293 lines
37 KiB
XML
293 lines
37 KiB
XML
<document id="ACA6CE88CAD53B42DF955273511E18D6" ID-CLB-Dataset="7835" ID-DOI="10.11646/zootaxa.4938.1.1" ID-GBIF-Dataset="045b347a-4a5b-474f-b69f-1cd3fedf7678" ID-ISSN="1175-5326" ID-Zenodo-Dep="4561258" ID-ZooBank="074E95CD-E7C0-4098-93F0-70DE03CAD0D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614236395824" checkinUser="plazi" docAuthor="Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De" docDate="2021" docId="CD475E66EC115A6A6BBF1906FBCDFB1C" docLanguage="en" docName="zootaxa.4938.1.1.pdf" docOrigin="Zootaxa 4938 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys isotarsis Cherman 2021, new species" docType="treatment" docUuid="0D51DD77-5048-4285-B0F6-A0785885A6F0" docUuidSource="ZooBank" docVersion="7" lastPageNumber="23" masterDocId="317E261EEC055A7C6B281C59FFAAFFBD" masterDocTitle="Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="21" updateTime="1698931070950" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2AF139A3BA083BE58ECE6EC7CCF08757">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="8C2DF464C7CA3EB6BF73DB1F773CC702">Cherman, Mariana Alejandra</mods:namePart>
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<mods:affiliation id="EE96EE170EF91A9F04B16A186A465038">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:namePart id="7809A6E898D59E6D90EA89B8B144F146">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="3E43A13D66D8597AB4EE9A778ECED286">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & basilio. dsb @ gmail. com; https: // orcid. org / 0000 - 0003 - 3556 - 5880</mods:affiliation>
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<mods:nameIdentifier id="503D73B9BA92DA18000501C2FA45E999" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3648197E258D5B93471E77D48EF49E33">Mise, Kleber Makoto</mods:namePart>
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<mods:affiliation id="509D0156A888A0F961337235E295E514">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & klebermise @ yahoo. com. br; https: // orcid. org / 0000 - 0003 - 4391 - 9589</mods:affiliation>
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<mods:nameIdentifier id="C45475B5ABFB30F7B4935792504A489D" type="email">klebermise@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="C4311B9B8BA465F5AFB97E01BDA7232E">Frisch, Johannes</mods:namePart>
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<mods:affiliation id="92A29C6B186B82CEF71C1CA9BA24A4E9">Museum für Naturkunde Berlin, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:namePart id="ECFAD286F8A5BD35BA40DAB12FA87C29">Almeida, Lúcia Massutti De</mods:namePart>
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<mods:affiliation id="73E1DDEF31DE43C31E59D41B74BFA8F8">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. & lalmeida @ ufpr. br; https: // orcid. org / 0000 - 0003 - 4277 - 711 X</mods:affiliation>
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<mods:nameIdentifier id="A5D15ABBAA1735F38F95BB0FC59BC14C" type="email">lalmeida@ufpr.br</mods:nameIdentifier>
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<treatment id="CD475E66EC115A6A6BBF1906FBCDFB1C" ID-DOI="http://doi.org/10.5281/zenodo.4561284" ID-GBIF-Taxon="180301836" ID-Zenodo-Dep="4561284" LSID="urn:lsid:zoobank.org:act:0D51DD77-5048-4285-B0F6-A0785885A6F0" httpUri="http://treatment.plazi.org/id/CD475E66EC115A6A6BBF1906FBCDFB1C" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<emphasis id="779A3362EC115A686BBF1906FD26FAC5" bold="true" box="[151,652,1374,1401]" pageId="20" pageNumber="21">
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<taxonomicName id="82EE94F3EC115A686BBF1906FE45FAC5" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[151,495,1374,1401]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="isotarsis" status="sp. nov.">
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<emphasis id="779A3362EC115A686BBF1906FEC5FAC4" bold="true" box="[151,367,1375,1401]" italics="true" pageId="20" pageNumber="21">Liogenys isotarsis</emphasis>
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Cherman
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,
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<taxonomicNameLabel id="6CA98E19EC115A686AD41906FD26FAC5" box="[508,652,1374,1401]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="0DF4BCFBEC115A686BBF19DAFCD3FA7C" pageId="20" pageNumber="21" type="description">
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<paragraph id="4551EF70EC115A686BBF19DAFEBFFA20" blockId="20.[151,889,1374,1473]" box="[151,277,1411,1437]" pageId="20" pageNumber="21">
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<figureCitation id="DDD5F3F5EC115A686BBF19DAFF4FFA20" box="[151,229,1411,1437]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="20" pageNumber="21">Figs. 9</figureCitation>
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;
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<figureCitation id="DDD5F3F5EC115A686BDA19DAFEBBFA20" box="[242,273,1411,1437]" captionStart="FIGURE 27" captionStartId="55.[151,250,1459,1483]" captionTargetBox="[171,1416,192,1422]" captionTargetId="figure-20@55.[151,1435,181,1434]" captionTargetPageId="55" captionText="FIGURE 27. Geographical distribution of the Liogenys new species. Shaped points: new species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561332" httpUri="https://zenodo.org/record/4561332/files/figure.png" pageId="20" pageNumber="21">27</figureCitation>
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.
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</paragraph>
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<paragraph id="4551EF70EC115A686BBF19FFFCD3FA7C" blockId="20.[151,889,1374,1473]" box="[151,889,1446,1473]" pageId="20" pageNumber="21">
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http://zoobank.org/
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<subSubSection id="0DF4BCFBEC115A686BBF19B6FDB7F9EC" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="4551EF70EC115A686BBF19B6FA9DF990" blockId="20.[151,1437,1518,2013]" pageId="20" pageNumber="21">
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<emphasis id="779A3362EC115A686BBF19B6FE94F9B5" bold="true" box="[151,318,1518,1545]" pageId="20" pageNumber="21">Type material</emphasis>
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:
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<materialsCitation id="F586E52DEC115A686A6619B6FA9DF990" ID-GBIF-Occurrence="3047558383" collectionCode="USNM" collectorName="M. S. Pennington" country="Trinidad and Tobago" location="Trinidad" pageId="20" pageNumber="21" specimenCount="2" typeStatus="holotype">
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<typeStatus id="9A5551D2EC115A686A6619B6FE1CF9B4" box="[334,438,1519,1545]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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labeled (
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<collectionCode id="23FF77B5EC115A68693519B6FDDFF9B4" box="[541,629,1519,1545]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
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): [white typeset] “
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<collectingCountry id="3DF9AFE0EC115A68681719B6FC0EF9B4" box="[831,932,1519,1545]" name="Trinidad and Tobago" pageId="20" pageNumber="21">Trinidad</collectingCountry>
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,
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<collectingCountry id="3DF9AFE0EC115A68688719B6FBABF9B4" box="[943,1025,1519,1545]" name="Bolivia" pageId="20" pageNumber="21">Bolivia</collectingCountry>
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/ 1922-23 /
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<collectorName id="E81B8AA6EC115A686FA219B6FAF9F9B4" box="[1162,1363,1519,1545]" pageId="20" pageNumber="21">M. S. Pennington</collectorName>
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”, [red, typeset and handwritten] “
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<taxonomicName id="82EE94F3EC115A686A941A4AFD77F990" authority="ISOTARSIS" authorityName="ISOTARSIS" box="[444,733,1555,1581]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">LIOGENYS ISOTARSIS</taxonomicName>
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/
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<typeStatus id="9A5551D2EC115A6869DC1A4AFC22F990" box="[756,904,1555,1581]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
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/ Cherman M. A.”, genitalia mounted.
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</materialsCitation>
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</paragraph>
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<paragraph id="4551EF70EC115A686BEF1A6EFDB7F9EC" blockId="20.[151,1437,1518,2013]" box="[199,541,1591,1617]" pageId="20" pageNumber="21">
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<typeStatus id="9A5551D2EC115A686BEF1A6EFE85F9EC" box="[199,303,1591,1617]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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deposited at USNM.
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</paragraph>
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</subSubSection>
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<subSubSection id="0DF4BCFBEC115A686BEF1A03FAD5F8B8" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="4551EF70EC115A686BEF1A03FAD5F8B8" blockId="20.[151,1437,1518,2013]" pageId="20" pageNumber="21">
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<emphasis id="779A3362EC115A686BEF1A03FE90F9C9" bold="true" box="[199,314,1626,1652]" pageId="20" pageNumber="21">Diagnosis</emphasis>
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. Body elongate; purplish brown, pronotum and elytra coarsely punctate (
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<figureCitation id="DDD5F3F5EC115A686F531A02FB7CF9C8" box="[1147,1238,1627,1653]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="20" pageNumber="21">Fig. 9A</figureCitation>
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); clypeal emargination angulate, teeth very short; outer sides of clypeal teeth subparallel; pronotal posterior corners rounded (
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<figureCitation id="DDD5F3F5EC115A686E471A26FF10F900" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="20" pageNumber="21">Fig. 9B</figureCitation>
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); in males metatibia not carinate along the inner margin, apical inner surface glabrous; tarsomeres almost equal in width, cylindrical (
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<figureCitation id="DDD5F3F5EC115A686AA41A9EFE65F95C" box="[396,463,1735,1761]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
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D–E); pygidium flat, glabrous (
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<figureCitation id="DDD5F3F5EC115A68681D1A9EFC24F95C" box="[821,910,1735,1761]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="20" pageNumber="21">Fig. 9C</figureCitation>
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); parameres narrowed at the midline; widened towards the apex, apex equal in width to the basal portion (
|
||
<figureCitation id="DDD5F3F5EC115A6868051AB2FC28F8B8" box="[813,898,1771,1797]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="20" pageNumber="21">Fig. 9F</figureCitation>
|
||
); parameres convex in lateral view (
|
||
<figureCitation id="DDD5F3F5EC115A686E301AB2FAD8F8B8" box="[1304,1394,1771,1797]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="20" pageNumber="21">Fig. 9G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC115A6A6BEF1B56FACCFD4C" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="materials_examined">
|
||
<paragraph id="4551EF70EC115A6A6BEF1B56FACCFD4C" blockId="20.[151,1437,1518,2013]" lastBlockId="22.[151,1437,151,1185]" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
|
||
<emphasis id="779A3362EC115A686BEF1B56FE9EF894" bold="true" box="[199,308,1807,1833]" pageId="20" pageNumber="21">
|
||
<typeStatus id="9A5551D2EC115A686BEF1B56FE9EF894" box="[199,308,1807,1833]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="779A3362EC115A686A681B56FED7F894" bold="true" box="[320,381,1807,1833]" pageId="20" pageNumber="21">Male</emphasis>
|
||
. Length:
|
||
<quantity id="82164295EC115A686ACE1B56FD97F894" box="[486,573,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="20" pageNumber="21" unit="mm" value="9.1">9.1 mm</quantity>
|
||
; width:
|
||
<quantity id="82164295EC115A6869BB1B57FD42F894" box="[659,744,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="20" pageNumber="21" unit="mm" value="5.1">5.1 mm</quantity>
|
||
. Purplish brown.
|
||
<emphasis id="779A3362EC115A6868821B56FC5AF894" box="[938,1008,1807,1833]" italics="true" pageId="20" pageNumber="21">Head:</emphasis>
|
||
distance between eyes more than three times the width of one eye; clypeal emargination deep, angulate and wide; teeth short, outer sides of anterior teeth subparallel; outer margin of anterior teeth noticeably shorter than the eye length; lateral margin slightly convex, not produced; distal maxillary palpomere, maximum width more than twice the apical width; fovea shallow, extending past the transverse midline; antenna with 10 antennomeres, club lighter in color and longer than the funicle.
|
||
<emphasis id="779A3362EC115A686E4F1BC6FF6FF861" italics="true" pageId="20" pageNumber="21">Thorax:</emphasis>
|
||
anterior margin of pronotum slightly produced medially; pronotal disc glabrous, punctures strongly coarse and very sparse; posterior corners rounded; hypomere with short bristles, mesepisternum and sides of metaventrite with sparse thick bristles; distance between mesocoxae and metacoxae up to twice the length of the metacoxa; scutellum ogival, with two longitudinal rows of punctures strongly coarse.
|
||
<emphasis id="779A3362EC135A6A685F1C86FC6CFF44" box="[887,966,223,249]" italics="true" pageId="22" pageNumber="23">Elytra:</emphasis>
|
||
shiny, glabrous, unicolored with the pronotum, coarsely punctate; elytral suture unicolored with the elytron and weakly elevated; ridges I, II, and IV more defined than ridge III.
|
||
<emphasis id="779A3362EC135A6A6ABA1D7EFE7AFEFD" box="[402,464,295,320]" italics="true" pageId="22" pageNumber="23">Legs:</emphasis>
|
||
three protibial teeth, the middle and apical equal in size; the three teeth equally spaced; mesofemural surface almost glabrous, setae on margins; mesotibia cylindrical in cross section, surface coarsely sculptured; two transverse carinae, the apical carina complete; metacoxa with sparse bristles, more abundant near the insertion of the metafemur; basal apophysis of metacoxa produced; metatibia not carinate along the inner margin, apical inner surface glabrous; two metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; protarsomeres and mesotarsomeres I to IV almost equal in width to the metatarsomeres I to IV (
|
||
<figureCitation id="DDD5F3F5EC135A6A680B1DA6FCD3FDA4" box="[803,889,511,537]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="22" pageNumber="23">Fig. 9E</figureCitation>
|
||
); metatarsomere I equal in size to metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and equal in width to the inferior, distance between teeth slightly longer than the inferior tooth.
|
||
<emphasis id="779A3362EC135A6A69151E1EFD1AFDDC" box="[573,688,583,609]" italics="true" pageId="22" pageNumber="23">Abdomen:</emphasis>
|
||
ventrites with scattered short bristles; propygidium slightly visible, glabrous; pygidium flat, subtrapezoidal, wide; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc glabrous; pygidial apex rounded.
|
||
<emphasis id="779A3362EC135A6A69D41ED6FCDCFD15" box="[764,886,655,680]" italics="true" pageId="22" pageNumber="23">Parameres</emphasis>
|
||
: narrowed at the midline; basal region wider than the parameres combined at the transverse midline, parameral split at 2/3; inner margins slightly convergent, after the midline widened towards the apex, apex equal in width to the basal portion; parameres convex in lateral view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1191BFF8EC105A696BBF1BCEFF74F845" ID-DOI="http://doi.org/10.5281/zenodo.4561286" ID-Zenodo-Dep="4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="21">
|
||
<paragraph id="4551EF70EC105A696BBF1BCEFF74F845" blockId="21.[151,1436,1943,2040]" pageId="21" pageNumber="22">
|
||
<emphasis id="779A3362EC105A696BBF1BCEFEBEF812" bold="true" box="[151,276,1943,1967]" pageId="21" pageNumber="22">FIGURE 9.</emphasis>
|
||
<taxonomicName id="82EE94F3EC105A696A331BC1FD96F80D" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[283,572,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="isotarsis" status="sp. nov.">
|
||
<emphasis id="779A3362EC105A696A331BC1FE7EF812" box="[283,468,1944,1967]" italics="true" pageId="21" pageNumber="22">Liogenys isotarsis</emphasis>
|
||
Cherman
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="779A3362EC105A69696E1BC1FD6EF812" bold="true" box="[582,708,1943,1968]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="6CA98E19EC105A69696E1BC1FD6EF812" box="[582,708,1943,1968]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0DF4BCFBEC135A6A6BEF1EA2FBC6FC84" pageId="22" pageNumber="23" type="etymology">
|
||
<paragraph id="4551EF70EC135A6A6BEF1EA2FBC6FC84" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="23">
|
||
<emphasis id="779A3362EC135A6A6BEF1EA2FEE2FCA8" bold="true" box="[199,328,763,789]" pageId="22" pageNumber="23">Etymology</emphasis>
|
||
. Adjective in the nominative singular. From Ancient Greek
|
||
<emphasis id="779A3362EC135A6A68DB1EA5FB88FCAA" box="[1011,1058,764,791]" italics="true" pageId="22" pageNumber="23">ἴΣΟΣ</emphasis>
|
||
(ísos, “equal”) +
|
||
<emphasis id="779A3362EC135A6A6FC51EA2FA9BFCA9" box="[1261,1329,763,788]" italics="true" pageId="22" pageNumber="23">tarsus</emphasis>
|
||
. In
|
||
<taxonomicName id="82EE94F3EC135A6A6E481EA2FE85FC85" authorityName="Cherman" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isotarsis">
|
||
<emphasis id="779A3362EC135A6A6E481EA2FE85FC85" italics="true" pageId="22" pageNumber="23">Liogenys isotarsis</emphasis>
|
||
</taxonomicName>
|
||
all the three pairs of tarsi have almost the same width, even being a male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC135A6A6BEF1F1AFD79FC3F" pageId="22" pageNumber="23" type="materials_examined">
|
||
<paragraph id="4551EF70EC135A6A6BEF1F1AFC11FCE0" blockId="22.[151,1437,151,1185]" box="[199,955,834,861]" pageId="22" pageNumber="23">
|
||
<emphasis id="779A3362EC135A6A6BEF1F1AFECCFCE1" bold="true" box="[199,358,834,861]" pageId="22" pageNumber="23">Type locality.</emphasis>
|
||
<materialsCitation id="F586E52DEC135A6A6A441F1AFC1DFCE0" ID-GBIF-Occurrence="3047558489" box="[364,951,834,861]" country="Bolivia" latitude="-14.832111" location="Trinidad" longLatPrecision="1" longitude="-64.90267" pageId="22" pageNumber="23" specimenCount="1">
|
||
<collectingCountry id="3DF9AFE0EC135A6A6A441F1AFE48FCE0" box="[364,482,835,861]" name="Bolivia" pageId="22" pageNumber="23">BOLIVIA</collectingCountry>
|
||
,
|
||
<location id="4031B9ABEC135A6A6AC51F1AFDE6FCE0" LSID="urn:lsid:plazi:treatment:CD475E66EC115A6A6BBF1906FBCDFB1C:4031B9ABEC135A6A6AC51F1AFDE6FCE0" box="[493,588,835,861]" country="Bolivia" latitude="-14.832111" longLatPrecision="1" longitude="-64.90267" name="Trinidad" pageId="22" pageNumber="23">Trinidad</location>
|
||
[
|
||
<geoCoordinate id="20DA89B7EC135A6A69721F1BFD50FCE0" box="[602,762,834,861]" degrees="14" direction="south" minutes="49" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="55.6" value="-14.832111">14°49’55.6’’S</geoCoordinate>
|
||
<geoCoordinate id="20DA89B7EC135A6A68291F1BFC1AFCE0" box="[769,944,834,861]" degrees="64" direction="west" minutes="54" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="09.6" value="-64.90267">64°54’09.6’’W</geoCoordinate>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4551EF70EC135A6A6BEF1F3FFD79FC3F" blockId="22.[151,1437,151,1185]" box="[199,723,870,898]" pageId="22" pageNumber="23">
|
||
<emphasis id="779A3362EC135A6A6BEF1F3FFE54FC3D" bold="true" box="[199,510,870,896]" pageId="22" pageNumber="23">Geographical distribution</emphasis>
|
||
.
|
||
<collectingCountry id="3DF9AFE0EC135A6A69241F3EFDD4FC3C" box="[524,638,871,897]" name="Bolivia" pageId="22" pageNumber="23">BOLIVIA</collectingCountry>
|
||
(
|
||
<collectingRegion id="872A2192EC135A6A69A41F3EFD6CFC3F" box="[652,710,871,898]" country="Bolivia" name="Beni" pageId="22" pageNumber="23">Beni</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DF4BCFBEC135A6A6BEF1FD2FBCDFB1C" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="4551EF70EC135A6A6BEF1FD2FAE9FBE4" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="23">
|
||
<emphasis id="779A3362EC135A6A6BEF1FD2FE9EFC18" bold="true" box="[199,308,907,933]" pageId="22" pageNumber="23">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="82EE94F3EC135A6A6A6B1FD2FDA4FC19" authorityName="Cherman" authorityYear="2021" box="[323,526,907,932]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isotarsis">
|
||
<emphasis id="779A3362EC135A6A6A6B1FD2FDA4FC19" box="[323,526,907,932]" italics="true" pageId="22" pageNumber="23">Liogenys isotarsis</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="82EE94F3EC135A6A69A61FD2FC34FC18" authority="Evans, 2003" authorityName="Evans" authorityYear="2003" box="[654,926,907,933]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="779A3362EC135A6A69A61FD2FCA1FC18" box="[654,779,907,933]" italics="true" pageId="22" pageNumber="23">L. unicolor</emphasis>
|
||
<bibRefCitation id="217F9281EC135A6A683A1FD2FC34FC18" author="Evans, A. V." box="[786,926,907,933]" pageId="22" pageNumber="23" pagination="1 - 458" refId="ref35927" refString="Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="217F9281EC135A6A68851FD2FB32FC18" author="Cherman, M. A. & Mise, K. M. & Moron, M. A. & Vaz-de-Mello, F. Z. & Almeida, L. M." box="[941,1176,907,933]" pageId="22" pageNumber="23" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. & Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
|
||
Cherman
|
||
<emphasis id="779A3362EC135A6A6F351FD2FBFCFC18" box="[1053,1110,907,933]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="217F9281EC135A6A6F8C1FD2FB40FC18" author="Cherman, M. A. & Basilio, D. S. & Mise, K. M. & Almeida, L. M." box="[1188,1258,907,933]" pageId="22" pageNumber="23" pagination="1 - 69" refId="ref35724" refString="Cherman, M. A., Basilio, D. S., Mise, K. M. & Almeida, L. M. (2019 a) Unraveling the puzzle of Liogenys biodiversity: fifteen new species, nomenclatural acts and new geographical records (Coleoptera: Scarabaeidae). Insect Systematics & Evolution, 51, 1 - 69. https: // doi. org / 10.1163 / 1876312 X- 00001040" type="journal article" year="2019">2019a</bibRefCitation>
|
||
) in the size and body color.
|
||
<taxonomicName id="82EE94F3EC135A6A6A341FF6FE4CFC75" authorityName="Cherman" authorityYear="2021" box="[284,486,943,968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isotarsis">
|
||
<emphasis id="779A3362EC135A6A6A341FF6FE4CFC75" box="[284,486,943,968]" italics="true" pageId="22" pageNumber="23">Liogenys isotarsis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="82EE94F3EC135A6A69521FF6FD5CFC74" authorityName="Evans" authorityYear="2003" box="[634,758,943,969]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="779A3362EC135A6A69521FF6FD5CFC74" box="[634,758,943,969]" italics="true" pageId="22" pageNumber="23">L. unicolor</emphasis>
|
||
</taxonomicName>
|
||
mainly in the outer sides of clypeal teeth subparallel (follow the lateral margin of clypeus); the pronotum wider; the punctures of scutellum and elytra coarser; the elytral ridges more defined; and in the pygidium flat (convex). In males, the protarsomeres are even more cylindrical and less enlarged than in
|
||
<taxonomicName id="82EE94F3EC135A6A6A1C1842FE1AFB88" authorityName="Evans" authorityYear="2003" box="[308,432,1051,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="779A3362EC135A6A6A1C1842FE1AFB88" box="[308,432,1051,1077]" italics="true" pageId="22" pageNumber="23">L. unicolor</emphasis>
|
||
</taxonomicName>
|
||
. The parameres resemble those of
|
||
<taxonomicName id="82EE94F3EC135A6A68181842FB25FB88" authority="Cherman, 2019" authorityName="Cherman" authorityYear="2019" box="[816,1167,1051,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="angustitarsis">
|
||
<emphasis id="779A3362EC135A6A68181842FC74FB89" box="[816,990,1051,1076]" italics="true" pageId="22" pageNumber="23">L. angustitarsis</emphasis>
|
||
Cherman, 2019
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="217F9281EC135A6A6FB41842FA3AFB88" author="Cherman, M. A. & Basilio, D. S. & Mise, K. M. & Almeida, L. M." box="[1180,1424,1051,1077]" pageId="22" pageNumber="23" pagination="1 - 69" refId="ref35724" refString="Cherman, M. A., Basilio, D. S., Mise, K. M. & Almeida, L. M. (2019 a) Unraveling the puzzle of Liogenys biodiversity: fifteen new species, nomenclatural acts and new geographical records (Coleoptera: Scarabaeidae). Insect Systematics & Evolution, 51, 1 - 69. https: // doi. org / 10.1163 / 1876312 X- 00001040" type="journal article" year="2019">
|
||
Cherman
|
||
<emphasis id="779A3362EC135A6A6E231842FAE8FB88" box="[1291,1346,1051,1077]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
|
||
2019a
|
||
</bibRefCitation>
|
||
), but they differ mainly in the lateral view, which are concave in the new species. Females remain unknown.
|
||
</paragraph>
|
||
<paragraph id="4551EF70EC135A6A6BBF183AFBCDFB1C" blockId="22.[151,1437,151,1185]" pageId="22" pageNumber="23">
|
||
<taxonomicName id="82EE94F3EC135A6A6BBF183AFEC8FBC1" authorityName="Cherman" authorityYear="2021" box="[151,354,1123,1148]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isotarsis">
|
||
<emphasis id="779A3362EC135A6A6BBF183AFEC8FBC1" box="[151,354,1123,1148]" italics="true" pageId="22" pageNumber="23">Liogenys isotarsis</emphasis>
|
||
</taxonomicName>
|
||
is possibly associated only to Bolivian Chiquitano Forests, and might occur in sympatry with
|
||
<taxonomicName id="82EE94F3EC135A6A6EAE183AFED6FB1C" authority="Frey, 1965" authorityName="Frey" authorityYear="1965" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="parallela">
|
||
<emphasis id="779A3362EC135A6A6EAE183AFF54FB1C" italics="true" pageId="22" pageNumber="23">L. parallela</emphasis>
|
||
<bibRefCitation id="217F9281EC135A6A6A2D18DEFED6FB1C" author="Frey, G." box="[261,380,1158,1185]" pageId="22" pageNumber="23" pagination="143 - 151" refId="ref36131" refString="Frey, G. (1965) Neue s ʾ damerikanische Melolonthidae (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 16, 143 - 151." type="journal article" year="1965">Frey, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82EE94F3EC135A6A6A9B18DEFD49FB1C" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[435,739,1159,1185]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mendozana">
|
||
<emphasis id="779A3362EC135A6A6A9B18DEFDFAFB1C" box="[435,592,1159,1185]" italics="true" pageId="22" pageNumber="23">L. mendozana</emphasis>
|
||
<bibRefCitation id="217F9281EC135A6A697F18DEFD49FB1C" author="Moser, J." box="[599,739,1159,1185]" pageId="22" pageNumber="23" pagination="95 - 167" refId="ref36796" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">
|
||
Moser,
|
||
<quantity id="82164295EC135A6A698318DEFD49FB1C" box="[683,739,1159,1185]" metricMagnitude="1" metricUnit="m" metricValue="4.87172" pageId="22" pageNumber="23" unit="in" value="1918.0">1918</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in their northernmost distribution.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |