazi:treatment:CFBEFA1D9678E75A54D66E9AB3CF3401" httpUri="http://treatment.plazi.org/id/CFBEFA1D9678E75A54D66E9AB3CF3401" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
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<subSubSection pageId="4" pageNumber="19" type="nomenclature">
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<paragraph pageId="4" pageNumber="19">
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<taxonomicName authority="Foerster, 1891" authorityName="Foerster" authorityYear="1891" class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">
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Escheria crassipunctata
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<normalizedToken originalValue="Förster">Foerster</normalizedToken>
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, 1891
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</taxonomicName>
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Fig. 7
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="19" type="reference_group">
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<paragraph pageId="4" pageNumber="19">
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des " Plattigen Steinmergels " von Brunnstatt" volume="5" year="1891">
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<normalizedToken originalValue="Förster">Foerster</normalizedToken>
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1891
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</bibRefCitation>
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: 364, plate XI, Fig. 11 (original description from Brunstatt);
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<bibRefCitation author="Handlirsch, A" journalOrPublisher="Wilhelm Engelmann, Leipzig" pageId="8" pageNumber="23" title="Die fossilen Insekten und die Phylogenie der rezenten Formen" volumeTitle="Ein Handbuch fuer Palaeontologen und Zoologen" year="1908">Handlirsch 1908</bibRefCitation>
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: 767 (catalogue).
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</paragraph>
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<paragraph pageId="4" pageNumber="19">
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius crassipunctatus" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctatus">Hydrobius crassipunctatus</taxonomicName>
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:
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<bibRefCitation author="Theobald, N" journalOrPublisher="G. Thomas, Nancy" pageId="8" pageNumber="23" title="Les insectes fossiles des terrains oligocenes de France" year="1937">
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<normalizedToken originalValue="Théobald">Theobald</normalizedToken>
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1937
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</bibRefCitation>
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: 169, plate II, Fig. 28 (transferred to
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Hydrobius</taxonomicName>
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, referred from Kleinkems);
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<bibRefCitation author="Hansen, M" journalOrPublisher="Apollo Books, Stenstrup" pageId="8" pageNumber="23" title="World Catalogue of Insects. Volume 2. Hydrophiloidea (s. str.) (Coleoptera)" year="1999">Hansen 1999</bibRefCitation>
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: 319 (catalogue).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="taxonomic notes">
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<paragraph pageId="4" pageNumber="19">Taxonomic notes.</paragraph>
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<paragraph pageId="4" pageNumber="19">
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As in the case of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus convexus" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="convexus">Copelatus convexus</taxonomicName>
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,
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<bibRefCitation author="Theobald, N" journalOrPublisher="G. Thomas, Nancy" pageId="8" pageNumber="23" title="Les insectes fossiles des terrains oligocenes de France" year="1937">
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<normalizedToken originalValue="Théobald">Theobald</normalizedToken>
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(1937)
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</bibRefCitation>
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transferred
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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to the hydrophilid genus
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Hydrobius</taxonomicName>
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and assigned fossil specimen no. R 707 from the locality of Kleinkems (deposited in NHMB) to this species. We have examined the specimen from Kleinkems for this study (Figs 10-11) but we cannot confirm that it is conspecific with
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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for the following reasons: (i) the elytra are slightly constricted sub-basally in the specimen from Kleinkems, but evenly rounded laterally in
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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; (ii) the body outline is distinctly interrupted between the pronotum and the elytra, but it is uninterrupted in
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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, (iii) eyes are large and globular in the specimen from Kleinkems, but relatively small in
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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. A more detailed comparison is impossible as the holotype of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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is lost and was moreover preserved in dorsal view based on the drawing by
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<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des " Plattigen Steinmergels " von Brunnstatt" volume="5" year="1891">
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<normalizedToken originalValue="Förster">Foerster</normalizedToken>
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(1891)
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</bibRefCitation>
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, whereas the specimen from Kleinkems is preserved in ventral view.
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</paragraph>
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<paragraph pageId="4" pageNumber="19">
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Based on the original drawing by
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<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des " Plattigen Steinmergels " von Brunnstatt" volume="5" year="1891">
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<normalizedToken originalValue="Förster">Foerster</normalizedToken>
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(1891)
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</bibRefCitation>
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,
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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does not bear any synapomorphy of the
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
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. For this reason, the species is removed from the fossil record of the
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
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and is placed in
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<taxonomicName class="Insecta" family="Corydiidae" genus="Polyphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphaga" order="Blattodea" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">Polyphaga</taxonomicName>
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incertae sedis.
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</paragraph>
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<paragraph pageId="5" pageNumber="20">
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<pageBreakToken pageId="5" pageNumber="20" start="start">Specimen</pageBreakToken>
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no. R707 from Kleinkems does not bear any synapomorphy of the
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
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, and moreover bears a combination of characters which excludes its assignment to the
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
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: (i) prosternal process wide, (ii) mesocoxal cavities rather wide apart, and (iii) eyes large and globular. The preserved characters of this specimen do not allow an unambiguous family assignment (see
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<bibRefCitation author="Lawrence, JF" journalOrPublisher="CSIRO Publishing, Melbourne" pageId="8" pageNumber="23" title="Beetles of the world. A key and information system for families and subfamilies. CD-ROM, Version 1.0 for MS-Windows" year="1999">Lawrence et al. 1999</bibRefCitation>
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).
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</paragraph>
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<caption pageId="5" pageNumber="20">
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<paragraph pageId="5" pageNumber="20">
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Figures 7-12. 7
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria crassipunctata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="crassipunctata">Escheria crassipunctata</taxonomicName>
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<normalizedToken originalValue="Förster">Foerster</normalizedToken>
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, 1891, original illustration of the holotype 8
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria dimidiata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="dimidiata">Escheria dimidiata</taxonomicName>
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<normalizedToken originalValue="Förster">Foerster</normalizedToken>
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, 1891, original illustration of the holotype 9
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria punctulata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="punctulata">Escheria punctulata</taxonomicName>
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<normalizedToken originalValue="Förster">Foerster</normalizedToken>
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, 1891, original illustration of the holotype 10-12 specimen NHMB R707 (10 photograph 11 drawing 12 original drawing by
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<bibRefCitation author="Theobald, N" journalOrPublisher="G. Thomas, Nancy" pageId="8" pageNumber="23" title="Les insectes fossiles des terrains oligocenes de France" year="1937">
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<normalizedToken originalValue="Théobald">Theobald</normalizedToken>
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(1937)
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</bibRefCitation>
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).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |