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<document id="F416D7D02285250B8B1CBE1EDCF17D53" ID-DOI="10.1590/1806-9665-RBENT-2023-0044" ID-ISSN="1806-9665" ID-Zenodo-Dep="13197554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722638410007" checkinUser="felipe" docAuthor="Tavares, Leon Gustavo de Miranda, Marquez-López, Yesenia, Machado, Renato Jose Pires, Martins, Caleb Califre &amp; Contreras-Ramos, Atilano" docDate="2023" docId="CE2CD627EC7AFFDBFFA8FBF8C59B1C43" docLanguage="en" docName="RevBrasEntomol.67.spe.e20230044.pdf" docOrigin="Revista Brasileira de Entomologia (e 20230044) 67" docSource="http://dx.doi.org/10.1590/1806-9665-rbent-2023-0044" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Menkeleon bellulus" docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="3215AE5FEC72FFD3FFD9FFFAC0051D1E" masterDocTitle="Antlions of formerly recognized tribe Gnopholeontini (Neuroptera: Myrmeleontidae: Brachynemurini) from Peninsula of Baja California, with a new species of Tyttholeon Adams" masterLastPageNumber="17" masterPageNumber="1" pageNumber="9" updateTime="1722686643456" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3DFD8DB9A0FD1A3A1A2A7C3F6BBE1759">Antlions of formerly recognized tribe Gnopholeontini (Neuroptera: Myrmeleontidae: Brachynemurini) from Peninsula of Baja California, with a new species of Tyttholeon Adams</mods:title>
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<mods:namePart id="5F2A706C390943F0D8A5C04E5332F328">Tavares, Leon Gustavo de Miranda</mods:namePart>
<mods:affiliation id="BCF92FF618A00E619D990A119EE5B2AA">Universidade Federal do Paraná (DZUP), Departamento de Zoologia, Curitiba, PR, Brasil.</mods:affiliation>
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<mods:namePart id="D934E0529C6970855087B41C51367C6E">Marquez-López, Yesenia</mods:namePart>
<mods:affiliation id="060A64F6455AC6FCA09B643A73581914">Universidad Autónoma Metropolitana-Iztapalapa, Departamento de Biología, Ciudad de México, México.</mods:affiliation>
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<mods:namePart id="8E4BFBDFA74330C82A98D2ECBF211B09">Machado, Renato Jose Pires</mods:namePart>
<mods:affiliation id="74C546CAFFB7B82769AD0829BAC4F107">Universidade Federal do Paraná (DZUP), Departamento de Zoologia, Curitiba, PR, Brasil.</mods:affiliation>
</mods:name>
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<mods:roleTerm id="C0B337FA7E01F4CE461BF85E29FD4488">Author</mods:roleTerm>
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<mods:namePart id="C492895115157E50D376226749E7FC1A">Martins, Caleb Califre</mods:namePart>
<mods:affiliation id="9BF278D9C72FD9E6D900F8728CCBF2C2">Universidade Estadual do Maranhão, Centro de Estudos Superiores de Caxias, Departamento de Química e Biologia, Caxias, MA, Brasil.</mods:affiliation>
</mods:name>
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<mods:namePart id="D3F1C63051F66A06075A7949375171AF">Contreras-Ramos, Atilano</mods:namePart>
<mods:affiliation id="8420DA95FF8A89AA969AB38DF2FEF473">* &amp; Universidad Nacional Autónoma de México, Instituto de Biología, Departamento de Zoología, Colección Nacional de Insectos, Ciudad de México, México. urn: lsid: zoobank. org: pub: 5978 D 8 C 0 - E 25 F- 452 A-A 526 - C 37 F 5 EDC 1 D 2 B</mods:affiliation>
<mods:nameIdentifier id="28C687D9493E8F76C5BECF65C5AE9422" type="email">acontreras@ib.unam.mx</mods:nameIdentifier>
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<mods:title id="03413A1C02A0B3BCC93259091CA12B4F">Revista Brasileira de Entomologia</mods:title>
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<mods:date id="E11DD74CC201F87694F280692171A8C8">2023</mods:date>
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<mods:title id="C7A5F3E2175F64FB513CF01FC3E5E6DD">e 20230044</mods:title>
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<mods:classification id="ED81ECDF37723225879D44B36A859A22">journal article</mods:classification>
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<treatment id="CE2CD627EC7AFFDBFFA8FBF8C59B1C43" LSID="urn:lsid:plazi:treatment:CE2CD627EC7AFFDBFFA8FBF8C59B1C43" httpUri="http://treatment.plazi.org/id/CE2CD627EC7AFFDBFFA8FBF8C59B1C43" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="0E9F34BAEC7AFFDBFFA8FBF8C1C9190A" box="[113,460,1025,1045]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="463A6731EC7AFFDBFFA8FBF8C1C9190A" blockId="8.[113,460,1025,1045]" box="[113,460,1025,1045]" pageId="8" pageNumber="9">
<heading id="1D72D05DEC7AFFDBFFA8FBF8C1C9190A" bold="true" box="[113,460,1025,1045]" fontSize="8" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="81851CB2EC7AFFDBFFA8FBF8C1C9190A" ID-CoL="3ZQZ4" authority="(Banks, 1905)" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[113,460,1025,1045]" class="Insecta" family="Myrmeleontidae" genus="Menkeleon" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bellulus">
<emphasis id="74F1BB23EC7AFFDBFFA8FBF8C1C9190A" bold="true" box="[113,460,1025,1045]" pageId="8" pageNumber="9">
Menkeleon bellulus (
<bibRefCitation id="22141AC0EC7AFFDBFE90FBFBC1C0190A" author="Banks, N." box="[329,453,1025,1044]" pageId="8" pageNumber="9" pagination="1 - 20" refId="ref11820" refString="Banks, N., 1905. Descriptions of new Nearctic neuropteroid insects. Trans. Am. Entomol. Soc. 32, 1 - 20." type="journal article" year="1905">Banks, 1905</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0E9F34BAEC7AFFDBFF4EFBE5C115192C" box="[151,272,1055,1074]" pageId="8" pageNumber="9" type="description">
<paragraph id="463A6731EC7AFFDBFF4EFBE5C115192C" blockId="8.[112,787,1055,1985]" box="[151,272,1055,1074]" pageId="8" pageNumber="9">
<figureCitation id="DEBE7BB4EC7AFFDBFF4EFBE5C0E7192C" box="[151,226,1055,1074]" captionStart="Figure 1" captionStartId="1.[83,138,1947,1964]" captionTargetBox="[169,1387,836,1931]" captionTargetId="figure-428@1.[169,1387,835,1931]" captionTargetPageId="1" captionText="Figure 1 Live specimens of species comprised in the former Gnopholeontini:A) Gnopholeon barberi (Currie, 1903), by BJ Stacey. B) Gnopholeon delicatulus (Currie, 1903), by CD Martinez. C) Menkeleon bellulus (Banks, 1905), by Sean Werle. D) Tyttholeon puerilis Adams, 1957, by BJ Stacey." figureDoi="http://doi.org/10.5281/zenodo.13197556" httpUri="https://zenodo.org/record/13197556/files/figure.png" pageId="8" pageNumber="9">Figs. 1C</figureCitation>
,
<figureCitation id="DEBE7BB4EC7AFFDBFF32FBE5C115192C" box="[235,272,1055,1074]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStartId-0="7.[83,136,1122,1139]" captionStartId-1="7.[83,136,1905,1922]" captionTargetBox-0="[378,1180,158,1094]" captionTargetBox-1="[231,1327,1216,1881]" captionTargetId-0="figure-141@7.[378,1180,151,1106]" captionTargetId-1="figure-13@7.[229,1327,1211,1889]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Figure 8 Menkeleon bellulus (Banks, 1905), female specimen. A) habitus, dorsal view. B) habitus, lateral view. C) head, thorax and legs, lateral view.D) head, frontal view.E) fore and hind wings. F) head and thorax, dorsal view." captionText-1="Figure 9 Menkeleon bellulus (Banks, 1905), male and female terminalia.A) male terminalia, ventral view.B) male terminalia, posterior view.C) male terminalia, lateral view.D) female terminalia, ventral view. E) female terminalia, lateral view. — Abbreviations: ag: anterior gonapophysis; ds: digging setae; et: ectoproct; etl: ectoproct postventral lobe; etm: ectoproct median process; etv: ectoproct ventral process; go: gonarcus; gp: gonapophyseal plate; lg: lateral gonapophysis; md: mediuncus; pap: paramere plates; pat: paramere tooth; pg: posterior gonapophysis; pgp: pregenital plate; stIX: sternite IX; tgIX: tergite IX." figureDoi-0="http://doi.org/10.5281/zenodo.13197572" figureDoi-1="http://doi.org/10.5281/zenodo.13197574" httpUri-0="https://zenodo.org/record/13197572/files/figure.png" httpUri-1="https://zenodo.org/record/13197574/files/figure.png" pageId="8" pageNumber="9">8–9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0E9F34BAEC7AFFDBFF4EFBC6C14018D3" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="463A6731EC7AFFDBFF4EFBC6C26919DB" blockId="8.[112,787,1055,1985]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="6695401FEC7AFFDBFF4EFBC6C26919DB" pageId="8" pageNumber="9">
<treatmentCitation id="C7244120EC7AFFDBFF4EFBC6C1E1194E" author="Banks, N." box="[151,484,1084,1104]" page="7" pageId="8" pageNumber="9" year="1905">
<taxonomicName id="81851CB2EC7AFFDBFF4EFBC6C1E1194E" ID-CoL="3XY4W" authority="Banks, 1905: 7" authorityName="Banks" authorityPageNumber="7" authorityYear="1905" box="[151,484,1084,1104]" class="Insecta" family="Myrmeleontidae" genus="Maracandula" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bellula">
Maracandula bellula
<bibRefCitation id="22141AC0EC7AFFDBFE85FBC6C1E1194E" author="Banks, N." box="[348,484,1084,1104]" pageId="8" pageNumber="9" pagination="1 - 20" refId="ref11820" refString="Banks, N., 1905. Descriptions of new Nearctic neuropteroid insects. Trans. Am. Entomol. Soc. 32, 1 - 20." type="journal article" year="1905">Banks, 1905: 7</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="993ED993EC7AFFDBFE35FBC7C24E194E" box="[492,587,1085,1104]" pageId="8" pageNumber="9" type="lectotype">Lectotype</typeStatus>
female, Three Rivers,
<collectingRegion id="8441A9D3EC7AFFDBFFA9FBA0C0D51973" box="[112,208,1114,1133]" country="United States of America" name="California" pageId="8" pageNumber="9">California</collectingRegion>
(MCZ), designated by
<treatmentCitation id="C7244120EC7AFFDBFE60FBA0C26A1973" author="Stange, L. A." box="[441,623,1114,1133]" page="155" pageId="8" pageNumber="9" year="1970">
<bibRefCitation id="22141AC0EC7AFFDBFE60FBA0C26A1973" author="Stange, L. A." box="[441,623,1114,1133]" pageId="8" pageNumber="9" pagination="1 - 192" refId="ref13410" refString="Stange, L. A., 1970. Revision of the ant-lion tribe Brachynemurini of North America (Neuroptera: myrmeleontidae). Univ. Calif. Publ. Entomol. 55 (6), 1 - 192." type="journal article" year="1970">Stange, 1970: 155</bibRefCitation>
</treatmentCitation>
]. – Baker, 1905: 90 [distribution]. –
<treatmentCitation id="C7244120EC7AFFDBFEEEFB8DC1D01994" author="Banks, N." box="[311,469,1143,1162]" page="32" pageId="8" pageNumber="9" year="1907">
<bibRefCitation id="22141AC0EC7AFFDBFEEEFB8DC1D01994" author="Banks, N." box="[311,469,1143,1162]" pageId="8" pageNumber="9" refId="ref11848" refString="Banks, N. 1907. Catalogue of the Neuropteroid Insects (Except Odonata) of the United States. Am. Entomol. Soc., Philadelphia, 53 pp." type="book" year="1907">Banks, 1907: 32</bibRefCitation>
</treatmentCitation>
[species list]. –
<treatmentCitation id="C7244120EC7AFFDBFDA3FB8DC3171994" author="Banks, N." box="[634,786,1143,1162]" page="59" pageId="8" pageNumber="9" year="1927">
<bibRefCitation id="22141AC0EC7AFFDBFDA3FB8DC3171994" author="Banks, N." box="[634,786,1143,1162]" pageId="8" pageNumber="9" pagination="1 - 84" refId="ref11958" refString="Banks, N., 1927. Revision of the Nearctic Myrmeleonidae [sic]. Bull. Mus. Comp. Zool. 68, 1 - 84." type="journal article" year="1927">Banks 1927: 59</bibRefCitation>
</treatmentCitation>
[redescription, taxonomy, distribution]. –
<treatmentCitation id="C7244120EC7AFFDBFE22FB6EC29919B6" author="Markl, W." box="[507,668,1172,1192]" page="224" pageId="8" pageNumber="9" year="1954">
<bibRefCitation id="22141AC0EC7AFFDBFE22FB6EC29919B6" author="Markl, W." box="[507,668,1172,1192]" pageId="8" pageNumber="9" pagination="178 - 263" refId="ref12660" refString="Markl, W., 1954. Vergleichend-morphologische Studien zur Systematik und Klassifikation der Myrmeleoniden (Insecta, Neuroptera). Verh. Natforsch. Ges. Basel 65, 178 - 263. [errata: 66, 140]" type="journal article" year="1954">Markl, 1954: 224</bibRefCitation>
</treatmentCitation>
[illustration of wings]. –
<treatmentCitation id="C7244120EC7AFFDBFF30FB48C18919DB" author="Adams, P. A." box="[233,396,1202,1221]" page="93" pageId="8" pageNumber="9" year="1956">
<bibRefCitation id="22141AC0EC7AFFDBFF30FB48C18919DB" author="Adams, P. A." box="[233,396,1202,1221]" pageId="8" pageNumber="9" pagination="82 - 108" refId="ref11738" refString="Adams, P. A., 1956. New ant-lions from the southwestern United States (Neuroptera: myrmeleontidae). Psyche (Stuttg.) 63, 82 - 108." type="journal article" year="1956">Adams, 1956: 93</bibRefCitation>
</treatmentCitation>
[genitalia illustration].
</treatmentCitationGroup>
</paragraph>
<paragraph id="463A6731EC7AFFDBFF4EFB2AC2381803" blockId="8.[112,787,1055,1985]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="6695401FEC7AFFDBFF4EFB2AC2381803" pageId="8" pageNumber="9">
=
<treatmentCitation id="C7244120EC7AFFDBFF72FB35C23E19FC" author="Banks, N." box="[171,571,1231,1251]" page="140" pageId="8" pageNumber="9" year="1942">
<taxonomicName id="81851CB2EC7AFFDBFF72FB35C23E19FC" ID-CoL="3XY4Z" authority="Banks, 1942: 140" authorityName="Banks" authorityPageNumber="140" authorityYear="1942" box="[171,571,1231,1251]" class="Insecta" family="Myrmeleontidae" genus="Maracandula" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="minima">
Maracandula minima
<bibRefCitation id="22141AC0EC7AFFDBFE55FB35C23E19FC" author="Banks, N." box="[396,571,1231,1251]" pageId="8" pageNumber="9" pagination="133 - 151" refId="ref12014" refString="Banks, N., 1942. Contributions toward a knowledge of the insect fauna of Lower California. No. 4. Neuroptera Myrmeleonidae. Proc. Calif. Acad. Sci. 24 (4), 133 - 151." type="journal article" year="1942">Banks, 1942: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="993ED993EC7AFFDBFD90FB35C2A219FC" box="[585,679,1231,1250]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
male, San Miguel, Lower
<collectingRegion id="8441A9D3EC7AFFDBFED0FB17C16E181E" box="[265,363,1261,1280]" country="United States of America" name="California" pageId="8" pageNumber="9">California</collectingRegion>
,
<date id="323B41F1EC7AFFDBFEAEFB17C1E6181E" box="[375,483,1261,1280]" pageId="8" pageNumber="9" value="1938-07-03">03.vii.1938</date>
, M &amp; R (CAS)]. –
<bibRefCitation id="22141AC0EC7AFFDBFD40FB17C317181E" author="Banks, N." box="[665,786,1261,1280]" pageId="8" pageNumber="9" pagination="133 - 151" refId="ref12014" refString="Banks, N., 1942. Contributions toward a knowledge of the insect fauna of Lower California. No. 4. Neuroptera Myrmeleonidae. Proc. Calif. Acad. Sci. 24 (4), 133 - 151." type="journal article" year="1942">Banks, 1942</bibRefCitation>
[illustrations]. –
<treatmentCitation id="C7244120EC7AFFDBFECBFAF0C1BB1803" author="Stange, L. A." box="[274,446,1290,1309]" page="155" pageId="8" pageNumber="9" year="1970">
<bibRefCitation id="22141AC0EC7AFFDBFECBFAF0C1BB1803" author="Stange, L. A." box="[274,446,1290,1309]" pageId="8" pageNumber="9" pagination="1 - 192" refId="ref13410" refString="Stange, L. A., 1970. Revision of the ant-lion tribe Brachynemurini of North America (Neuroptera: myrmeleontidae). Univ. Calif. Publ. Entomol. 55 (6), 1 - 192." type="journal article" year="1970">Stange, 1970: 155</bibRefCitation>
</treatmentCitation>
[synonymy].
</treatmentCitationGroup>
</paragraph>
<paragraph id="463A6731EC7AFFDBFF4EFAD2C14018D3" blockId="8.[112,787,1055,1985]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="6695401FEC7AFFDBFF4EFAD2C14018D3" pageId="8" pageNumber="9">
=
<taxonomicName id="81851CB2EC7AFFDBFF72FADDC2011825" ID-CoL="3ZQZ4" authority="(Banks, 1905)" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[171,516,1319,1339]" class="Insecta" family="Myrmeleontidae" genus="Menkeleon" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bellulus">
Menkeleon bellulus (
<bibRefCitation id="22141AC0EC7AFFDBFE5BFADDC1FF1824" author="Banks, N." box="[386,506,1319,1338]" pageId="8" pageNumber="9" pagination="1 - 20" refId="ref11820" refString="Banks, N., 1905. Descriptions of new Nearctic neuropteroid insects. Trans. Am. Entomol. Soc. 32, 1 - 20." type="journal article" year="1905">Banks, 1905</bibRefCitation>
)
</taxonomicName>
. –
<treatmentCitation id="C7244120EC7AFFDBFDFCFAD2C2D21824" author="Stange, L. A." box="[549,727,1319,1339]" page="155" pageId="8" pageNumber="9" year="1970">
<bibRefCitation id="22141AC0EC7AFFDBFDFCFAD2C2D21824" author="Stange, L. A." box="[549,727,1319,1339]" pageId="8" pageNumber="9" pagination="1 - 192" refId="ref13410" refString="Stange, L. A., 1970. Revision of the ant-lion tribe Brachynemurini of North America (Neuroptera: myrmeleontidae). Univ. Calif. Publ. Entomol. 55 (6), 1 - 192." type="journal article" year="1970">Stange, 1970: 155</bibRefCitation>
</treatmentCitation>
[new combination, redescription, taxonomy, distribution]. –
<treatmentCitation id="C7244120EC7AFFDBFDA5FABFC3161846" author="Penny, N. D." box="[636,787,1349,1368]" page="45" pageId="8" pageNumber="9" year="1977">
<bibRefCitation id="22141AC0EC7AFFDBFDA5FABFC3161846" author="Penny, N. D." box="[636,787,1349,1368]" pageId="8" pageNumber="9" pagination="1 - 61" refId="ref13263" refString="Penny, N. D., 1977. Lista de Megaloptera, Neuroptera e Raphidioptera do Mexico, America Central, ilhas Caraibas e America do Sul. Acta Amazon. 7, 1 - 61." type="journal article" year="1977">Penny, 1977: 45</bibRefCitation>
</treatmentCitation>
[distribution].–
<treatmentCitation id="C7244120EC7AFFDBFED1FA99C1C9186B" author="Penny, N. D. &amp; Adams, P. A. &amp; Stange, L. A." box="[264,460,1378,1398]" page="85" pageId="8" pageNumber="9" year="1997">
<bibRefCitation id="22141AC0EC7AFFDBFED1FA99C1C9186B" author="Penny, N. D. &amp; Adams, P. A. &amp; Stange, L. A." box="[264,460,1378,1398]" pageId="8" pageNumber="9" pagination="39 - 114" refId="ref13301" refString="Penny, N. D., Adams, P. A., Stange, L. A., 1997. Species catalog of the Neuroptera, Megaloptera, and Raphidioptera of America North of Mexico. Proc. Calif. Acad. Sci. 50 (4), 39 - 114." type="journal article" year="1997">Penny et al., 1997: 85</bibRefCitation>
</treatmentCitation>
[taxonomy, distribution].–
<treatmentCitation id="C7244120EC7AFFDBFD17FA98C0C6188D" author="Stange, L. A." page="92" pageId="8" pageNumber="9" year="1994">
<bibRefCitation id="22141AC0EC7AFFDBFD17FA98C0C6188D" author="Stange, L. A." pageId="8" pageNumber="9" pagination="67 - 119" refId="ref13563" refString="Stange, L. A., 1994. Reclassification of the New World antlion genera formerly included in the tribe Brachynemurini (Neuroptera: myrmeleontidae). Insecta Mundi 8, 67 - 119." type="journal article" year="1994">Stange, 1994: 92</bibRefCitation>
</treatmentCitation>
, 112, 117 [larvae description, illustration, larvae photograph]. –
<treatmentCitation id="C7244120EC7AFFDBFFA9FA67C11F18AE" author="Stange, L. A." box="[112,282,1437,1456]" page="251" pageId="8" pageNumber="9" year="2004">
<bibRefCitation id="22141AC0EC7AFFDBFFA9FA67C11F18AE" author="Stange, L. A." box="[112,282,1437,1456]" pageId="8" pageNumber="9" pagination="1 - 565" refId="ref13599" refString="Stange, L. A., 2004. A systematic catalog, bibliography and classification of the world antlions (Insecta: Neuroptera: Myrmeleontidae). Mem. Am. Entomol. Inst. 74 (4), 1 - 565." type="journal article" year="2004">Stange, 2004: 251</bibRefCitation>
</treatmentCitation>
[taxonomy,
<typeStatus id="993ED993EC7AFFDBFE54FA67C1C718AE" box="[397,450,1437,1456]" pageId="8" pageNumber="9">types</typeStatus>
, distribution]. –
<treatmentCitation id="C7244120EC7AFFDBFD82FA67C09218D3" author="Badano, D. &amp; Miller, R. &amp; Stange, L. A." page="936" pageId="8" pageNumber="9" year="2018">
<bibRefCitation id="22141AC0EC7AFFDBFD82FA67C09218D3" author="Badano, D. &amp; Miller, R. &amp; Stange, L. A." pageId="8" pageNumber="9" pagination="933 - 949" refId="ref11771" refString="Badano, D., Miller, R., Stange, L. A., 2018. Rediscovery and revision of the antlion genus Ripalda Navas within a phylogeny of Nemoleontini (Neuroptera, Myrmeleontidae). Invertebr. Syst. 32, 933 - 949." type="journal article" year="2018">Badano et al., 2018: 936</bibRefCitation>
</treatmentCitation>
, 937 [phylogeny].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0E9F34BAEC7AFFDBFF4EFA22C30518F5" box="[151,768,1496,1515]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="463A6731EC7AFFDBFF4EFA22C30518F5" blockId="8.[112,787,1055,1985]" box="[151,768,1496,1515]" pageId="8" pageNumber="9">
<emphasis id="74F1BB23EC7AFFDBFF4EFA22C11E18F5" bold="true" box="[151,283,1496,1515]" pageId="8" pageNumber="9">Distribution.</emphasis>
<collectingCountry id="3E9227A1EC7AFFDBFEF9FA22C13F18F5" box="[288,314,1496,1515]" name="United States of America" pageId="8" pageNumber="9">US</collectingCountry>
(AZ, CA, NM, TX, UT),
<collectingCountry id="3E9227A1EC7AFFDBFDCBFA22C25C18F5" box="[530,601,1496,1515]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
(BC, CHIH, SON).
</paragraph>
</subSubSection>
<subSubSection id="0E9F34BAEC7AFFDBFF4EFA0FC1981B85" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="463A6731EC7AFFDBFF4EFA0FC1981B85" blockId="8.[112,787,1055,1985]" pageId="8" pageNumber="9">
<emphasis id="74F1BB23EC7AFFDBFF4EFA0FC1051B16" bold="true" box="[151,256,1525,1544]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
Adults. Abdomen banded with pale and dark brown; wings in rest in a moth-like position; male gonarcus broad, not twisted laterally; paramere plates sclerotized ventrally; paramere teeth basally expanded, broad, slightly curved and pointing posterodorsally; female lateral gonapophysis with long digging setae; posterior gonapophysis cylindrical, with hairlike setae.
</paragraph>
</subSubSection>
<subSubSection id="0E9F34BAEC7AFFDBFF4EF95CC0F41A98" pageId="8" pageNumber="9" type="discussion">
<paragraph id="463A6731EC7AFFDBFF4EF95CC0F41A98" blockId="8.[112,787,1055,1985]" pageId="8" pageNumber="9">
<emphasis id="74F1BB23EC7AFFDBFF4EF95CC0FC1BA7" bold="true" box="[151,249,1702,1721]" pageId="8" pageNumber="9">Remarks.</emphasis>
This species was first recorded from
<collectingCountry id="3E9227A1EC7AFFDBFDB1F95FC2B41BA6" box="[616,689,1701,1720]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
(and
<collectingRegion id="8441A9D3EC7AFFDBFD31F95FC0D01BC8" country="Mexico" name="Baja California" pageId="8" pageNumber="9">Baja California</collectingRegion>
) by
<bibRefCitation id="22141AC0EC7AFFDBFEDDF939C18B1BC8" author="Stange, L. A." box="[260,398,1731,1750]" pageId="8" pageNumber="9" pagination="1 - 192" refId="ref13410" refString="Stange, L. A., 1970. Revision of the ant-lion tribe Brachynemurini of North America (Neuroptera: myrmeleontidae). Univ. Calif. Publ. Entomol. 55 (6), 1 - 192." type="journal article" year="1970">Stange (1970)</bibRefCitation>
. The general pale and dark coloration with a banded abdomen and setose body are very characteristic for this species. Additionally, when perched,
<taxonomicName id="81851CB2EC7AFFDBFDDEF904C2711A0F" baseAuthorityName="Banks" baseAuthorityYear="1905" box="[519,628,1790,1809]" class="Insecta" family="Myrmeleontidae" genus="Menkeleon" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bellulus">M. bellulus</taxonomicName>
individulas rest their wings in a characteristic moth-like position (
<figureCitation id="DEBE7BB4EC7AFFDBFD83F8E1C29F1A30" box="[602,666,1819,1838]" captionStart="Figure 1" captionStartId="1.[83,138,1947,1964]" captionTargetBox="[169,1387,836,1931]" captionTargetId="figure-428@1.[169,1387,835,1931]" captionTargetPageId="1" captionText="Figure 1 Live specimens of species comprised in the former Gnopholeontini:A) Gnopholeon barberi (Currie, 1903), by BJ Stacey. B) Gnopholeon delicatulus (Currie, 1903), by CD Martinez. C) Menkeleon bellulus (Banks, 1905), by Sean Werle. D) Tyttholeon puerilis Adams, 1957, by BJ Stacey." figureDoi="http://doi.org/10.5281/zenodo.13197556" httpUri="https://zenodo.org/record/13197556/files/figure.png" pageId="8" pageNumber="9">Fig. 1C</figureCitation>
), a behavior also seen in
<taxonomicName id="81851CB2EC7AFFDBFF21F8C2C16E1A55" authorityName="Banks" authorityYear="1920" box="[248,363,1848,1867]" class="Insecta" family="Myrmeleontidae" genus="Chaetoleon" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Chaetoleon</taxonomicName>
tripunctatus (
<bibRefCitation id="22141AC0EC7AFFDBFDDDF8C2C2871A52" author="Banks, N." box="[516,642,1848,1868]" pageId="8" pageNumber="9" pagination="58 - 60" refId="ref11933" refString="Banks, N., 1922. South America Glenurus and some other Myrmeleonidae. Can. Entomol. 54, 58 - 60." type="journal article" year="1922">Banks, 1922</bibRefCitation>
), C. variabilis
<bibRefCitation id="22141AC0EC7AFFDBFFA8F8ACC0EF1A77" author="Banks, N." box="[113,234,1878,1897]" pageId="8" pageNumber="9" pagination="133 - 151" refId="ref12014" refString="Banks, N., 1942. Contributions toward a knowledge of the insect fauna of Lower California. No. 4. Neuroptera Myrmeleonidae. Proc. Calif. Acad. Sci. 24 (4), 133 - 151." type="journal article" year="1942">Banks, 1942</bibRefCitation>
and in a few
<taxonomicName id="81851CB2EC7AFFDBFEABF8ACC2101A77" box="[370,533,1878,1897]" class="Insecta" family="Myrmeleontidae" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Brachynemurini">Brachynemurini</taxonomicName>
formerly included in the Lemolemini.
</paragraph>
</subSubSection>
<subSubSection id="0E9F34BAEC7AFFDBFF4EF86AC59B1C43" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="463A6731EC7AFFDBFF4EF86AC3171ADF" blockId="8.[112,787,1055,1985]" pageId="8" pageNumber="9">
<emphasis id="74F1BB23EC7AFFDBFF4EF86AC1581ABD" bold="true" box="[151,349,1936,1955]" pageId="8" pageNumber="9">Material examined.</emphasis>
(
<specimenCount id="5083ACB8EC7AFFDBFEB5F86BC1821AB8" box="[364,391,1937,1958]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
,
<specimenCount id="5083ACB8EC7AFFDBFE49F86BC1A81AB8" box="[400,429,1937,1958]" count="2" pageId="8" pageNumber="9" type="female">2♀</specimenCount>
)
<materialsCitation id="F6ED6D6CEC7AFFDBFE60F86BC30B1ADF" collectingDate="2022-07-01" country="Mexico" county="Ensenada" elevation="1205" latitude="31.109484" location="Hotel" longLatPrecision="1" longitude="-115.63665" municipality="Rancho Mike's Sky" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Baja California">
<collectingCountry id="3E9227A1EC7AFFDBFE60F86BC2041ABA" box="[441,513,1937,1956]" name="Mexico" pageId="8" pageNumber="9">México</collectingCountry>
–
<collectingRegion id="8441A9D3EC7AFFDBFDCEF86BC2AC1ABA" box="[535,681,1937,1956]" country="Mexico" name="Baja California" pageId="8" pageNumber="9">Baja California</collectingRegion>
:
<collectingCounty id="AF5B1FBDEC7AFFDBFD69F86BC30A1ABA" box="[688,783,1937,1956]" pageId="8" pageNumber="9">Ensenada</collectingCounty>
,
<collectingMunicipality id="A65EFD4BEC7AFFDBFFA8F854C11C1ADF" box="[113,281,1966,1985]" pageId="8" pageNumber="9">Rancho Mike’s Sky</collectingMunicipality>
,
<location id="435A31EAEC7AFFDBFEF9F854C1561ADF" LSID="urn:lsid:plazi:treatment:CE2CD627EC7AFFDBFFA8FBF8C59B1C43:435A31EAEC7AFFDBFEF9F854C1561ADF" box="[288,339,1966,1985]" country="Mexico" county="Ensenada" latitude="31.109484" longLatPrecision="1" longitude="-115.63665" municipality="Rancho Mike's Sky" name="Hotel" pageId="8" pageNumber="9" stateProvince="Baja California">Hotel</location>
,
<quantity id="817DCAD4EC7AFFDBFE83F854C1A41ADF" box="[346,417,1966,1985]" metricMagnitude="3" metricUnit="m" metricValue="1.205" pageId="8" pageNumber="9" unit="m" value="1205.0">
<elevation id="CDA88002EC7AFFDBFE83F854C1A41ADF" box="[346,417,1966,1985]" metricMagnitude="3" metricUnit="m" metricValue="1.205" pageId="8" pageNumber="9" unit="m" value="1205.0">1205 m</elevation>
</quantity>
;
<geoCoordinate id="23B101F6EC7AFFDBFE71F854C21E1ADF" box="[424,539,1966,1985]" degrees="31" direction="north" minutes="06.569" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="31.109484">31°06.569’N</geoCoordinate>
,
<geoCoordinate id="23B101F6EC7AFFDBFDF8F854C2A51ADF" box="[545,672,1966,1985]" degrees="115" direction="west" minutes="38.199" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="-115.63665">115º38.199’W</geoCoordinate>
;
<date id="323B41F1EC7AFFDBFD7EF854C30B1ADF" box="[679,782,1966,1985]" pageId="8" pageNumber="9" value="2022-07-01">
<collectingDate id="227FB819EC7AFFDBFD7EF854C30B1ADF" box="[679,782,1966,1985]" pageId="8" pageNumber="9" value="2022-07-01">01.vii.2022</collectingDate>
</date>
</materialsCitation>
;
</paragraph>
<paragraph id="463A6731EC7AFFDBFCE7FF60C59B1C43" blockId="8.[830,1506,154,351]" pageId="8" pageNumber="9">
<materialsCitation id="F6ED6D6CEC7AFFDBFCE7FF60C4B81C1B" collectingDate="2022-06-29" collectionCode="CNIN" collectorName="A. Contreras &amp; A. Gomez &amp; Y. Marquez &amp; A. Ramirez &amp; M. Ramirez &amp; N Rancho El Piedra" elevation="1857" latitude="31.8972" location="Sierra Juarez" longLatPrecision="1" longitude="-115.9656" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1">
<specimenCount id="5083ACB8EC7AFFDBFCE7FF60C3641DB1" box="[830,865,154,175]" count="1" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
; colecta manual;
<collectorName id="EB7002E7EC7AFFDBFBC9FF60C48C1DB3" box="[1040,1161,154,173]" pageId="8" pageNumber="9">A. Contreras</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFB4BFF60C4F41DB3" box="[1170,1265,154,173]" pageId="8" pageNumber="9">A. Gómez</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFB23FF60C56C1DB3" box="[1274,1385,154,173]" pageId="8" pageNumber="9">Y. Marquez</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFAA8FF60C5D91DB3" box="[1393,1500,154,173]" pageId="8" pageNumber="9">A. Ramírez</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFC99FF42C3B01DD4" box="[832,949,183,203]" pageId="8" pageNumber="9">M. Ramírez</collectorName>
;
<collectionCode id="2094FFF4EC7AFFDBFC18FF4DC3F11DD4" box="[961,1012,183,202]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="8" pageNumber="9">CNIN</collectionCode>
NEU:MYR:4008 •
<location id="435A31EAEC7AFFDBFB74FF4DC5291DD5" LSID="urn:lsid:plazi:treatment:CE2CD627EC7AFFDBFFA8FBF8C59B1C43:435A31EAEC7AFFDBFB74FF4DC5291DD5" box="[1197,1324,183,203]" latitude="31.8972" longLatPrecision="1" longitude="-115.9656" name="Sierra Juarez" pageId="8" pageNumber="9">Sierra Juarez</location>
, 1.1 km
<collectorName id="EB7002E7EC7AFFDBFA58FF42C39D1DF9" pageId="8" pageNumber="9">N Rancho El Piedra</collectorName>
, ca.
<locationDeviation id="57887E13EC7AFFDBFC18FF2EC5331DF9" box="[961,1334,212,232]" location="Parque Nacional Constitucion" pageId="8" pageNumber="9">
<quantity id="817DCAD4EC7AFFDBFC18FF2EC3FB1DF9" box="[961,1022,212,231]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="km" value="15.0">15 km</quantity>
S Parque Nacional Constitución
</locationDeviation>
de
<quantity id="817DCAD4EC7AFFDBFA83FF2EC5D91DF6" box="[1370,1500,212,232]" metricMagnitude="3" metricUnit="m" metricValue="1.8571512" pageId="8" pageNumber="9" unit="m" value="1857.1512">
<elevation id="CDA88002EC7AFFDBFA83FF2EC5D91DF6" box="[1370,1500,212,232]" metricMagnitude="3" metricUnit="m" metricValue="1.8571512" pageId="8" pageNumber="9" unit="m" value="1857.1512">1857, 1512 m</elevation>
</quantity>
;
<geoCoordinate id="23B101F6EC7AFFDBFC99FF08C3BC1C1B" box="[832,953,242,261]" degrees="31" direction="north" minutes="53.832" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="31.8972">31°53.832’N</geoCoordinate>
,
<geoCoordinate id="23B101F6EC7AFFDBFC1AFF08C44C1C1B" box="[963,1097,242,261]" degrees="115" direction="west" minutes="57.936" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="-115.9656">115º57.936’W</geoCoordinate>
;
<date id="323B41F1EC7AFFDBFB8AFF08C4B81C1B" box="[1107,1213,242,261]" pageId="8" pageNumber="9" value="2022-06-29">
<collectingDate id="227FB819EC7AFFDBFB8AFF08C4B81C1B" box="[1107,1213,242,261]" pageId="8" pageNumber="9" value="2022-06-29">29.vi.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="F6ED6D6CEC7AFFDBFB11FF08C39A1C43" collectionCode="UV, CNIN" collectorName="A. Contreras &amp; A. Gomez &amp; Y. Marquez &amp; A. Ramirez &amp; M. Ramirez &amp; La Aduana" location="La Aduana" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Sonora">
<specimenCount id="5083ACB8EC7AFFDBFB11FF08C4E61C19" box="[1224,1251,242,263]" count="1" pageId="8" pageNumber="9" type="male">1♂</specimenCount>
; trampa de luz
<collectionCode id="2094FFF4EC7AFFDBFA58FF08C59A1C1B" box="[1409,1439,242,261]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="8" pageNumber="9">UV</collectionCode>
, vapor de mercurio, aditivos metálicos;
<collectorName id="EB7002E7EC7AFFDBFB5AFEF5C4F91C3C" box="[1155,1276,271,290]" pageId="8" pageNumber="9">A. Contreras</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFADFFEF5C5601C3C" box="[1286,1381,271,290]" pageId="8" pageNumber="9">A. Gómez</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFAB6FEEAC5DB1C3D" box="[1391,1502,272,291]" pageId="8" pageNumber="9">Y. Marquez</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFC99FED7C3AA1C5E" box="[832,943,301,320]" pageId="8" pageNumber="9">A. Ramírez</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFC63FED7C4341C5E" box="[954,1073,301,320]" pageId="8" pageNumber="9">M. Ramírez</collectorName>
;
<collectionCode id="2094FFF4EC7AFFDBFBE4FED7C4751C5E" box="[1085,1136,301,320]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="8" pageNumber="9">CNIN</collectionCode>
NEU:MYR:4091.
<collectingRegion id="8441A9D3EC7AFFDBFAC4FED7C5631C5E" box="[1309,1382,301,320]" country="Mexico" name="Sonora" pageId="8" pageNumber="9">Sonora</collectingRegion>
:
<collectorName id="EB7002E7EC7AFFDBFAA8FED7C5DB1C5E" box="[1393,1502,301,320]" pageId="8" pageNumber="9">
<location id="435A31EAEC7AFFDBFAA8FED7C5DB1C5E" LSID="urn:lsid:plazi:treatment:CE2CD627EC7AFFDBFFA8FBF8C59B1C43:435A31EAEC7AFFDBFAA8FED7C5DB1C5E" box="[1393,1502,301,320]" name="La Aduana" pageId="8" pageNumber="9" stateProvince="Sonora">La Aduana</location>
</collectorName>
;
<date id="323B41F1EC7AFFDBFCE6FEB0C39A1C43" box="[831,927,330,349]" pageId="8" pageNumber="9" value="1062-05-22">22.v.1062</date>
</materialsCitation>
;
<materialsCitation id="F6ED6D6CEC7AFFDBFC71FEB0C59C1C43" box="[936,1433,330,351]" collectionCode="CNIN" collectorName="F. D. Parker &amp; L. A. Stange" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Sonora">
<specimenCount id="5083ACB8EC7AFFDBFC71FEB0C3CF1C41" box="[936,970,330,351]" count="1" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
;
<collectorName id="EB7002E7EC7AFFDBFC0EFEB0C4461C43" box="[983,1091,330,349]" pageId="8" pageNumber="9">F. D. Parker</collectorName>
,
<collectorName id="EB7002E7EC7AFFDBFB95FEB0C4BA1C43" box="[1100,1215,330,349]" pageId="8" pageNumber="9">L. A. Stange</collectorName>
;
<collectionCode id="2094FFF4EC7AFFDBFB11FEB0C4FE1C43" box="[1224,1275,330,349]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="8" pageNumber="9">CNIN</collectionCode>
NEU:MYR:2637
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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