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<document id="575FD20E8BF4E1C31A5F87AEFD4CA601" ID-DOI="10.11646/zootaxa.3718.1.1" ID-GBIF-Dataset="038e61f6-f50a-439a-bc5a-249a2cb4164f" ID-ISSN="1175-5326" ID-Zenodo-Dep="284980" ID-ZooBank="B770D766-62D2-472C-A8F2-CF994F35C95D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460685340328" checkinUser="plazi" docAuthor="Do Nascimento, Jeane M. C. &amp; Salles, Frederico F." docDate="2013" docId="CE5587F3FFEA2E5A9496F9FCFCECE91E" docLanguage="en" docName="zt03718p027.pdf" docOrigin="Zootaxa 3718 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hylister obliquus Nascimento &amp; Salles, 2013, sp. nov." docType="treatment" docVersion="6" lastPageNumber="16" masterDocId="326CFF8BFFE12E489401FFA5FF80E821" masterDocTitle="New species of Hermanella complex (Ephemeroptera: Leptophlebiidae) from Brazilian Atlantic Forest" masterLastPageNumber="27" masterPageNumber="1" pageNumber="12" updateTime="1698351042985" updateUser="plazi">
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<mods:titleInfo id="6AA2875845FC2E45AD0BE1E192DE5AED">
<mods:title id="69E9738C396E6459406D5CA84501493D">New species of Hermanella complex (Ephemeroptera: Leptophlebiidae) from Brazilian Atlantic Forest</mods:title>
</mods:titleInfo>
<mods:name id="126768FEBC0294EB75051E0D025CDDC7" type="personal">
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<mods:namePart id="65D60C198409660AA7A41068AAE3231F">Do Nascimento, Jeane M. C.</mods:namePart>
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<mods:namePart id="C10C87EA7B37B7CCD59075A8BE8A5237">Salles, Frederico F.</mods:namePart>
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<mods:title id="FA396236D64EEB14BC507B4AC79BCDE8">Zootaxa</mods:title>
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<mods:date id="9C4C6EFB121C1983913AAB976463C70A">2013</mods:date>
<mods:detail id="1ADCA0D9DD2A4D296BE8063AEB8F1565" type="volume">
<mods:number id="FBCFE8EE78B0DC8571C327D0D39FF00F">3718</mods:number>
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<mods:number id="5857AE3D68297FFA131D236C6F759616">1</mods:number>
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<mods:identifier id="51C56DBC5D836C37A40FAD9A19272DA9" type="DOI">10.11646/zootaxa.3718.1.1</mods:identifier>
<mods:identifier id="9E6876238B438E39B493EF72DBF0583F" type="GBIF-Dataset">038e61f6-f50a-439a-bc5a-249a2cb4164f</mods:identifier>
<mods:identifier id="B5EEF9B0DB132BAFF550A667AD060937" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E1792C389D94D8EA3931E95562DB33F4" type="Zenodo-Dep">284980</mods:identifier>
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</mods:mods>
<treatment id="CE5587F3FFEA2E5A9496F9FCFCECE91E" ID-DOI="http://doi.org/10.5281/zenodo.6146191" ID-GBIF-Taxon="127665968" ID-Zenodo-Dep="6146191" LSID="urn:lsid:plazi:treatment:CE5587F3FFEA2E5A9496F9FCFCECE91E" httpUri="http://treatment.plazi.org/id/CE5587F3FFEA2E5A9496F9FCFCECE91E" lastPageId="18" lastPageNumber="16" pageId="11" pageNumber="12">
<subSubSection id="0EE6656EFFEA2E439496F9FCFE9FEEB5" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="464336E5FFEA2E439496F9FCFE4BEE52" blockId="11.[151,459,1625,1685]" box="[151,459,1625,1651]" pageId="11" pageNumber="12">
<heading id="1D0B8189FFEA2E439496F9FCFE4BEE52" bold="true" box="[151,459,1625,1651]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="7488EAF7FFEA2E439496F9FCFE4BEE52" bold="true" box="[151,459,1625,1651]" pageId="11" pageNumber="12">
<taxonomicName id="81FC4D66FFEA2E439496F9FCFEE6EE52" box="[151,358,1625,1651]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="obliquus" status="sp. nov.">
<emphasis id="7488EAF7FFEA2E439496F9FCFEE6EE52" bold="true" box="[151,358,1625,1651]" italics="true" pageId="11" pageNumber="12">Hylister obliquus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FBB578CFFEA2E43956FF9FCFE4BEE52" box="[366,459,1625,1651]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="464336E5FFEA2E439496F9D9FE9FEEB5" blockId="11.[151,459,1625,1685]" box="[151,287,1660,1685]" pageId="11" pageNumber="12">(Figs. 8–12)</paragraph>
</subSubSection>
<subSubSection id="0EE6656EFFEA2E439496F961FB2EEFF9" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="464336E5FFEA2E439496F961FCF4EF95" blockId="11.[151,1436,1732,2009]" pageId="11" pageNumber="12">
<emphasis id="7488EAF7FFEA2E439496F961FE91EEFC" bold="true" box="[151,273,1732,1757]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Male imago: (1) fore-wing hyaline, brown at base (Figs. 8A, B and 9A, B); (2) subgenital plate yellowish brown, with pointed projection near inner base of forceps (Figs. 8D and 9D); (3) penes divided on distal 1/3 (Fig. 9D); (4) penis lobe with ventral, short, narrow, posteromedially directed spine (Fig. 9D); (5) abdomen with contrasting color pattern (segments II–VI translucent, segment I dark brown, and terga VII–X reddish brown) (Figs. 8A, B). Nymph: (1) clypeus with lateral margin slightly concave, without anteromedial projection; (2) galealacinia with a prominent tusk on inner apical angle (1/3 the apical width of galea-lacinia) (
<figureCitation id="DEC72A60FFEA2E4390B4F8D2FA88EFB1" box="[1205,1288,1911,1936]" captionStart="FIGURE 11" captionStartId="16.[151,250,1974,1996]" captionTargetBox="[223,1368,215,1952]" captionTargetId="figure@16.[194,1392,193,1965]" captionTargetPageId="16" captionText="FIGURE 11. Hylister obliquus, sp. nov., nymphal mouthparts. A) labrum: left: dorsal view, right: ventral view; B) mandible; C) maxilla; D) hypopharynx; E) labium: right: ventral view, left: dorsal view." httpUri="https://zenodo.org/record/284989/files/figure.png" pageId="11" pageNumber="12">Fig. 11</figureCitation>
C); (3) tarsal claws with subapical denticle much larger than others (
<figureCitation id="DEC72A60FFEA2E4396FEF839FCD0EF95" box="[767,848,1948,1973]" captionStart="FIGURE 12" captionStartId="17.[151,250,1995,2017]" captionTargetBox="[250,1339,216,1993]" captionTargetId="figure@17.[221,1365,193,2004]" captionTargetPageId="17" captionText="FIGURE 12. Hylister obliquus, sp. nov., nymph. A) fore leg; B) middle leg; C) hind leg; D) tarsal claw." httpUri="https://zenodo.org/record/284990/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
D).
</paragraph>
<paragraph id="464336E5FFEA2E4394C6F865FB2EEFF9" blockId="11.[151,1436,1732,2009]" box="[199,1198,1984,2009]" pageId="11" pageNumber="12">
<emphasis id="7488EAF7FFEA2E4394C6F865FEDBEFF8" bold="true" box="[199,347,1984,2009]" pageId="11" pageNumber="12">Male imago.</emphasis>
Length: body,
<quantity id="81049B00FFEA2E439605F865FD0EEFF9" box="[516,654,1984,2009]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.5" metricValueMin="6.8" pageId="11" pageNumber="12" unit="mm" value="7.65" valueMax="8.5" valueMin="6.8">6.8–8.5 mm</quantity>
; forewing,
<quantity id="81049B00FFEA2E43970DF865FC16EFF9" box="[780,918,1984,2009]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.5" metricValueMin="6.2" pageId="11" pageNumber="12" unit="mm" value="6.85" valueMax="7.5" valueMin="6.2">6.2–7.5 mm</quantity>
; hind wing, 1.0–
<quantity id="81049B00FFEA2E439052F865FB2AEFF9" box="[1107,1194,1984,2008]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0EE6656EFFED2E4494C6FF32FDA8EF6C" pageId="12" pageNumber="13" type="description">
<paragraph id="464336E5FFED2E4494C6FF32FD37E8D9" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6FF32FE8DE891" bold="true" box="[199,269,151,176]" pageId="12" pageNumber="13">Head.</emphasis>
Dorsal region orangish yellow with brown marks; ventral region dark brown. Upper portion of compound eye orangish brown, lower portion black; ocelli whitish, surrounded by dark brown.
<emphasis id="7488EAF7FFED2E4490BEFF19FA9CE8F2" box="[1215,1308,188,211]" italics="true" pageId="12" pageNumber="13">Antenna</emphasis>
. Scape and pedicel dark brown, flagellum translucent white.
</paragraph>
<paragraph id="464336E5FFED2E4494C6FEA1FC3FEA1C" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6FEA1FEA6E93C" bold="true" box="[199,294,260,285]" pageId="12" pageNumber="13">Thorax.</emphasis>
Pronotum yellowish brown, with medial strip, submedial mark and lateral margin black; mesonotum brown, medioparapsidal, mediolongitudinal and anterolateral parapsidal sutures yellow, scuto-scutellar impression whitish, scutellum blackish brown; metanotum yellowish brown with brown areas; pleurae brown, with sclerites dark brown, membranous region lighter; sterna dark brown, prosternal carina narrow (Fig. 8C).
<emphasis id="7488EAF7FFED2E4490C7FED4FA8BE9A9" box="[1222,1291,369,392]" italics="true" pageId="12" pageNumber="13">Wings</emphasis>
(Figs. 8A, B and 9A–C). Membrane hyaline, brown at base. Forewing with longitudinal veins light yellow and cross veins translucent; hind wing with longitudinal and cross veins translucent, vein C brown in basal third.
<emphasis id="7488EAF7FFED2E4490E0FE1CFA96E9F1" box="[1249,1302,441,464]" italics="true" pageId="12" pageNumber="13">Legs</emphasis>
. Coxae and trochanters brown washed with black. Leg I: femur orangish brown; tibia dark orange with a basal slender brown band and an apical slender black band; tarsomeres white, with apical black band. Legs II and III: whitish, femora with apical orangish brown band, tibiae with a small apical brown band.
</paragraph>
<paragraph id="464336E5FFED2E4494C6FDE2FCACEB19" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6FDE2FEC3EA41" bold="true" box="[199,323,583,608]" pageId="12" pageNumber="13">Abdomen.</emphasis>
Tergum I dark brown washed with black, terga II–VI translucent, posterior margin orangish brown; terga VII–X reddish brown, terga VII and IX with submedial white mark. Abdominal sternum I brown, sterna II– VI translucent, medial region with gray mark; VII–IX reddish brown.
<emphasis id="7488EAF7FFED2E4497C8FD2AFBB3EA89" box="[969,1075,655,680]" italics="true" pageId="12" pageNumber="13">Genitalia</emphasis>
(Figs. 8D and 9D). Subgenital plate yellowish brown, with pointed projection near inner base of forceps.
<emphasis id="7488EAF7FFED2E4497F2FD10FBCEEAED" box="[1011,1102,693,716]" italics="true" pageId="12" pageNumber="13">Forceps</emphasis>
. Forceps whitish, segment II 0.17 length of segment I, 1.5 length of segment III. Penis divided on distal 1/3, each lobe with a ventral, short, narrow and posteromedially directed spine (Figs. 8D and 9D); penis lobe whitish; spine orangish brown. Caudal filaments white, with apical margin of each segment black.
</paragraph>
<paragraph id="464336E5FFED2E4494C6FCE1FD58EB84" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6FCE1FE20EB7C" bold="true" box="[199,416,836,861]" pageId="12" pageNumber="13">Female subimago.</emphasis>
Length: body, 7.0–
<quantity id="81049B00FFED2E44967DFCE1FD55EB7D" box="[636,725,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="12" pageNumber="13" unit="mm" value="7.8">7.8 mm</quantity>
; fore wing, 8.0–
<quantity id="81049B00FFED2E44978FFCE1FC66EB7D" box="[910,998,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="13" unit="mm" value="8.5">8.5 mm</quantity>
; hind wing, 1.0–
<quantity id="81049B00FFED2E4490A5FCE1FB7CEB7D" box="[1188,1276,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
. Color pattern similar to male imago except for the color ligther. Abdomen completely orangish brown, without the male contrasting color pattern. Sternum IX cleft apically.
</paragraph>
<paragraph id="464336E5FFED2E4494C6FC0AFBA2EBCD" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6FC0AFE00EBE9" bold="true" box="[199,384,943,968]" pageId="12" pageNumber="13">Mature Nymph</emphasis>
. Length: body,
<quantity id="81049B00FFED2E449630FC15FD3BEBE6" box="[561,699,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="7.9" metricValueMin="4.8" pageId="12" pageNumber="13" unit="mm" value="6.35" valueMax="7.9" valueMin="4.8">4.8–7.9 mm</quantity>
; antenna,
<quantity id="81049B00FFED2E44972AFC15FC36EBE6" box="[811,950,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" pageId="12" pageNumber="13" unit="mm" value="3.05" valueMax="3.4" valueMin="2.7">2.7–3.4 mm</quantity>
; cerci, 6.0–
<quantity id="81049B00FFED2E449036FC0AFB10EBE6" box="[1079,1168,943,968]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="12" pageNumber="13" unit="mm" value="8.2">8.2 mm</quantity>
; terminal filament,
<quantity id="81049B00FFED2E44916AFC15FF7EEBCD" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.35" metricValueMin="0.93" pageId="12" pageNumber="13" unit="mm" value="11.4" valueMax="13.5" valueMin="9.3">9.3– 13.5 mm</quantity>
. General coloration dark orangish brown, female lighter (
<figureCitation id="DEC72A60FFED2E449780FC71FC5DEBCD" box="[897,989,980,1005]" captionStart="FIGURE 10" captionStartId="15.[151,250,1653,1675]" captionTargetBox="[171,1425,213,1612]" captionTargetId="figure@15.[154,1439,201,1622]" captionTargetPageId="15" captionText="FIGURE 10. Hylister obliquus, sp. nov. A) and B) habitus of nymph; C) gill IV." httpUri="https://zenodo.org/record/284988/files/figure.png" pageId="12" pageNumber="13">Figs. 10</figureCitation>
A, B).
</paragraph>
<paragraph id="464336E5FFED2E4494C6FC52FB0EED2D" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6FC52FE8DEC31" bold="true" box="[199,269,1015,1040]" pageId="12" pageNumber="13">Head.</emphasis>
Yellowish orangish brown, with black marks. Upper portion of male compound eye dark brown, lower portion black. Eye of female black.
<emphasis id="7488EAF7FFED2E449634FBB9FD13EC12" box="[565,659,1052,1075]" italics="true" pageId="12" pageNumber="13">Antenna</emphasis>
yellowish white. Clypeus without a anteromedial projection, lateral margin strongly concave, maximum width of labrum 1.3 times maximum width of clypeus.
<emphasis id="7488EAF7FFED2E4490CFFB9AFAD1EC79" box="[1230,1361,1087,1112]" italics="true" pageId="12" pageNumber="13">Mouthparts</emphasis>
(
<figureCitation id="DEC72A60FFED2E449165FB9AFF30EC5D" captionStart="FIGURE 11" captionStartId="16.[151,250,1974,1996]" captionTargetBox="[223,1368,215,1952]" captionTargetId="figure@16.[194,1392,193,1965]" captionTargetPageId="16" captionText="FIGURE 11. Hylister obliquus, sp. nov., nymphal mouthparts. A) labrum: left: dorsal view, right: ventral view; B) mandible; C) maxilla; D) hypopharynx; E) labium: right: ventral view, left: dorsal view." httpUri="https://zenodo.org/record/284989/files/figure.png" pageId="12" pageNumber="13">Figs. 11</figureCitation>
A–E). Labrum (
<figureCitation id="DEC72A60FFED2E449565FBC1FE32EC5D" box="[356,434,1124,1149]" captionStart="FIGURE 11" captionStartId="16.[151,250,1974,1996]" captionTargetBox="[223,1368,215,1952]" captionTargetId="figure@16.[194,1392,193,1965]" captionTargetPageId="16" captionText="FIGURE 11. Hylister obliquus, sp. nov., nymphal mouthparts. A) labrum: left: dorsal view, right: ventral view; B) mandible; C) maxilla; D) hypopharynx; E) labium: right: ventral view, left: dorsal view." httpUri="https://zenodo.org/record/284989/files/figure.png" pageId="12" pageNumber="13">Fig. 11</figureCitation>
A) brown, U-shaped anteromedial emargination; with dorsal proximal row of 27–29 long setae interrupted medially; numerous ventral setae directed obliquely to sagittal plane. Mandible (
<figureCitation id="DEC72A60FFED2E4490DDFB22FAABEC81" box="[1244,1323,1159,1184]" captionStart="FIGURE 11" captionStartId="16.[151,250,1974,1996]" captionTargetBox="[223,1368,215,1952]" captionTargetId="figure@16.[194,1392,193,1965]" captionTargetPageId="16" captionText="FIGURE 11. Hylister obliquus, sp. nov., nymphal mouthparts. A) labrum: left: dorsal view, right: ventral view; B) mandible; C) maxilla; D) hypopharynx; E) labium: right: ventral view, left: dorsal view." httpUri="https://zenodo.org/record/284989/files/figure.png" pageId="12" pageNumber="13">Fig. 11</figureCitation>
B) brown, except for central region translucent; outer margin strongly curved, forming a right angle; with 3–4 thick and short setae near to basal articulation. Maxilla (
<figureCitation id="DEC72A60FFED2E449662FB6AFD34ECC9" box="[611,692,1231,1256]" captionStart="FIGURE 11" captionStartId="16.[151,250,1974,1996]" captionTargetBox="[223,1368,215,1952]" captionTargetId="figure@16.[194,1392,193,1965]" captionTargetPageId="16" captionText="FIGURE 11. Hylister obliquus, sp. nov., nymphal mouthparts. A) labrum: left: dorsal view, right: ventral view; B) mandible; C) maxilla; D) hypopharynx; E) labium: right: ventral view, left: dorsal view." httpUri="https://zenodo.org/record/284989/files/figure.png" pageId="12" pageNumber="13">Fig. 11</figureCitation>
C). Galea-lacinia with a prominent tusk on inner apical angle (1/3 the apical width of galea-lacinia). Hypopharynx as in figure 11D. Labium as in figure 11E.
</paragraph>
<paragraph id="464336E5FFED2E4494C6FAB2FEF6EE9C" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6FAB2FEA6ED11" bold="true" box="[199,294,1303,1328]" pageId="12" pageNumber="13">Thorax.</emphasis>
Terga orangish brown, irregularly washed with gray; pronotum with a dark gray submedial stripe, apico-lateral angle with 2–3thick setae; pleura bownish, washed with gray; sterna whitish yellow. Legs. General coloration brown. Leg I (
<figureCitation id="DEC72A60FFED2E4495AFFAFAFE7FED59" box="[430,511,1375,1400]" captionStart="FIGURE 12" captionStartId="17.[151,250,1995,2017]" captionTargetBox="[250,1339,216,1993]" captionTargetId="figure@17.[221,1365,193,2004]" captionTargetPageId="17" captionText="FIGURE 12. Hylister obliquus, sp. nov., nymph. A) fore leg; B) middle leg; C) hind leg; D) tarsal claw." httpUri="https://zenodo.org/record/284990/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
A). Femur with yellowish translucent marks in dorsal region, proximal region gray, outer margin with long and short thick spatulate setae, long spine-like setae and few thin setae; tibia with a medial translucent mark and apical black band, with few setae along outer margins and short spine-like setae in inner margin; tarsi with three apical black bands. Legs II and III (
<figureCitation id="DEC72A60FFED2E44974CFA69FC2CEDC5" box="[845,940,1484,1509]" captionStart="FIGURE 12" captionStartId="17.[151,250,1995,2017]" captionTargetBox="[250,1339,216,1993]" captionTargetId="figure@17.[221,1365,193,2004]" captionTargetPageId="17" captionText="FIGURE 12. Hylister obliquus, sp. nov., nymph. A) fore leg; B) middle leg; C) hind leg; D) tarsal claw." httpUri="https://zenodo.org/record/284990/files/figure.png" pageId="12" pageNumber="13">Figs. 12</figureCitation>
B, C). Femora yellowish translucent with a basal and subapical brown bands. Outer and inner margin and dorsal region with long and short thick spatulate setae, outer margin with few thin setae, absent in femur III. Tibia yellowish translucent with medial brown band; tibia II with few short thick spatulate setae in dorsal surface and numerous thin setae in outer margin, tibia III with short thick spatulate setae along outer and inner margin and few long spatulate setae along outer margin. Tarsal claws (
<figureCitation id="DEC72A60FFED2E4494E6F9DAFEB9EEB9" box="[231,313,1663,1688]" captionStart="FIGURE 12" captionStartId="17.[151,250,1995,2017]" captionTargetBox="[250,1339,216,1993]" captionTargetId="figure@17.[221,1365,193,2004]" captionTargetPageId="17" captionText="FIGURE 12. Hylister obliquus, sp. nov., nymph. A) fore leg; B) middle leg; C) hind leg; D) tarsal claw." httpUri="https://zenodo.org/record/284990/files/figure.png" pageId="12" pageNumber="13">Fig. 12</figureCitation>
D) with subapical denticle much larger than others, 4–6 median denticles subequal in size, without accessory denticles.
</paragraph>
<paragraph id="464336E5FFED2E4494C6F962FD9FEF09" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6F962FEC3EEC1" bold="true" box="[199,323,1735,1760]" pageId="12" pageNumber="13">Abdomen.</emphasis>
Terga dark orangish brown, with a submedial yellow spot. Sterna orangish yellow. Gills (
<figureCitation id="DEC72A60FFED2E44912EF962FA00EEC1" box="[1327,1408,1735,1760]" captionStart="FIGURE 10" captionStartId="15.[151,250,1653,1675]" captionTargetBox="[171,1425,213,1612]" captionTargetId="figure@15.[154,1439,201,1622]" captionTargetPageId="15" captionText="FIGURE 10. Hylister obliquus, sp. nov. A) and B) habitus of nymph; C) gill IV." httpUri="https://zenodo.org/record/284988/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
C) gray, with marks and apical region translucent; plate-like, with apex excavated, bearing long slender processes. Caudal filaments yellowish brown.
</paragraph>
<paragraph id="464336E5FFED2E4494C6F891FDA8EF6C" blockId="12.[151,1437,151,1941]" box="[199,552,1844,1869]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6F891FE4AEF6C" bold="true" box="[199,458,1844,1869]" pageId="12" pageNumber="13">Life cycle association.</emphasis>
Reared.
</paragraph>
</subSubSection>
<subSubSection id="0EE6656EFFED2E4594C6F8F2FE9AEF97" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="etymology">
<paragraph id="464336E5FFED2E4494C6F8F2FE64EFB4" blockId="12.[151,1437,151,1941]" pageId="12" pageNumber="13">
<emphasis id="7488EAF7FFED2E4494C6F8F2FECEEF51" bold="true" box="[199,334,1879,1904]" pageId="12" pageNumber="13">Etymology.</emphasis>
From the Latin word
<taxonomicName id="81FC4D66FFED2E44964FF8F2FD2EEF51" box="[590,686,1879,1904]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="7488EAF7FFED2E44964FF8F2FD2EEF51" box="[590,686,1879,1904]" italics="true" pageId="12" pageNumber="13">obliquus</emphasis>
</taxonomicName>
, oblique; a reference to the inner margin of the apex of the gills, which is obliquely excavated.
</paragraph>
<paragraph id="464336E5FFEC2E459496F824FE9AEF97" blockId="13.[151,1436,1921,1974]" pageId="13" pageNumber="14">
<emphasis id="7488EAF7FFEC2E459496F824FE9FEFB7" bold="true" box="[151,287,1921,1943]" pageId="13" pageNumber="14">FIGUREs 8.</emphasis>
<taxonomicName id="81FC4D66FFEC2E45952BF824FE5CEFB6" box="[298,476,1921,1943]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obliquus" status="sp. nov.">
<emphasis id="7488EAF7FFEC2E45952BF824FE5CEFB6" box="[298,476,1921,1943]" italics="true" pageId="13" pageNumber="14">Hylister obliquus</emphasis>
</taxonomicName>
,
<emphasis id="7488EAF7FFEC2E4595EDF827FDBFEFB6" bold="true" box="[492,575,1922,1943]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="6FBB578CFFEC2E4595EDF827FDBFEFB6" box="[492,575,1922,1943]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male imago.
<emphasis id="7488EAF7FFEC2E4596DAF824FD75EFB9" bold="true" box="[731,757,1921,1944]" pageId="13" pageNumber="14">A)</emphasis>
lateral view;
<emphasis id="7488EAF7FFEC2E45978BF824FC24EFB9" bold="true" box="[906,932,1921,1944]" pageId="13" pageNumber="14">B)</emphasis>
dorsal view;
<emphasis id="7488EAF7FFEC2E459037F824FBD2EFB9" bold="true" box="[1078,1106,1921,1944]" pageId="13" pageNumber="14">C)</emphasis>
prosternum;
<emphasis id="7488EAF7FFEC2E4590DEF824FB79EFB9" bold="true" box="[1247,1273,1921,1944]" pageId="13" pageNumber="14">D)</emphasis>
male genitalia, ventral view.
</paragraph>
</subSubSection>
<subSubSection id="0EE6656EFFEF2E469496F801FE9AEFF8" pageId="14" pageNumber="15" type="description">
<paragraph id="464336E5FFEF2E469496F801FE9AEFF8" blockId="14.[151,1436,1955,2009]" pageId="14" pageNumber="15">
<emphasis id="7488EAF7FFEF2E469496F801FE9DEF98" bold="true" box="[151,285,1956,1978]" pageId="14" pageNumber="15">FIGUREs 9.</emphasis>
<taxonomicName id="81FC4D66FFEF2E469524F801FE55EF9B" box="[293,469,1956,1978]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obliquus" status="sp. nov.">
<emphasis id="7488EAF7FFEF2E469524F801FE55EF9B" box="[293,469,1956,1978]" italics="true" pageId="14" pageNumber="15">Hylister obliquus</emphasis>
</taxonomicName>
,
<emphasis id="7488EAF7FFEF2E4695E5F801FDB5EF98" bold="true" box="[484,565,1956,1977]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="6FBB578CFFEF2E4695E5F801FDB5EF98" box="[484,565,1956,1977]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male imago.
<emphasis id="7488EAF7FFEF2E4696CBF806FD64EF9B" bold="true" box="[714,740,1955,1978]" pageId="14" pageNumber="15">A)</emphasis>
forewing;
<emphasis id="7488EAF7FFEF2E469757F806FCEEEF9B" bold="true" box="[854,878,1955,1978]" pageId="14" pageNumber="15">B)</emphasis>
hind wing;
<emphasis id="7488EAF7FFEF2E4697ECF806FB88EF9B" bold="true" box="[1005,1032,1955,1978]" pageId="14" pageNumber="15">C)</emphasis>
enlarged hind wing;
<emphasis id="7488EAF7FFEF2E4690E2F806FB7DEF9B" bold="true" box="[1251,1277,1955,1978]" pageId="14" pageNumber="15">D)</emphasis>
male genitalia, ventral view.
</paragraph>
</subSubSection>
<caption id="1283666DFFEE2E479496F9D0FC4BEEAA" box="[151,971,1652,1675]" httpUri="https://zenodo.org/record/284988/files/figure.png" pageId="15" pageNumber="16" targetBox="[171,1425,213,1612]" targetPageId="15">
<paragraph id="464336E5FFEE2E479496F9D0FC4BEEAA" blockId="15.[151,971,1652,1675]" box="[151,971,1652,1675]" pageId="15" pageNumber="16">
<emphasis id="7488EAF7FFEE2E479496F9D0FE9FEEAB" bold="true" box="[151,287,1653,1675]" pageId="15" pageNumber="16">FIGURE 10.</emphasis>
<taxonomicName id="81FC4D66FFEE2E479524F9D0FE54EEAA" box="[293,468,1653,1675]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="obliquus" status="sp. nov.">
<emphasis id="7488EAF7FFEE2E479524F9D0FE54EEAA" box="[293,468,1653,1675]" italics="true" pageId="15" pageNumber="16">Hylister obliquus</emphasis>
</taxonomicName>
,
<emphasis id="7488EAF7FFEE2E4795E0F9D0FDD4EEAA" bold="true" box="[481,596,1652,1675]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="6FBB578CFFEE2E4795E0F9D0FDB2EEAB" box="[481,562,1653,1674]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
A)
</emphasis>
and
<emphasis id="7488EAF7FFEE2E479685F9D1FD1CEEAA" bold="true" box="[644,668,1652,1675]" pageId="15" pageNumber="16">B)</emphasis>
habitus of nymph;
<emphasis id="7488EAF7FFEE2E479762F9D1FCFDEEAA" bold="true" box="[867,893,1652,1675]" pageId="15" pageNumber="16">C)</emphasis>
gill IV.
</paragraph>
</caption>
<subSubSection id="0EE6656EFFEE2E4794C6F917FD92EF5E" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="464336E5FFEE2E4794C6F917FD92EF5E" blockId="15.[151,1436,1714,2027]" pageId="15" pageNumber="16">
<emphasis id="7488EAF7FFEE2E4794C6F917FDB7EEEA" bold="true" box="[199,567,1714,1739]" pageId="15" pageNumber="16">
Material examined.
<typeStatus id="99478847FFEE2E4795C1F917FDB1EEEA" box="[448,561,1714,1739]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male imago (reared),
<collectingCountry id="3EEB7675FFEE2E479744F917FC0CEEEA" box="[837,908,1714,1739]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
, Espírito Santo, Sooretama, Rio São José (
<geoCoordinate id="23C85022FFEE2E47918DF916FE9CEECE" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="-19.125862">S 19°07'33.1''</geoCoordinate>
,
<geoCoordinate id="23C85022FFEE2E47952AF972FE56EECE" box="[299,470,1750,1775]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-40.240585">W 40°14'26.1''</geoCoordinate>
),
<quantity id="81049B00FFEE2E4795ECF972FDA2EECE" box="[493,546,1751,1775]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="15" pageNumber="16" unit="m" value="24.0">24m</quantity>
,
<date id="32421025FFEE2E479631F973FD2CEECE" box="[560,684,1750,1775]" pageId="15" pageNumber="16" value="2010-09-09">09/ix/2010</date>
, FF Salles, JMC Nascimento cols. (INPA).
<emphasis id="7488EAF7FFEE2E4790A9F973FAA9EECE" bold="true" box="[1192,1321,1750,1775]" pageId="15" pageNumber="16">
<typeStatus id="99478847FFEE2E4790A9F973FAA4EECE" box="[1192,1316,1750,1775]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
:
</emphasis>
one male imago (reared), three male imagos (light trap), three female (reared), 12 nymphs same date of
<typeStatus id="99478847FFEE2E4790BBF95FFA9CEF32" box="[1210,1308,1786,1811]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(two male, one female, four nymphs at CZNC; one male, one female, four nymphs at IBN; one male, one female, four nymphs at INPA), five male imago, three female
<date id="32421025FFEE2E479660F8E7FD52EF7A" box="[609,722,1858,1883]" pageId="15" pageNumber="16" value="2011-02-14">14/ii/2011</date>
(one male, one female at CZNC; one male, one female at IBN; three male, one female at DZRJ).
</paragraph>
</subSubSection>
<subSubSection id="0EE6656EFFEE2E5A94C6F82FFCECE91E" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16" type="discussion">
<paragraph id="464336E5FFEE2E5A94C6F82FFCECE91E" blockId="15.[151,1436,1714,2027]" lastBlockId="18.[151,1436,151,320]" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
<emphasis id="7488EAF7FFEE2E4794C6F82FFECFEF82" bold="true" box="[199,335,1930,1955]" pageId="15" pageNumber="16">Comments.</emphasis>
The imago of
<taxonomicName id="81FC4D66FFEE2E4795FDF829FD0DEF82" box="[508,653,1930,1955]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="obliquus" status="sp. nov.">
<emphasis id="7488EAF7FFEE2E4795FDF829FD0DEF82" box="[508,653,1930,1955]" italics="true" pageId="15" pageNumber="16">Hy. obliquus</emphasis>
</taxonomicName>
<emphasis id="7488EAF7FFEE2E479697F82FFD71EF82" bold="true" box="[662,753,1930,1955]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="6FBB578CFFEE2E479697F82FFD71EF82" box="[662,753,1930,1955]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
appears very similar to
<taxonomicName id="81FC4D66FFEE2E47900BF829FABCEF82" authority="Kluge, 2007" authorityName="Kluge" authorityYear="2007" box="[1034,1340,1930,1955]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="chimaera">
<emphasis id="7488EAF7FFEE2E47900BF829FB24EF82" box="[1034,1188,1930,1955]" italics="true" pageId="15" pageNumber="16">Hy. chimaera</emphasis>
Kluge, 2007
</taxonomicName>
, sharing with this species the same pattern of coloration on body and wings. However, both species can be distinguished by the styliger plate, which has a pair of pointed projections near base of forceps in the new species and a pair of large protuberances, each bearing a very small non-sclerotized point on its apex, in
<taxonomicName id="81FC4D66FFF32E5A9015FF3CFB2DE891" box="[1044,1197,151,176]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chimaera">
<emphasis id="7488EAF7FFF32E5A9015FF3CFB2DE891" box="[1044,1197,151,176]" italics="true" pageId="18" pageNumber="19">Hy. chimaera</emphasis>
</taxonomicName>
. The nymphs can be distinguished based on the presence, in the new species, of a subapical denticle much larger than others in the tarsal claw. Besides that, nymphs of the new species possess a unique gill shape. While in
<taxonomicName id="81FC4D66FFF32E5A904EFF44FEB8E93D" authority="Dominguez &amp; Flowers, 1989" authorityName="Dominguez &amp; Flowers" authorityYear="1989" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="7488EAF7FFF32E5A904EFF44FB74E8D9" box="[1103,1268,223,248]" italics="true" pageId="18" pageNumber="19">Hy. plaumanni</emphasis>
Domínguez &amp; Flowers, 1989
</taxonomicName>
and
<taxonomicName id="81FC4D66FFF32E5A9571FEA0FD85E93C" box="[368,517,260,285]" class="Insecta" family="Leptophlebiidae" genus="Hylister" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="chimerae">
<emphasis id="7488EAF7FFF32E5A9571FEA0FD85E93C" box="[368,517,260,285]" italics="true" pageId="18" pageNumber="19">Hy. chimerae</emphasis>
</taxonomicName>
the apex is truncate, bearing long slender processes, on the new species the apex also bears these processes, but it is excavate rather than truncate.
</paragraph>
</subSubSection>
</treatment>
</document>
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