258 lines
34 KiB
XML
258 lines
34 KiB
XML
<document id="5C0C462AD2916B7493C955EFCA81EDAD" ID-DOI="10.15298/rusentj.28.1.02" ID-ISSN="0132-8069" ID-Zenodo-Dep="10919914" checkinTime="1712198186010" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2019" docId="CD4787B3FFCD416CFC09FE9BFDE1FA95" docLanguage="en" docName="RussEntomolJ.28.1.5-16.pdf" docOrigin="Russian Entomological Journal 28 (1)" docSource="http://dx.doi.org/10.15298/rusentj.28.1.02" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docTitle="Eustra parallela Fedorenko 2019, sp.n." docType="treatment" docVersion="1" lastPageNumber="15" masterDocId="317EFFCBFFC44166FF93FFA7FFB3FFF4" masterDocTitle="New or little-known species of the genus Eustra Schmidt-Göbel, 1846 (Coleoptera: Carabidae: Ozaenini)" masterLastPageNumber="16" masterPageNumber="5" pageNumber="14" updateTime="1712198501952" updateUser="ExternalLinkService">
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<mods:title id="AC3ACCF3484EFF086BBD16AF4F5D3424">New or little-known species of the genus Eustra Schmidt-Göbel, 1846 (Coleoptera: Carabidae: Ozaenini)</mods:title>
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<mods:name id="2084A5E65E7EA3EC8568C491DDA6D9B3" type="personal">
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<mods:namePart id="88C8739482ABB773E8BE17CE57DA8169">Fedorenko, D. N.</mods:namePart>
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<mods:title id="9DB622E055F6CEEDF1C07282A580A90E">Russian Entomological Journal</mods:title>
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<mods:part id="210D1A74EFD0E004F55A18B7714D50BB">
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<mods:date id="EEF8288A1052219DEFC820E24CBC12F2">2019</mods:date>
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<mods:number id="DD27C413957DEB652B2C1D4E5ACF1A67">2019-09-30</mods:number>
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<mods:number id="DC7D91BA95B0346E8629DA70E0C3A4FA">28</mods:number>
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<mods:url id="34A52312A51AF8B31FB7748841D1CEE4">http://dx.doi.org/10.15298/rusentj.28.1.02</mods:url>
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<mods:identifier id="C4A2029B6A02D4A77FF93E7CA91E8388" type="DOI">10.15298/rusentj.28.1.02</mods:identifier>
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<mods:identifier id="19B6A16BB4520BB436CFD2723FF3DDEB" type="ISSN">0132-8069</mods:identifier>
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<treatment id="CD4787B3FFCD416CFC09FE9BFDE1FA95" LSID="urn:lsid:plazi:treatment:CD4787B3FFCD416CFC09FE9BFDE1FA95" httpUri="http://treatment.plazi.org/id/CD4787B3FFCD416CFC09FE9BFDE1FA95" lastPageId="10" lastPageNumber="15" pageId="9" pageNumber="14">
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<subSubSection id="0DF4652EFFCD416FFC09FE9BFABBFEA7" box="[922,1288,315,339]" pageId="9" pageNumber="14" type="nomenclature">
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<paragraph id="455136A5FFCD416FFC09FE9BFABBFEA7" blockId="9.[812,1399,315,982]" box="[922,1288,315,339]" pageId="9" pageNumber="14">
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<heading id="1E1981C9FFCD416FFC09FE9BFABBFEA7" box="[922,1288,315,339]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="14" reason="8">
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<taxonomicName id="82EE4D26FFCD416FFC09FE9BFB7BFEA7" ID-CoL="8MDLW" authority="Fedorenko" authorityName="Fedorenko" authorityYear="2019" box="[922,1224,315,339]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="parallela" status="sp. nov.">
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<emphasis id="779AEAB7FFCD416FFC09FE9BFBF8FEA7" box="[922,1099,315,339]" italics="true" pageId="9" pageNumber="14">Eustra parallela</emphasis>
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Fedorenko
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</taxonomicName>
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,
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<emphasis id="779AEAB7FFCD416FFB40FE9CFABBFEA7" bold="true" box="[1235,1288,315,339]" pageId="9" pageNumber="14">
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<taxonomicNameLabel id="6CA957CCFFCD416FFB40FE9CFABBFEA7" box="[1235,1288,315,339]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0DF4652EFFCD416FFC54FEFEFB6FFE84" box="[967,1244,344,369]" pageId="9" pageNumber="14" type="description">
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<paragraph id="455136A5FFCD416FFC54FEFEFB6FFE84" blockId="9.[812,1399,315,982]" box="[967,1244,344,369]" pageId="9" pageNumber="14">
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<figureCitation id="DDD52A20FFCD416FFC54FEFEFBBAFE85" box="[967,1033,345,369]" captionStart="Figs 1–6" captionStartId="2.[226,263,1969,1988]" captionTargetBox="[200,1389,657,1951]" captionTargetId="figure-306@2.[188,1398,652,1954]" captionTargetPageId="2" captionText="Figs 1–6. Eustra spp., dorsal habitus: 1 — E. parallela sp.n.; 2 — E. cariniceps sp.n.; 3 — E. posteroguttata sp.n.; 4 — E. nageli subglabra ssp.n.; 5 — E. dalatensis sp.n.; 6 — E. csikii. Рис. 1–6. Eustra spp., габитус дорЗально: 1 — E. parallela sp.n.; 2 — E. cariniceps sp.n.; 3 — E. posteroguttata sp.n.; 4 — E. nageli subglabra ssp.n.; 5 — E. dalatensis sp.n.; 6 — E. csikii." figureDoi="http://doi.org/10.5281/zenodo.10919916" httpUri="https://zenodo.org/record/10919916/files/figure.png" pageId="9" pageNumber="14">Figs 1</figureCitation>
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,
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<figureCitation id="DDD52A20FFCD416FFB8BFEFEFBEEFE85" box="[1048,1117,345,369]" captionStart="Figs 7–18" captionStartId="5.[226,262,1969,1988]" captionTargetBox="[204,1393,463,1941]" captionTargetId="figure-121@5.[188,1398,450,1949]" captionTargetPageId="5" captionText="Figs 7–18. Median lobe of aedeagus: 7–9 — Eustra matanga matanga; 10–12 — E. posteroguttata sp.n.; 13–15 — E. csikii; 16–18 — E. parallela sp.n.; 7, 10, 13, 16 — right aspect; 8, 17 — ventral aspect; 9, 12, 15, 18 — left aspect; 11, 14 — dorsal aspect;. Scale bars: 0.5 mm. Рис. 7–18. СреднЯЯ долЯ Эдеагуса: 7–9 — Eustra matanga matanga; 10–12 — E. posteroguttata sp.n.; 13–15 — E. csikii; 16–18 — E. parallela sp.n.; 7, 10, 13, 16 — справа; 8, 17 — сниЗу; 9, 12, 15, 18 — слева; 11, 14 — сверху. Масштаб: 0.5 мм." figureDoi="http://doi.org/10.5281/zenodo.10919920" httpUri="https://zenodo.org/record/10919920/files/figure.png" pageId="9" pageNumber="14">16–18</figureCitation>
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,
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<figureCitation id="DDD52A20FFCD416FFBFAFEFEFB35FE85" box="[1129,1158,345,369]" captionStart="Figs 26–32" captionStartId="8.[226,263,1917,1936]" captionTargetBox="[192,1395,999,1881]" captionTargetId="figure-10@8.[189,1399,998,1876]" captionTargetPageId="8" captionText="Figs 26–32. Median lobe of aedeagus in plane of apex (~left ventrobasal aspect): 26 — Eustra nageli subglabra ssp.n.; 27 — E. posteroguttata sp.n.; 28 — E. cariniceps sp.n.; 29 — E. dalatensis sp.n.; 30 — E. csikii; 31 — E. parallela; 32 — E. matanga matanga. Scale bar 0.5 mm. Рис. 26–32. СреднЯЯ долЯ Эдеагуса в плоскости вершины (~вентробаЗально слева): 26 — Eustra nageli subglabra ssp.n.; 27 — E. posteroguttata sp.n.; 28 — E. cariniceps sp.n.; 29 — E. dalatensis sp.n.; 30 — E. csikii; 31 — E. parallela; 32 — E. matanga matanga. Масштаб 0.5 мм." figureDoi="http://doi.org/10.5281/zenodo.10919924" httpUri="https://zenodo.org/record/10919924/files/figure.png" pageId="9" pageNumber="14">31</figureCitation>
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,
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<figureCitation id="DDD52A20FFCD416FFB01FEFFFB6BFE84" box="[1170,1240,344,368]" captionStart="Figs 33–46" captionStartId="9.[226,263,1918,1937]" captionTargetBox="[197,1392,1006,1883]" captionTargetId="figure-565@9.[188,1398,996,1901]" captionTargetPageId="9" captionText="Figs 33–46. Parameres: 33–34 — Eustra cariniceps sp.n.; 35–36 — E. matanga matanga; 37–38 — E. csikii; 39–40 — E. posteroguttata sp.n.; 41–42 — E. dalatensis sp.n.; 43–44 — E. nageli subglabra ssp.n.; 45–46 — E. parallela sp.n.; 33, 35, 37, 39, 41, 43, 45 — right paramere, left aspect; 34, 36, 38, 40, 42, 44, 46 — left paramere, right aspect. Scale bar 0.5 mm. Рис. 33–46. Парамеры: 33–34 — Eustra cariniceps sp.n.; 35–36 — E. matanga matanga; 37–38 — E. csikii; 39–40 — E. posteroguttata sp.n.; 41–42 — E. dalatensis sp.n.; 43–44 — E. nageli subglabra ssp.n.; 45–46 — E. parallela sp.n.; 33, 35, 37, 39, 41, 43, 45 — праваЯ парамера, вид слева; 34, 36, 38, 40, 42, 44, 46 — леваЯ парамера, вид справа. Масштаб 0.5 мм." figureDoi="http://doi.org/10.5281/zenodo.10919928" httpUri="https://zenodo.org/record/10919928/files/figure.png" pageId="9" pageNumber="14">45–46</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="0DF4652EFFCD416FFCC1FED9FBD4FE35" pageId="9" pageNumber="14" type="materials_examined">
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<paragraph id="455136A5FFCD416FFCC1FED9FBD4FE35" blockId="9.[812,1399,315,982]" pageId="9" pageNumber="14">
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<materialsCitation id="F5863CF8FFCD416FFCC1FED9FBD4FE35" collectorName="Mt. Kinabalu N. P. & Kurbatov & Zimina'" country="Malaysia" elevation="1700" location="Mt. Kinabalu" pageId="9" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
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MATERIAL.
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<typeStatus id="9A558807FFCD416FFC5CFED9FBACFE65" box="[975,1055,382,401]" pageId="9" pageNumber="14" type="holotype">Holotype</typeStatus>
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<specimenCount id="53E8FD2CFFCD416FFBB4FEDCFB88FE65" box="[1063,1083,379,401]" count="1" pageId="9" pageNumber="14" type="male">♂</specimenCount>
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(ZMMU, formerly kept at MPSU) with label: ‘E
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<collectingCountry id="3DF97635FFCD416FFC22FE31FBB1FE5D" box="[945,1026,406,425]" name="Malaysia" pageId="9" pageNumber="14">Malaysia</collectingCountry>
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,
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<collectingRegion id="872AF847FFCD416FFB9DFE31FBF6FE5D" box="[1038,1093,406,425]" country="Malaysia" name="Sabah" pageId="9" pageNumber="14">Sabah</collectingRegion>
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,
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<collectorName id="E81B5373FFCD416FFBC2FE31FAB3FE5D" box="[1105,1280,406,425]" pageId="9" pageNumber="14">
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<location id="4031607EFFCD416FFBC2FE31FB74FE5D" LSID="urn:lsid:plazi:treatment:CD4787B3FFCD416CFC09FE9BFDE1FA95:4031607EFFCD416FFBC2FE31FB74FE5D" box="[1105,1223,406,425]" country="Malaysia" name="Mt. Kinabalu" pageId="9" pageNumber="14" stateProvince="Sabah">Mt. Kinabalu</location>
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N. P.
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</collectorName>
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,
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<quantity id="82169B40FFCD416FFA9FFE31FAF9FE5D" box="[1292,1354,406,425]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="14" unit="m" value="1700.0">
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<elevation id="CEC3D196FFCD416FFA9FFE31FAF9FE5D" box="[1292,1354,406,425]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="14" unit="m" value="1700.0">1700m</elevation>
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</quantity>
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||
, 16– 30.07.[20]02,
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<collectorName id="E81B5373FFCD416FFC3AFE09FC49FE35" box="[937,1018,430,449]" pageId="9" pageNumber="14">Kurbatov</collectorName>
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&
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<collectorName id="E81B5373FFCD416FFB88FE09FBD0FE35" box="[1051,1123,430,449]" pageId="9" pageNumber="14">Zimina’</collectorName>
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.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="0DF4652EFFCD416CFCC1FE61FE12FBC8" lastPageId="10" lastPageNumber="15" pageId="9" pageNumber="14" type="description">
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<paragraph id="455136A5FFCD416FFCC1FE61FB74FCC2" blockId="9.[812,1399,315,982]" pageId="9" pageNumber="14">
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REDESCRIPTION.
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<collectionCode id="23FFAE60FFCD416FFB8AFE61FB8BFE2F" box="[1049,1080,454,475]" pageId="9" pageNumber="14">BL</collectionCode>
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<quantity id="82169B40FFCD416FFBAEFE61FB3AFE2F" box="[1085,1161,454,475]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="14" unit="mm" value="3.4">3.4 mm</quantity>
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, body (
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<figureCitation id="DDD52A20FFCD416FFB5DFE62FAB5FE28" box="[1230,1286,453,476]" captionStart="Figs 1–6" captionStartId="2.[226,263,1969,1988]" captionTargetBox="[200,1389,657,1951]" captionTargetId="figure-306@2.[188,1398,652,1954]" captionTargetPageId="2" captionText="Figs 1–6. Eustra spp., dorsal habitus: 1 — E. parallela sp.n.; 2 — E. cariniceps sp.n.; 3 — E. posteroguttata sp.n.; 4 — E. nageli subglabra ssp.n.; 5 — E. dalatensis sp.n.; 6 — E. csikii. Рис. 1–6. Eustra spp., габитус дорЗально: 1 — E. parallela sp.n.; 2 — E. cariniceps sp.n.; 3 — E. posteroguttata sp.n.; 4 — E. nageli subglabra ssp.n.; 5 — E. dalatensis sp.n.; 6 — E. csikii." figureDoi="http://doi.org/10.5281/zenodo.10919916" httpUri="https://zenodo.org/record/10919916/files/figure.png" pageId="9" pageNumber="14">Fig. 1</figureCitation>
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) shiny pale brown, elytra each with a cloudy brown spot extending from 2/3 to 3/4 elytra; the spots widely separated from both each other and lateral margin. Dorsal microsculpture distinct on labrum, imperceptible on clypeus, obliterate behind, including pronotum and elytra; elytral aciculate microsculpture outside and including greater part of glabrous submarginal area.Head moderately densely pubescent only laterally, widely glabrous medially, pronotum glabrous except very few setae in apical third; elytra very sparsely pubescent on disc, more densely along base and behind humeri; pilosity moderately long, discal setae half as long as distance between neighbouring two in a longitudinal row or barely longer; setigerous punctures indistinctly tuberculate on disc.
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</paragraph>
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<paragraph id="455136A5FFCD416CFCC1FC9CFEEAFE98" blockId="9.[812,1399,315,982]" lastBlockId="10.[189,776,315,1377]" lastPageId="10" lastPageNumber="15" pageId="9" pageNumber="14">Head with neck constriction shallow laterally and nearly imperceptible medially, vertex almost flat. Supra-ocular carina conspicuous above eye, obliterate behind, leaving genae even and straight. Frontoclypeal suture very gently concave. Labrum apically truncate, with 6 close setigerous tubercles. Antennomeres 5–10 increasingly transverse, about as long as wide, with only 9 and 10 distinctly transverse. Mandibular scrobe bisetose.</paragraph>
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<caption id="1191662DFFCD416FFF71F8D9FB7EF7F9" ID-DOI="http://doi.org/10.5281/zenodo.10919928" ID-Zenodo-Dep="10919928" httpUri="https://zenodo.org/record/10919928/files/figure.png" pageId="9" pageNumber="14" startId="9.[226,263,1918,1937]" targetBox="[197,1392,1006,1883]" targetPageId="9" targetType="figure">
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<paragraph id="455136A5FFCD416FFF71F8D9FBA0F835" blockId="9.[189,1399,1917,2061]" pageId="9" pageNumber="14">
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Figs 33–46. Parameres: 33–34 —
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<taxonomicName id="82EE4D26FFCD416FFD96F8D9FD2EF864" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[517,669,1918,1936]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="cariniceps" status="sp. nov.">
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<emphasis id="779AEAB7FFCD416FFD96F8D9FD2EF864" box="[517,669,1918,1936]" italics="true" pageId="9" pageNumber="14">Eustra cariniceps</emphasis>
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</taxonomicName>
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<emphasis id="779AEAB7FFCD416FFD32F8D9FD7FF865" bold="true" box="[673,716,1918,1937]" pageId="9" pageNumber="14">
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<taxonomicNameLabel id="6CA957CCFFCD416FFD32F8D9FD7FF865" box="[673,716,1918,1937]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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; 35–36 —
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<taxonomicName id="82EE4D26FFCD416FFCBFF8D9FC6DF864" authorityName="Andrewes" authorityYear="1919" box="[812,990,1918,1936]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="matanga" subSpecies="matanga">
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<emphasis id="779AEAB7FFCD416FFCBFF8D9FC6DF864" box="[812,990,1918,1936]" italics="true" pageId="9" pageNumber="14">E. matanga matanga</emphasis>
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</taxonomicName>
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; 37–38 —
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<taxonomicName id="82EE4D26FFCD416FFBAEF8D9FB31F865" authorityName="Jedlicka" authorityYear="1968" box="[1085,1154,1918,1937]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="csikii">
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<emphasis id="779AEAB7FFCD416FFBAEF8D9FB31F865" box="[1085,1154,1918,1937]" italics="true" pageId="9" pageNumber="14">E. csikii</emphasis>
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</taxonomicName>
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; 39–40 —
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<taxonomicName id="82EE4D26FFCD416FFB72F8D9FAC4F864" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[1249,1399,1918,1936]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="posteroguttata" status="sp.n.">
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<emphasis id="779AEAB7FFCD416FFB72F8D9FAC4F864" box="[1249,1399,1918,1936]" italics="true" pageId="9" pageNumber="14">E. posteroguttata</emphasis>
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</taxonomicName>
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<emphasis id="779AEAB7FFCD416FFF2EF831FF5BF85D" bold="true" box="[189,232,1942,1961]" pageId="9" pageNumber="14">
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||
<taxonomicNameLabel id="6CA957CCFFCD416FFF2EF831FF5BF85D" box="[189,232,1942,1961]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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; 41–42 —
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<taxonomicName id="82EE4D26FFCD416FFEDEF831FE0CF85D" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[333,447,1942,1961]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="dalatensis" status="sp.n.">
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<emphasis id="779AEAB7FFCD416FFEDEF831FE0CF85D" box="[333,447,1942,1961]" italics="true" pageId="9" pageNumber="14">E. dalatensis</emphasis>
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</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFE56F831FE43F85D" bold="true" box="[453,496,1942,1961]" pageId="9" pageNumber="14">
|
||
<taxonomicNameLabel id="6CA957CCFFCD416FFE56F831FE43F85D" box="[453,496,1942,1961]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 43–44 —
|
||
<taxonomicName id="82EE4D26FFCD416FFDC5F831FCB0F85D" authorityName="Fedorenko" authorityYear="2019" box="[598,771,1942,1961]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="nageli" subSpecies="subglabra">
|
||
<emphasis id="779AEAB7FFCD416FFDC5F831FCB0F85D" box="[598,771,1942,1961]" italics="true" pageId="9" pageNumber="14">E. nageli subglabra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFC99F831FC8EF85D" bold="true" box="[778,829,1942,1961]" pageId="9" pageNumber="14">ssp.n.</emphasis>
|
||
; 45–46 —
|
||
<taxonomicName id="82EE4D26FFCD416FFC30F831FBB8F85D" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[931,1035,1942,1961]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="parallela" status="sp.n.">
|
||
<emphasis id="779AEAB7FFCD416FFC30F831FBB8F85D" box="[931,1035,1942,1961]" italics="true" pageId="9" pageNumber="14">E. parallela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFB81F831FB8EF85D" bold="true" box="[1042,1085,1942,1961]" pageId="9" pageNumber="14">
|
||
<taxonomicNameLabel id="6CA957CCFFCD416FFB81F831FB8EF85D" box="[1042,1085,1942,1961]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 33, 35, 37, 39, 41, 43, 45 — right paramere, left aspect; 34, 36, 38, 40, 42, 44, 46 — left paramere, right aspect. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
<paragraph id="455136A5FFCD416FFF71F86DFB7EF7F9" blockId="9.[189,1399,1917,2061]" pageId="9" pageNumber="14">
|
||
Рис. 33–46. Парамеры: 33–34 —
|
||
<taxonomicName id="82EE4D26FFCD416FFDB8F86DFD79F828" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[555,714,1994,2012]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="cariniceps" status="sp. nov.">
|
||
<emphasis id="779AEAB7FFCD416FFDB8F86DFD79F828" box="[555,714,1994,2012]" italics="true" pageId="9" pageNumber="14">Eustra cariniceps</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFD46F86EFCB3F828" bold="true" box="[725,768,1993,2012]" pageId="9" pageNumber="14">
|
||
<taxonomicNameLabel id="6CA957CCFFCD416FFD46F86EFCB3F828" box="[725,768,1993,2012]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 35–36 —
|
||
<taxonomicName id="82EE4D26FFCD416FFCE0F86DFB80F828" authorityName="Andrewes" authorityYear="1919" box="[883,1075,1994,2012]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="matanga" subSpecies="matanga">
|
||
<emphasis id="779AEAB7FFCD416FFCE0F86DFB80F828" box="[883,1075,1994,2012]" italics="true" pageId="9" pageNumber="14">E. matanga matanga</emphasis>
|
||
</taxonomicName>
|
||
; 37–38 —
|
||
<taxonomicName id="82EE4D26FFCD416FFB36F86DFB42F828" authorityName="Jedlicka" authorityYear="1968" box="[1189,1265,1993,2012]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="csikii">
|
||
<emphasis id="779AEAB7FFCD416FFB36F86DFB42F828" box="[1189,1265,1993,2012]" italics="true" pageId="9" pageNumber="14">E. csikii</emphasis>
|
||
</taxonomicName>
|
||
; 39–40 —
|
||
<taxonomicName id="82EE4D26FFCD416FFAF7F86DFE88F800" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="posteroguttata" status="sp.n.">
|
||
<emphasis id="779AEAB7FFCD416FFAF7F86DFE88F800" italics="true" pageId="9" pageNumber="14">E. posteroguttata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFED2F846FEDFF800" bold="true" box="[321,364,2017,2036]" pageId="9" pageNumber="14">
|
||
<taxonomicNameLabel id="6CA957CCFFCD416FFED2F846FEDFF800" box="[321,364,2017,2036]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 41–42 —
|
||
<taxonomicName id="82EE4D26FFCD416FFE40F845FDF7F800" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[467,580,2017,2036]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="dalatensis" status="sp.n.">
|
||
<emphasis id="779AEAB7FFCD416FFE40F845FDF7F800" box="[467,580,2017,2036]" italics="true" pageId="9" pageNumber="14">E. dalatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFDD8F846FDC5F800" bold="true" box="[587,630,2017,2036]" pageId="9" pageNumber="14">
|
||
<taxonomicNameLabel id="6CA957CCFFCD416FFDD8F846FDC5F800" box="[587,630,2017,2036]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 43–44 —
|
||
<taxonomicName id="82EE4D26FFCD416FFD4FF845FC39F800" authorityName="Fedorenko" authorityYear="2019" box="[732,906,2017,2036]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="nageli" subSpecies="subglabra">
|
||
<emphasis id="779AEAB7FFCD416FFD4FF845FC39F800" box="[732,906,2017,2036]" italics="true" pageId="9" pageNumber="14">E. nageli subglabra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFC02F846FC76F800" bold="true" box="[913,965,2017,2036]" pageId="9" pageNumber="14">ssp.n.</emphasis>
|
||
; 45–46 —
|
||
<taxonomicName id="82EE4D26FFCD416FFBB8F845FB27F800" authority="Fedorenko, 2019" authorityName="Fedorenko" authorityYear="2019" box="[1067,1172,2017,2036]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="14" phylum="Arthropoda" rank="species" species="parallela" status="sp.n.">
|
||
<emphasis id="779AEAB7FFCD416FFBB8F845FB27F800" box="[1067,1172,2017,2036]" italics="true" pageId="9" pageNumber="14">E. parallela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="779AEAB7FFCD416FFB08F846FB75F800" bold="true" box="[1179,1222,2017,2036]" pageId="9" pageNumber="14">
|
||
<taxonomicNameLabel id="6CA957CCFFCD416FFB08F846FB75F800" box="[1179,1222,2017,2036]" pageId="9" pageNumber="14" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 33, 35, 37, 39, 41, 43, 45 — праваЯ парамера, вид слева; 34, 36, 38, 40, 42, 44, 46 — леваЯ парамера, вид справа. Масштаб 0.5 мм.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="455136A5FFCE416CFF71FED6FE27FD58" blockId="10.[189,776,315,1377]" pageId="10" pageNumber="15">
|
||
Pronotum cordate,
|
||
<collectionCode id="23FFAE60FFCE416CFE0DFED6FE71FE72" box="[414,450,369,390]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="15">PW</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFCE416CFE5AFED6FE56FE72" box="[457,485,369,390]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="15" type="Museum">PL</collectionCode>
|
||
1.59,
|
||
<collectionCode id="23FFAE60FFCE416CFDB7FED6FDFBFE72" box="[548,584,369,390]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="15">PW</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFCE416CFDDCFED6FDC4FE72" box="[591,631,369,390]" pageId="10" pageNumber="15">HW</collectionCode>
|
||
1.12,
|
||
<collectionCode id="23FFAE60FFCE416CFD25FED6FD50FE72" box="[694,739,369,390]" pageId="10" pageNumber="15">PLw</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFCE416CFD79FED6FCB5FE72" box="[746,774,369,390]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="15" type="Museum">PL</collectionCode>
|
||
0.28, sides sinuate in front of right and sharp basal angles, subsinuate behind apical angles. Base barely narrower than apex,
|
||
<collectionCode id="23FFAE60FFCE416CFF66FE66FEA2FE22" box="[245,273,449,470]" pageId="10" pageNumber="15">PB</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFCE416CFE8AFE66FE8BFE22" box="[281,312,449,470]" pageId="10" pageNumber="15">PA</collectionCode>
|
||
0.92, its sides slightly oblique and sinuate; apex rather deeply concave. Basal foveae round, large and deep, each extended apicad into a deep submarginal lines almost reaching apex. Lateral margin widely explanate and slightly reflexed outside this line in apical half, more reflexed behind; lateral bead strongly reflexed; lateral edge with 8–9 minute denticles or tubercles. Basal bead fine to missing. Median line and anterior transverse impression deep, basal transverse impression very deep.
|
||
</paragraph>
|
||
<paragraph id="455136A5FFCE416CFF71FD16FEF4FCC5" blockId="10.[189,776,315,1377]" pageId="10" pageNumber="15">
|
||
Elytra long,
|
||
<collectionCode id="23FFAE60FFCE416CFECFFD16FECAFD32" box="[348,377,689,710]" pageId="10" pageNumber="15">EL</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFCE416CFE12FD16FE15FD32" box="[385,422,689,710]" pageId="10" pageNumber="15">EW</collectionCode>
|
||
1.49,
|
||
<collectionCode id="23FFAE60FFCE416CFE77FD16FDBAFD32" box="[484,521,689,710]" pageId="10" pageNumber="15">EW</collectionCode>
|
||
/
|
||
<collectionCode id="23FFAE60FFCE416CFD83FD16FD87FD32" box="[528,564,689,710]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="15">PW</collectionCode>
|
||
1.61, nearly parallel-sided in middle third; base slightly oblique towards peduncle, humeri rounded apically. Lateral edge setulose yet smooth. Submarginal glabrous area rather large. USS consisting of 12+5+5 setae.
|
||
</paragraph>
|
||
<paragraph id="455136A5FFCE416CFF71FC91FEDEFC43" blockId="10.[189,776,315,1377]" pageId="10" pageNumber="15">Legs. Mesotarsomere 1 distinctly shorter than mesotarsomeres 2–4 combined and slightly shorter than mesotarsomere 5 (TF2 0.92: 1.18); metatarsomere 1 distinctly longer than metatarsomeres 2–4 combined and mesotarsomere 5 (TF3 1.38: 1.13).</paragraph>
|
||
<paragraph id="455136A5FFCE416CFF71FC1CFE12FBC8" blockId="10.[189,776,315,1377]" pageId="10" pageNumber="15">
|
||
Aedeagus (
|
||
<figureCitation id="DDD52A20FFCE416CFEDCFC1CFE0CFC25" box="[335,447,955,977]" captionStart="Figs 7–18" captionStartId="5.[226,262,1969,1988]" captionTargetBox="[204,1393,463,1941]" captionTargetId="figure-121@5.[188,1398,450,1949]" captionTargetPageId="5" captionText="Figs 7–18. Median lobe of aedeagus: 7–9 — Eustra matanga matanga; 10–12 — E. posteroguttata sp.n.; 13–15 — E. csikii; 16–18 — E. parallela sp.n.; 7, 10, 13, 16 — right aspect; 8, 17 — ventral aspect; 9, 12, 15, 18 — left aspect; 11, 14 — dorsal aspect;. Scale bars: 0.5 mm. Рис. 7–18. СреднЯЯ долЯ Эдеагуса: 7–9 — Eustra matanga matanga; 10–12 — E. posteroguttata sp.n.; 13–15 — E. csikii; 16–18 — E. parallela sp.n.; 7, 10, 13, 16 — справа; 8, 17 — сниЗу; 9, 12, 15, 18 — слева; 11, 14 — сверху. Масштаб: 0.5 мм." figureDoi="http://doi.org/10.5281/zenodo.10919920" httpUri="https://zenodo.org/record/10919920/files/figure.png" pageId="10" pageNumber="15">Figs 16–18</figureCitation>
|
||
,
|
||
<figureCitation id="DDD52A20FFCE416CFE59FC1CFE57FC24" box="[458,484,955,976]" captionStart="Figs 26–32" captionStartId="8.[226,263,1917,1936]" captionTargetBox="[192,1395,999,1881]" captionTargetId="figure-10@8.[189,1399,998,1876]" captionTargetPageId="8" captionText="Figs 26–32. Median lobe of aedeagus in plane of apex (~left ventrobasal aspect): 26 — Eustra nageli subglabra ssp.n.; 27 — E. posteroguttata sp.n.; 28 — E. cariniceps sp.n.; 29 — E. dalatensis sp.n.; 30 — E. csikii; 31 — E. parallela; 32 — E. matanga matanga. Scale bar 0.5 mm. Рис. 26–32. СреднЯЯ долЯ Эдеагуса в плоскости вершины (~вентробаЗально слева): 26 — Eustra nageli subglabra ssp.n.; 27 — E. posteroguttata sp.n.; 28 — E. cariniceps sp.n.; 29 — E. dalatensis sp.n.; 30 — E. csikii; 31 — E. parallela; 32 — E. matanga matanga. Масштаб 0.5 мм." figureDoi="http://doi.org/10.5281/zenodo.10919924" httpUri="https://zenodo.org/record/10919924/files/figure.png" pageId="10" pageNumber="15">31</figureCitation>
|
||
,
|
||
<figureCitation id="DDD52A20FFCE416CFE7DFC1CFD9CFC25" box="[494,559,955,977]" captionStart="Figs 33–46" captionStartId="9.[226,263,1918,1937]" captionTargetBox="[197,1392,1006,1883]" captionTargetId="figure-565@9.[188,1398,996,1901]" captionTargetPageId="9" captionText="Figs 33–46. Parameres: 33–34 — Eustra cariniceps sp.n.; 35–36 — E. matanga matanga; 37–38 — E. csikii; 39–40 — E. posteroguttata sp.n.; 41–42 — E. dalatensis sp.n.; 43–44 — E. nageli subglabra ssp.n.; 45–46 — E. parallela sp.n.; 33, 35, 37, 39, 41, 43, 45 — right paramere, left aspect; 34, 36, 38, 40, 42, 44, 46 — left paramere, right aspect. Scale bar 0.5 mm. Рис. 33–46. Парамеры: 33–34 — Eustra cariniceps sp.n.; 35–36 — E. matanga matanga; 37–38 — E. csikii; 39–40 — E. posteroguttata sp.n.; 41–42 — E. dalatensis sp.n.; 43–44 — E. nageli subglabra ssp.n.; 45–46 — E. parallela sp.n.; 33, 35, 37, 39, 41, 43, 45 — праваЯ парамера, вид слева; 34, 36, 38, 40, 42, 44, 46 — леваЯ парамера, вид справа. Масштаб 0.5 мм." figureDoi="http://doi.org/10.5281/zenodo.10919928" httpUri="https://zenodo.org/record/10919928/files/figure.png" pageId="10" pageNumber="15">45–46</figureCitation>
|
||
): apex of median lobe toothed, right paramere slender, clavate, and longer than left one. Internal sac with several, large, spiniform, apical sclerites, including smaller dorsal ones in couple with large and arcuate ventral sclerite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4652EFFCE416CFF71FBE6FE76FAD8" pageId="10" pageNumber="15" type="diagnosis">
|
||
<paragraph id="455136A5FFCE416CFF71FBE6FE76FAD8" blockId="10.[189,776,315,1377]" pageId="10" pageNumber="15">
|
||
DIAGNOSIS. The new species is recognizable chiefly in the combination of almost flat vertex before very shallow neck constriction, fairly long elytra with very sparse elytral pubescence, combined with a distinctive aedeagus. This latter is different from that of
|
||
<emphasis id="779AEAB7FFCE416CFE43FB0BFDD2FB35" box="[464,609,1196,1217]" italics="true" pageId="10" pageNumber="15">
|
||
E. m.
|
||
<taxonomicName id="82EE4D26FFCE416CFD9FFB0BFDD2FB35" authorityName="Andrewes" authorityYear="1919" box="[524,609,1196,1217]" class="Insecta" family="Carabidae" genus="Eustra" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="15" phylum="Arthropoda" rank="species" species="matanga">matanga</taxonomicName>
|
||
</emphasis>
|
||
in the following points: right paramere very slender and subclavate, base of median lobe (that within its laterobasal extensions) as long as apical part (
|
||
<emphasis id="779AEAB7FFCE416CFEBDFB5BFEFAFAE5" box="[302,329,1276,1297]" italics="true" pageId="10" pageNumber="15">vs.</emphasis>
|
||
distinctly shorter), and internal sac with apical sclerites different in shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4652EFFCE416CFF71FA96FDE1FA95" pageId="10" pageNumber="15" type="distribution">
|
||
<paragraph id="455136A5FFCE416CFF71FA96FD41FAB3" blockId="10.[189,776,315,1377]" box="[226,754,1329,1351]" pageId="10" pageNumber="15">
|
||
DISTRIBUTION. Known from the
|
||
<typeStatus id="9A558807FFCE416CFDD3FA96FDD9FAB2" box="[576,618,1329,1350]" pageId="10" pageNumber="15">type</typeStatus>
|
||
locality only.
|
||
</paragraph>
|
||
<paragraph id="455136A5FFCE416CFF71FAECFDE1FA95" blockId="10.[189,776,315,1377]" box="[226,594,1355,1377]" pageId="10" pageNumber="15">HABITATS AND HABITS. No data.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |