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<document id="8F5831ACBF610BA2A81DD2CCEA831A8F" ID-CLB-Dataset="7835" ID-DOI="10.11646/zootaxa.4938.1.1" ID-GBIF-Dataset="045b347a-4a5b-474f-b69f-1cd3fedf7678" ID-ISSN="1175-5326" ID-Zenodo-Dep="4561258" ID-ZooBank="074E95CD-E7C0-4098-93F0-70DE03CAD0D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614236395824" checkinUser="plazi" docAuthor="Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes &amp; Almeida, Lúcia Massutti De" docDate="2021" docId="CD475E66EC075A7F6BBF194FFA36FAB5" docLanguage="en" docName="zootaxa.4938.1.1.pdf" docOrigin="Zootaxa 4938 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys Guerin-Meneville 1831" docType="key" docVersion="5" lastPageNumber="4" masterDocId="317E261EEC055A7C6B281C59FFAAFFBD" masterDocTitle="Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="3" updateTime="1698931070950" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EA4BA5E2C7C254FDCEA657F148560C7C">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="FB2576B25AE5455D02CD16C0AFF8FAC5">Cherman, Mariana Alejandra</mods:namePart>
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<mods:namePart id="B03A72755D1DF4E1701A5EE435EB9DDB">Basílio, Daniel Silva</mods:namePart>
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<mods:namePart id="EC23222DBF34AB202017614D8921780D">Mise, Kleber Makoto</mods:namePart>
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<mods:namePart id="BBD0A0756666984A3C5A945EA807548F">Frisch, Johannes</mods:namePart>
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<mods:namePart id="F1B36895195407990D718924DFD0F532">Almeida, Lúcia Massutti De</mods:namePart>
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Identification key to
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<emphasis id="779A3362EC075A7E6AB3194EFDA2FA8C" bold="true" box="[411,520,1303,1329]" italics="true" pageId="2" pageNumber="3">Liogenys</emphasis>
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species from the Chacoan province and its boundaries
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1. Antenna with 10 antennomeres; clypeal emargination wide or narrow; outer sides of anterior clypeal teeth parallel (
<figureCitation id="DDD5F3F5EC075A7E6E6B1905FA32FACC" box="[1347,1432,1372,1394]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="2" pageNumber="3">Figs. 2B</figureCitation>
,
<figureCitation id="DDD5F3F5EC075A7E6BCA1920FF54FA32" box="[226,254,1401,1423]" captionStart="FIGURE 3" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[215,1373,181,1884]" captionTargetId="figure-20@9.[215,1373,181,1884]" captionTargetPageId="9" captionText="FIGURE 3. Liogenys cribricollis Moser, 1921 male lectotype views: A) dorsal, B) head and pronotum, C) variation in shape of clypeus, D) labels, E) lateral, F) mesotibia and metatibia, G) mesotarsomere I, white arrow indicates ventral side modified, H) pygidium, I) female pygidium, J) apex of metatibia, white arrow indicates projection below the spurs, and the parameres in views: K) dorsal and L) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561268" httpUri="https://zenodo.org/record/4561268/files/figure.png" pageId="2" pageNumber="3">3B</figureCitation>
) or following the lateral margin of clypeus (
<figureCitation id="DDD5F3F5EC075A7E69801920FD56FA33" box="[680,764,1401,1423]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="2" pageNumber="3">Figs. 9B</figureCitation>
,
<figureCitation id="DDD5F3F5EC075A7E682F1920FC85FA32" box="[775,815,1401,1423]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="2" pageNumber="3">21B</figureCitation>
).................................................. 2
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- Antenna with 9 antennomeres; clypeal emargination narrow and semicircular (
<figureCitation id="DDD5F3F5EC075A7E68F419CFFB9EFA11" box="[988,1076,1430,1452]" captionStart="FIGURE 12" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@27.[203,1385,181,1920]" captionTargetPageId="27" captionText="FIGURE 12. Liogenys mendozana Moser, 1918 male lectotype views: A) dorsal, C) labels, E) head and pronotum, F) lateral, G) pygidium, H) female pygidium, and parameres in views: I) dorsal and J) lateral; L. mendozanus incisus Frey, 1969 female neotype views: B) dorsal, D) labels." figureDoi="http://doi.org/10.5281/zenodo.4561294" httpUri="https://zenodo.org/record/4561294/files/figure.png" pageId="2" pageNumber="3">Fig. 12E</figureCitation>
); outer sides of anterior clypeal teeth following the lateral margin of clypeus..........................................
<taxonomicName id="82EE94F3EC075A7E6F7C19EDFA36FA74" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[1108,1436,1460,1481]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mendozana">
<emphasis id="779A3362EC075A7E6F7C19EDFAB4FA74" box="[1108,1310,1460,1481]" italics="true" pageId="2" pageNumber="3">Liogenys mendozana</emphasis>
<bibRefCitation id="217F9281EC075A7E6E0C19EDFA36FA74" author="Moser, J." box="[1316,1436,1460,1481]" pageId="2" pageNumber="3" pagination="95 - 167" refId="ref36796" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">Moser, 1918</bibRefCitation>
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2. Clypeal teeth very short (
<figureCitation id="DDD5F3F5EC075A7E6AFF1988FD89FA5A" box="[471,547,1489,1511]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="2" pageNumber="3">Fig. 4B</figureCitation>
), outer sides of teeth not forming an angle with the lateral margin of clypeus; lateral margin convex or straight, not produced......................................................................... 3
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- Clypeal teeth long or short (
<figureCitation id="DDD5F3F5EC075A7E6AC61A55FDE8F99C" box="[494,578,1548,1570]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="2" pageNumber="3">Figs. 2B</figureCitation>
,
<figureCitation id="DDD5F3F5EC075A7E69641A55FDC0F99F" box="[588,618,1548,1570]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="2" pageNumber="3">8B</figureCitation>
), outer sides of teeth forming an angle with the lateral margin of clypeus, lateral margin convex and sometimes strongly produced (
<figureCitation id="DDD5F3F5EC075A7E698A1A70FD46F982" box="[674,748,1577,1599]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="2" pageNumber="3">Fig. 6B</figureCitation>
)....................................................... 8
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3. Pronotal disc densely punctate (
<figureCitation id="DDD5F3F5EC075A7E693C1A1FFDF7F9E1" box="[532,605,1606,1628]" captionStart="FIGURE 3" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[215,1373,181,1884]" captionTargetId="figure-20@9.[215,1373,181,1884]" captionTargetPageId="9" captionText="FIGURE 3. Liogenys cribricollis Moser, 1921 male lectotype views: A) dorsal, B) head and pronotum, C) variation in shape of clypeus, D) labels, E) lateral, F) mesotibia and metatibia, G) mesotarsomere I, white arrow indicates ventral side modified, H) pygidium, I) female pygidium, J) apex of metatibia, white arrow indicates projection below the spurs, and the parameres in views: K) dorsal and L) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561268" httpUri="https://zenodo.org/record/4561268/files/figure.png" pageId="2" pageNumber="3">Fig. 3B</figureCitation>
)................................................................... 4
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- Pronotal disc sparsely punctate (
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,
<figureCitation id="DDD5F3F5EC075A7E695F1A3DFD0AF9C7" box="[631,672,1636,1658]" captionStart="FIGURE 10" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@24.[203,1385,181,1920]" captionTargetPageId="24" captionText="FIGURE 10. Liogenys latitarsis Moser, 1918 male lectotype views: A) dorsal, B) labels, C) head and pronotum, D) lateral, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561288" httpUri="https://zenodo.org/record/4561288/files/figure.png" pageId="2" pageNumber="3">10C</figureCitation>
).............................................................. 6
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<paragraph id="4551EF70EC075A7E6BBF1AD8FA36F909" blockId="2.[151,1437,1372,2039]" pageId="2" pageNumber="3">
<keyLead id="FE1F4C45EC075A7E6BBF1AD8FA36F909" pageId="2" pageNumber="3">
4. Pronotum and elytra glabrous; punctures of pronotum very fine and dense (
<figureCitation id="DDD5F3F5EC075A7E68EE1AD8FBB9F92A" box="[966,1043,1665,1687]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="2" pageNumber="3">Fig. 4B</figureCitation>
); pygidium with dense scales (
<figureCitation id="DDD5F3F5EC075A7E6E6A1AD8FA3BF92A" box="[1346,1425,1665,1687]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="2" pageNumber="3">Fig. 4D</figureCitation>
), scales sparse only at the apex......................................................
<taxonomicName id="82EE94F3EC075A7E6FA01AC6FA36F909" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1160,1436,1694,1716]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="779A3362EC075A7E6FA01AC6FA9AF90E" box="[1160,1328,1694,1716]" italics="true" pageId="2" pageNumber="3">Liogenys densata</emphasis>
<bibRefCitation id="217F9281EC075A7E6E1E1AC6FA36F909" author="Frey, G." box="[1334,1436,1694,1716]" pageId="2" pageNumber="3" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
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</paragraph>
<paragraph id="4551EF70EC075A7E6BBF1AE5FA36F96C" blockId="2.[151,1437,1372,2039]" box="[151,1436,1723,1746]" pageId="2" pageNumber="3">
<keyLead id="FE1F4C45EC075A7E6BBF1AE5FA36F96C" box="[151,1436,1723,1746]" pageId="2" pageNumber="3">
- Pronotum and elytra setose; punctures of pronotum coarse and dense (
<figureCitation id="DDD5F3F5EC075A7E68AC1AE5FC65F96F" box="[900,975,1724,1746]" captionStart="FIGURE 3" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[215,1373,181,1884]" captionTargetId="figure-20@9.[215,1373,181,1884]" captionTargetPageId="9" captionText="FIGURE 3. Liogenys cribricollis Moser, 1921 male lectotype views: A) dorsal, B) head and pronotum, C) variation in shape of clypeus, D) labels, E) lateral, F) mesotibia and metatibia, G) mesotarsomere I, white arrow indicates ventral side modified, H) pygidium, I) female pygidium, J) apex of metatibia, white arrow indicates projection below the spurs, and the parameres in views: K) dorsal and L) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561268" httpUri="https://zenodo.org/record/4561268/files/figure.png" pageId="2" pageNumber="3">Fig. 3B</figureCitation>
); pygidium with sparse bristles and scales... 5
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<paragraph id="4551EF70EC075A7E6BBF1A80FA36F8B6" blockId="2.[151,1437,1372,2039]" pageId="2" pageNumber="3">
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5. Clypeal emargination semicircular, narrow, lateral margin of clypeus never produced; in males inner margin of metatibia concave subapically (
<figureCitation id="DDD5F3F5EC075A7E6AA61AAFFE48F8B1" box="[398,482,1782,1804]" captionStart="FIGURE 16" captionStartId="37.[151,250,1904,1928]" captionTargetBox="[216,1372,181,1880]" captionTargetId="figure-20@37.[216,1372,181,1880]" captionTargetPageId="37" captionText="FIGURE 16. Liogenys opacicollis Fairmaire, 1892 female lectotype views: A) dorsal, C) labels; L. densicollis Moser, 1921 male lectotype views: B) dorsal, D) labels, E) lateral, F) mesotibia and metatibia, white arrow indicates shape of metatibial margin, G) mesotarsomere I, arrow indicates modified inner margin and H) its variation, I) apex of metatibia, white arrow indicates projection below the spurs, J) pygidium, and parameres in views: K) dorsal and L) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561304" httpUri="https://zenodo.org/record/4561304/files/figure.png" pageId="2" pageNumber="3">Fig. 16F</figureCitation>
)................................................
<taxonomicName id="82EE94F3EC075A7E6F1F1AAEFA36F8B6" authority="Fairmaire, 1892" authorityName="Fairmaire" authorityYear="1892" box="[1079,1436,1782,1804]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="opacicollis">
<emphasis id="779A3362EC075A7E6F1F1AAEFB57F8B6" box="[1079,1277,1782,1804]" italics="true" pageId="2" pageNumber="3">Liogenys opacicollis</emphasis>
<bibRefCitation id="217F9281EC075A7E6E2B1AAFFA36F8B6" author="Fairmaire, M. L." box="[1283,1436,1782,1804]" pageId="2" pageNumber="3" pagination="242 - 253" refId="ref36073" refString="Fairmaire, M. L. (1892) Descriptions de quelques coleopteres Argentins. Annales de la Societe Entomologique de Belgique, 36, 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
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</paragraph>
<paragraph id="4551EF70EC075A7E6BBF1B4DFA36F8FB" blockId="2.[151,1437,1372,2039]" pageId="2" pageNumber="3">
<keyLead id="FE1F4C45EC075A7E6BBF1B4DFA36F8FB" pageId="2" pageNumber="3">
- Clypeal emargination rounded, wide, lateral margin of clypeus concave, produced in most of the cases, forming a blunt projection (
<figureCitation id="DDD5F3F5EC075A7E6A3C1B68FEF5F8FA" box="[276,351,1841,1863]" captionStart="FIGURE 3" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[215,1373,181,1884]" captionTargetId="figure-20@9.[215,1373,181,1884]" captionTargetPageId="9" captionText="FIGURE 3. Liogenys cribricollis Moser, 1921 male lectotype views: A) dorsal, B) head and pronotum, C) variation in shape of clypeus, D) labels, E) lateral, F) mesotibia and metatibia, G) mesotarsomere I, white arrow indicates ventral side modified, H) pygidium, I) female pygidium, J) apex of metatibia, white arrow indicates projection below the spurs, and the parameres in views: K) dorsal and L) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561268" httpUri="https://zenodo.org/record/4561268/files/figure.png" pageId="2" pageNumber="3">Fig. 3B</figureCitation>
); in males inner margin of metatibia convex subapically (
<figureCitation id="DDD5F3F5EC075A7E68A01B68FC65F8FA" box="[904,975,1841,1863]" captionStart="FIGURE 3" captionStartId="9.[151,250,1908,1932]" captionTargetBox="[215,1373,181,1884]" captionTargetId="figure-20@9.[215,1373,181,1884]" captionTargetPageId="9" captionText="FIGURE 3. Liogenys cribricollis Moser, 1921 male lectotype views: A) dorsal, B) head and pronotum, C) variation in shape of clypeus, D) labels, E) lateral, F) mesotibia and metatibia, G) mesotarsomere I, white arrow indicates ventral side modified, H) pygidium, I) female pygidium, J) apex of metatibia, white arrow indicates projection below the spurs, and the parameres in views: K) dorsal and L) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561268" httpUri="https://zenodo.org/record/4561268/files/figure.png" pageId="2" pageNumber="3">Fig. 3F</figureCitation>
)..........
<taxonomicName id="82EE94F3EC075A7E6F7F1B6BFA36F8FB" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[1111,1436,1841,1863]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cribricollis">
<emphasis id="779A3362EC075A7E6F7F1B6BFAB4F8FB" box="[1111,1310,1841,1863]" italics="true" pageId="2" pageNumber="3">Liogenys cribricollis</emphasis>
<bibRefCitation id="217F9281EC075A7E6E0C1B68FA36F8FB" author="Moser, J." box="[1316,1436,1841,1862]" pageId="2" pageNumber="3" pagination="133 - 182" refId="ref36844" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921</bibRefCitation>
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6. Lateral margin of clypeus concave, in males tarsomeres almost equal in width in all legs, cylindrical (
<figureCitation id="DDD5F3F5EC075A7E6F981B17FB53F8D9" box="[1200,1273,1870,1892]" captionStart="FIGURE 9" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@21.[203,1385,181,1920]" captionTargetPageId="21" captionText="FIGURE 9. Liogenys isotarsis Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) pygidium, D) lateral, E) metatibia and tarsi, black rows indicate not enlarged tarsi of all legs, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561286" httpUri="https://zenodo.org/record/4561286/files/figure.png" pageId="2" pageNumber="3">Fig. 9E</figureCitation>
)...............................................................................
<taxonomicName id="82EE94F3EC075A7E6F261B35FAB7F83F" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[1038,1309,1900,1922]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="isotarsis" status="sp. nov.">
<emphasis id="779A3362EC075A7E6F261B35FB16F83C" box="[1038,1212,1900,1921]" italics="true" pageId="2" pageNumber="3">Liogenys isotarsis</emphasis>
Cherman
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,
<emphasis id="779A3362EC075A7E6E0E1B35FA37F83C" bold="true" box="[1318,1437,1899,1921]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="6CA98E19EC075A7E6E0E1B35FA37F83C" box="[1318,1437,1899,1921]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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</paragraph>
<paragraph id="4551EF70EC075A7E6BBF1BD0FA36F823" blockId="2.[151,1437,1372,2039]" box="[151,1436,1929,1951]" pageId="2" pageNumber="3">
<keyLead id="FE1F4C45EC075A7E6BBF1BD0FA36F823" box="[151,1436,1929,1951]" pageId="2" pageNumber="3">
- Lateral margin of clypeus straight (
<figureCitation id="DDD5F3F5EC075A7E691C1BD0FD21F822" box="[564,651,1929,1951]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="2" pageNumber="3">Fig. 21B</figureCitation>
), in males protarsomeres and mesotarsomeres enlarged.................... 7
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<paragraph id="4551EF70EC075A7E6BBF1BFEFA37F801" blockId="2.[151,1437,1372,2039]" box="[151,1437,1958,1980]" pageId="2" pageNumber="3">
<keyLead id="FE1F4C45EC075A7E6BBF1BFEFA37F801" box="[151,1437,1958,1980]" pageId="2" pageNumber="3">
7. Elytra setose, shiny; body length less than
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.............................
<taxonomicName id="82EE94F3EC075A7E6F261BFEFAB7F801" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[1038,1309,1958,1980]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="truncata" status="sp. nov.">
<emphasis id="779A3362EC075A7E6F261BFEFB16F801" box="[1038,1212,1959,1980]" italics="true" pageId="2" pageNumber="3">Liogenys truncata</emphasis>
Cherman
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,
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</paragraph>
<paragraph id="4551EF70EC075A7E6BBF1B9DFA36F867" blockId="2.[151,1437,1372,2039]" box="[151,1436,1988,2010]" pageId="2" pageNumber="3">
<keyLead id="FE1F4C45EC075A7E6BBF1B9DFA36F867" box="[151,1436,1988,2010]" pageId="2" pageNumber="3">
- Elytra glabrous, semiopaque; body length
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..................................
<taxonomicName id="82EE94F3EC075A7E6F441B9DFA36F867" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1132,1436,1988,2010]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC075A7E6F441B9DFA9AF864" box="[1132,1328,1988,2009]" italics="true" pageId="2" pageNumber="3">Liogenys rectangula</emphasis>
<bibRefCitation id="217F9281EC075A7E6E1E1B9DFA36F867" author="Frey, G." box="[1334,1436,1988,2010]" pageId="2" pageNumber="3" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
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<keyStep id="FE1AF7D5EC075A7F6BBF1BB8FA36FF77" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<paragraph id="4551EF70EC075A7F6BBF1BB8FA36FF10" blockId="2.[151,1437,1372,2039]" lastBlockId="3.[151,1437,151,1288]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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8. Pronotum roughly punctate; lateral sides of abdomen, propygidium, and pygidium covered by wide scales (
<figureCitation id="DDD5F3F5EC075A7E6FC11BB8FA95F84A" box="[1257,1343,2017,2039]" captionStart="FIGURE 20" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@45.[203,1385,181,1920]" captionTargetPageId="45" captionText="FIGURE 20. Liogenys rugosicollis Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561318" httpUri="https://zenodo.org/record/4561318/files/figure.png" pageId="2" pageNumber="3">Fig. 20C</figureCitation>
).................................................................................
<taxonomicName id="82EE94F3EC065A7F6F4A1CC1FA36FF10" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1122,1436,151,173]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rugosicollis">
<emphasis id="779A3362EC065A7F6F4A1CC1FA85FF11" box="[1122,1327,151,173]" italics="true" pageId="3" pageNumber="4">Liogenys rugosicollis</emphasis>
<bibRefCitation id="217F9281EC065A7F6E1E1CC1FA36FF10" author="Frey, G." box="[1334,1436,151,173]" pageId="3" pageNumber="4" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
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</paragraph>
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- Pronotum somewhat coarsely punctate; sides of abdomen, propygidium, and pygidium with bristles (
<figureCitation id="DDD5F3F5EC065A7F6F841CECFAA8FF76" box="[1196,1282,181,203]" captionStart="FIGURE 10" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@24.[203,1385,181,1920]" captionTargetPageId="24" captionText="FIGURE 10. Liogenys latitarsis Moser, 1918 male lectotype views: A) dorsal, B) labels, C) head and pronotum, D) lateral, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561288" httpUri="https://zenodo.org/record/4561288/files/figure.png" pageId="3" pageNumber="4">Fig. 10D</figureCitation>
)........... 9
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</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1C8BFA36FEB8" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1C8BFA36FF5A" blockId="3.[151,1437,151,1288]" box="[151,1436,210,232]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1C8BFA36FF5A" box="[151,1436,210,232]" pageId="3" pageNumber="4">
9. Width of clypeal teeth combined three or more times longer than the length of one tooth (
<figureCitation id="DDD5F3F5EC065A7F6F141C8BFB36FF5A" box="[1084,1180,210,232]" captionStart="FIGURE 14" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[203,1385,186,1920]" captionTargetId="figure-20@31.[203,1385,181,1920]" captionTargetPageId="31" captionText="FIGURE 14. Liogenys nigrofusca Moser, 1918 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561298" httpUri="https://zenodo.org/record/4561298/files/figure.png" pageId="3" pageNumber="4">Figs. 14B</figureCitation>
,
<figureCitation id="DDD5F3F5EC065A7F6F8E1C8BFB65FF55" box="[1190,1231,210,232]" captionStart="FIGURE 18" captionStartId="41.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@41.[203,1385,181,1920]" captionTargetPageId="41" captionText="FIGURE 18. Liogenys parallela Frey, 1965 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, G) dorsolateral, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561314" httpUri="https://zenodo.org/record/4561314/files/figure.png" pageId="3" pageNumber="4">18B</figureCitation>
).............. 10
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1CA9FA36FEB8" blockId="3.[151,1437,151,1288]" box="[151,1436,239,262]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1CA9FA36FEB8" box="[151,1436,239,262]" pageId="3" pageNumber="4">
- Width of clypeal teeth combined less than three times longer than the length of one tooth (
<figureCitation id="DDD5F3F5EC065A7F6F6C1CB6FB32FEB8" box="[1092,1176,239,261]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="3" pageNumber="4">Figs. 6B</figureCitation>
,
<figureCitation id="DDD5F3F5EC065A7F6F8A1CA9FB15FEBB" box="[1186,1215,240,262]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="3" pageNumber="4">8B</figureCitation>
)............... 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1D54FA36FEE0" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1D54FA36FEFD" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1D54FA36FEFD" pageId="3" pageNumber="4">
10. Epipleuron of elytra uniformly clothed by fringed-like, long, erect bristles (
<figureCitation id="DDD5F3F5EC065A7F68931D54FB9AFE9F" box="[955,1072,269,291]" captionStart="FIGURE 18" captionStartId="41.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@41.[203,1385,181,1920]" captionTargetPageId="41" captionText="FIGURE 18. Liogenys parallela Frey, 1965 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, G) dorsolateral, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561314" httpUri="https://zenodo.org/record/4561314/files/figure.png" pageId="3" pageNumber="4">Fig. 18A, E</figureCitation>
); legs with tarsomere II twice as long as tarsomere I, or almost so, especially in male.......................................
<taxonomicName id="82EE94F3EC065A7F6F531D72FA36FEFD" authority="Frey, 1965" authorityName="Frey" authorityYear="1965" box="[1147,1436,298,320]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parallela">
<emphasis id="779A3362EC065A7F6F531D72FA9AFE82" box="[1147,1328,298,320]" italics="true" pageId="3" pageNumber="4">Liogenys parallela</emphasis>
<bibRefCitation id="217F9281EC065A7F6E1E1D72FA36FEFD" author="Frey, G." box="[1334,1436,298,320]" pageId="3" pageNumber="4" pagination="143 - 151" refId="ref36131" refString="Frey, G. (1965) Neue s ʾ damerikanische Melolonthidae (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 16, 143 - 151." type="journal article" year="1965">Frey, 1965</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1D11FA36FEE0" blockId="3.[151,1437,151,1288]" box="[151,1436,327,349]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1D11FA36FEE0" box="[151,1436,327,349]" pageId="3" pageNumber="4">- Epipleuron of elytra with sparse, short bristles; legs with tarsomere II less than twice as long as tarsomere I............. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1D3CFA36FE6F" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1D3CFA36FE2A" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1D3CFA36FE2A" pageId="3" pageNumber="4">
11. Pygidium subquadrate (
<figureCitation id="DDD5F3F5EC065A7F6AEE1D3CFDB5FEC6" box="[454,543,357,379]" captionStart="FIGURE 14" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[203,1385,186,1920]" captionTargetId="figure-20@31.[203,1385,181,1920]" captionTargetPageId="31" captionText="FIGURE 14. Liogenys nigrofusca Moser, 1918 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561298" httpUri="https://zenodo.org/record/4561298/files/figure.png" pageId="3" pageNumber="4">Fig. 14D</figureCitation>
), apex angulate, punctures very coarse; clypeal emargination deeper than the outer margin of anterior teeth (
<figureCitation id="DDD5F3F5EC065A7F6A461DDBFE6EFE25" box="[366,452,386,408]" captionStart="FIGURE 14" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[203,1385,186,1920]" captionTargetId="figure-20@31.[203,1385,181,1920]" captionTargetPageId="31" captionText="FIGURE 14. Liogenys nigrofusca Moser, 1918 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561298" httpUri="https://zenodo.org/record/4561298/files/figure.png" pageId="3" pageNumber="4">Fig. 14B</figureCitation>
)......................................................
<taxonomicName id="82EE94F3EC065A7F6F731DDAFA36FE2A" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[1115,1436,386,408]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nigrofusca">
<emphasis id="779A3362EC065A7F6F731DDAFAB4FE2A" box="[1115,1310,386,408]" italics="true" pageId="3" pageNumber="4">Liogenys nigrofusca</emphasis>
<bibRefCitation id="217F9281EC065A7F6E0C1DDAFA36FE2A" author="Moser, J." box="[1316,1436,386,408]" pageId="3" pageNumber="4" pagination="95 - 167" refId="ref36796" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">Moser, 1918</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1DF9FA36FE6F" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1DF9FA36FE6F" pageId="3" pageNumber="4">
- Pygidium trapezoidal (
<figureCitation id="DDD5F3F5EC065A7F6A961DC6FDABFE08" box="[446,513,415,437]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="3" pageNumber="4">Figs. 2</figureCitation>
DE; 5GH; 15G), apex subangulate or rounded; clypeal emargination shallower than the outer margin of anterior teeth (
<figureCitation id="DDD5F3F5EC065A7F6AFA1DE4FD82FE6E" box="[466,552,445,467]" captionStart="FIGURE 10" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@24.[203,1385,181,1920]" captionTargetPageId="24" captionText="FIGURE 10. Liogenys latitarsis Moser, 1918 male lectotype views: A) dorsal, B) labels, C) head and pronotum, D) lateral, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561288" httpUri="https://zenodo.org/record/4561288/files/figure.png" pageId="3" pageNumber="4">Fig. 10C</figureCitation>
)....................................................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1D83FA36FDFA" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1D83FA36FDB0" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1D83FA36FDB0" pageId="3" pageNumber="4">
12. Lateral margin of clypeus weakly produced, outer sides of anterior teeth obtuse or subparallel, without angle among anterior teeth and lateral margin of clypeus (
<figureCitation id="DDD5F3F5EC065A7F69131DAEFD25FDB0" box="[571,655,503,525]" captionStart="FIGURE 15" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@34.[203,1385,181,1920]" captionTargetPageId="34" captionText="FIGURE 15. Liogenys obscura Blanchard, 1851 female lectotype views: A) dorsal, C) labels; L. bruchi Moser, 1921 male lectotype views: B) dorsal, D) labels, E) head and pronotum, F) lateral, G) pygidium, and parameres in views: H) dorsal, I) dorsolateral and J) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561302" httpUri="https://zenodo.org/record/4561302/files/figure.png" pageId="3" pageNumber="4">Fig. 15E</figureCitation>
).............................................................. 13
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1E4CFA36FDFA" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1E4CFA36FDFA" pageId="3" pageNumber="4">
- Lateral margin of clypeus strongly produced, outer sides of anterior teeth almost parallel, right or acute angle between anterior and lateral teeth (
<figureCitation id="DDD5F3F5EC065A7F6AA01E6BFE74FDF5" box="[392,478,562,584]" captionStart="FIGURE 10" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@24.[203,1385,181,1920]" captionTargetPageId="24" captionText="FIGURE 10. Liogenys latitarsis Moser, 1918 male lectotype views: A) dorsal, B) labels, C) head and pronotum, D) lateral, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561288" httpUri="https://zenodo.org/record/4561288/files/figure.png" pageId="3" pageNumber="4">Fig. 10C</figureCitation>
)............................................................................. 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1E09FA36FD3E" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1E09FA36FDD8" blockId="3.[151,1437,151,1288]" box="[151,1436,591,613]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1E09FA36FDD8" box="[151,1436,591,613]" pageId="3" pageNumber="4">
13. Propygidium scaly; pygidium with erect and thick setae............................
<taxonomicName id="82EE94F3EC065A7F6F661E09FA36FDD8" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[1102,1436,591,613]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="779A3362EC065A7F6F661E09FB5DFDD9" box="[1102,1271,591,613]" italics="true" pageId="3" pageNumber="4">Liogenys obscura</emphasis>
<bibRefCitation id="217F9281EC065A7F6FD61E16FA36FDD8" author="Blanchard, E." box="[1278,1436,591,613]" pageId="3" pageNumber="4" pagination="129 - 240" refId="ref35099" refString="Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240." type="book chapter" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1E34FA36FD3E" blockId="3.[151,1437,151,1288]" box="[151,1436,621,643]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1E34FA36FD3E" box="[151,1436,621,643]" pageId="3" pageNumber="4">
- Propygidium with sparse bristles; pygidium with decumbent bristles....................
<taxonomicName id="82EE94F3EC065A7F6F4F1E34FA36FD3E" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1127,1436,621,643]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC065A7F6F4F1E34FA9AFD3F" box="[1127,1328,621,642]" italics="true" pageId="3" pageNumber="4">Liogenys cartwrighti</emphasis>
<bibRefCitation id="217F9281EC065A7F6E1E1E34FA36FD3E" author="Frey, G." box="[1334,1436,621,643]" pageId="3" pageNumber="4" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1ED3FA36FD66" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1ED3FA36FD00" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1ED3FA36FD00" pageId="3" pageNumber="4">14. Pygidium glabrous, or with few bristles on apex; distance between teeth of each tarsal claw as long as the length of superior tooth.............................................................................................. 15</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1E9CFA36FD66" blockId="3.[151,1437,151,1288]" box="[151,1436,709,731]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1E9CFA36FD66" box="[151,1436,709,731]" pageId="3" pageNumber="4">- Pygidium setose, distance between teeth of each tarsal claw shorter than the length of inferior tooth................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1EBBFA36FC36" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1EBBFA36FC8E" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1EBBFA36FC8E" pageId="3" pageNumber="4">
15. Body shiny, head and pronotum unicolored brown or dark brown in both sexes; clypeal emargination narrow, without clear sexual dimorphism in the clypeal margin; males with protarsomere II longer than wide (
<figureCitation id="DDD5F3F5EC065A7F6F061EA6FBDBFCA8" box="[1070,1137,767,789]" captionStart="FIGURE 5" captionStartId="14.[151,250,1933,1957]" captionTargetBox="[151,1435,181,1910]" captionTargetId="figure-21@14.[151,1435,181,1910]" captionTargetPageId="14" captionText="FIGURE 5. Liogenys denticeps Blanchard, 1851 female lectotype views: C) dorsal, F) labels, I) pygidium; L. denticulata Moser, 1918 male lectotype: A) dorsal, D) labels, G) pygidium, J) lateral, and parameres views: K) dorsal and L) lateral; L. denticulata female paralectotype views: B) dorsal, E) labels." figureDoi="http://doi.org/10.5281/zenodo.4561274" httpUri="https://zenodo.org/record/4561274/files/figure.png" pageId="3" pageNumber="4">Fig. 5J</figureCitation>
)...............................................................................................
<taxonomicName id="82EE94F3EC065A7F6F681F44FA36FC8E" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[1088,1436,797,819]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="denticeps">
<emphasis id="779A3362EC065A7F6F681F44FB52FC8F" box="[1088,1272,797,818]" italics="true" pageId="3" pageNumber="4">Liogenys denticeps</emphasis>
<bibRefCitation id="217F9281EC065A7F6FD61F44FA36FC8E" author="Blanchard, E." box="[1278,1436,797,819]" pageId="3" pageNumber="4" pagination="129 - 240" refId="ref35099" refString="Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240." type="book chapter" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1F62FA36FC36" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1F62FA36FC36" pageId="3" pageNumber="4">
- Body opaque, head and pronotum dull yellow unicolored in males, not unicolored in females, head dark brown or black; clypeal emargination very wide, in females emargination as wide as one half the distance between eyes, in males wider; males with protarsomere II as long as wide or wider than long (
<figureCitation id="DDD5F3F5EC065A7F69FC1F2CFC80FC36" box="[724,810,885,907]" captionStart="FIGURE 10" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@24.[203,1385,181,1920]" captionTargetPageId="24" captionText="FIGURE 10. Liogenys latitarsis Moser, 1918 male lectotype views: A) dorsal, B) labels, C) head and pronotum, D) lateral, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561288" httpUri="https://zenodo.org/record/4561288/files/figure.png" pageId="3" pageNumber="4">Fig. 10D</figureCitation>
)........................
<taxonomicName id="82EE94F3EC065A7F6F7E1F2CFA36FC36" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[1110,1436,885,907]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="779A3362EC065A7F6F7E1F2CFB5DFC37" box="[1110,1271,885,906]" italics="true" pageId="3" pageNumber="4">Liogenys pallens</emphasis>
<bibRefCitation id="217F9281EC065A7F6FD61F2CFA36FC36" author="Blanchard, E." box="[1278,1436,885,907]" pageId="3" pageNumber="4" pagination="129 - 240" refId="ref35099" refString="Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240." type="book chapter" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1FCBFA36FBBD" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1FCBFA36FC78" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1FCBFA36FC78" pageId="3" pageNumber="4">
16. Metafemur glabrous, row of bristles along its margins; males with parameres angulate at the midline, without subapical constriction (
<figureCitation id="DDD5F3F5EC065A7F6A161FF6FE28FC7B" box="[318,386,943,966]" captionStart="FIGURE 11" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@25.[203,1385,181,1920]" captionTargetPageId="25" captionText="FIGURE 11. Liogenys latitarsis Moser, 1918 intraspecific variations in shape of parameres, views: AE) dorsal, FJ) dorsolateral." figureDoi="http://doi.org/10.5281/zenodo.4561292" httpUri="https://zenodo.org/record/4561292/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
).............................................................
<taxonomicName id="82EE94F3EC065A7F6F441FE9FA36FC78" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[1132,1436,943,965]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="latitarsis">
<emphasis id="779A3362EC065A7F6F441FE9FAB4FC79" box="[1132,1310,943,965]" italics="true" pageId="3" pageNumber="4">Liogenys latitarsis</emphasis>
<bibRefCitation id="217F9281EC065A7F6E0C1FE9FA36FC78" author="Moser, J." box="[1316,1436,944,965]" pageId="3" pageNumber="4" pagination="95 - 167" refId="ref36796" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">Moser, 1918</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF1F94FA36FBBD" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1F94FA36FBBD" pageId="3" pageNumber="4">
- Metafemur setose throughout; males with parameres globose at the midline, with subapical constriction (
<figureCitation id="DDD5F3F5EC065A7F6FF91F94FA8FFC5E" box="[1233,1317,973,995]" captionStart="FIGURE 22" captionStartId="50.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@50.[203,1385,181,1920]" captionTargetPageId="50" captionText="FIGURE 22. Liogenys vicina Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, F) dorsolateral, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561322" httpUri="https://zenodo.org/record/4561322/files/figure.png" pageId="3" pageNumber="4">Fig. 22E</figureCitation>
).......................................................................................
<taxonomicName id="82EE94F3EC065A7F6FB01FB2FA36FBBD" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1176,1436,1002,1024]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="779A3362EC065A7F6FB01FB2FA9AFBBD" box="[1176,1328,1003,1024]" italics="true" pageId="3" pageNumber="4">Liogenys vicina</emphasis>
<bibRefCitation id="217F9281EC065A7F6E1E1FB2FA36FBBD" author="Frey, G." box="[1334,1436,1002,1024]" pageId="3" pageNumber="4" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1851FA36FBEA" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1851FA37FB86" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1851FA37FB86" pageId="3" pageNumber="4">
17. Clypeal emargination angulate (
<figureCitation id="DDD5F3F5EC065A7F693A185EFDF7FBA3" box="[530,605,1031,1054]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="3" pageNumber="4">Fig. 8B</figureCitation>
); fovea of distal maxillary palp occupying the entire dorsal surface, almost reaching the apex (
<figureCitation id="DDD5F3F5EC065A7F6A34187CFECFFB86" box="[284,357,1061,1083]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys foveata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) distal maxillary palp, D) lateral, E) mesotibia and metatibia, F) pygidium, and parameres in views: G) dorsal, and H) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561282" httpUri="https://zenodo.org/record/4561282/files/figure.png" pageId="3" pageNumber="4">Fig. 8C</figureCitation>
).........................................................
<taxonomicName id="82EE94F3EC065A7F6F37187CFA8AFB86" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[1055,1312,1061,1083]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="foveata" status="sp. nov.">
<emphasis id="779A3362EC065A7F6F37187CFB69FB87" box="[1055,1219,1061,1082]" italics="true" pageId="3" pageNumber="4">Liogenys foveata</emphasis>
Cherman
</taxonomicName>
<emphasis id="779A3362EC065A7F6E0E187CFA37FB86" bold="true" box="[1318,1437,1061,1083]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="6CA98E19EC065A7F6E0E187CFA37FB86" box="[1318,1437,1061,1083]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF181AFA36FBEA" blockId="3.[151,1437,151,1288]" box="[151,1436,1090,1112]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF181AFA36FBEA" box="[151,1436,1090,1112]" pageId="3" pageNumber="4">- Clypeal emargination rounded; fovea of distal maxillary palp extending past the midline but not reaching the apex....... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF1839FA36FB12" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF1839FA36FB2F" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF1839FA36FB2F" pageId="3" pageNumber="4">
18. Base of pygidium less than twice as wide as the apex (
<figureCitation id="DDD5F3F5EC065A7F69C41806FC9DFBC8" box="[748,823,1119,1141]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="3" pageNumber="4">Fig. 7E</figureCitation>
); in males, the three pairs of tarsi equally enlarged......................................................................................
<taxonomicName id="82EE94F3EC065A7F6F511824FA36FB2F" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[1145,1436,1149,1170]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="779A3362EC065A7F6F511824FAB4FB2F" box="[1145,1310,1149,1170]" italics="true" pageId="3" pageNumber="4">Liogenys tarsalis</emphasis>
<bibRefCitation id="217F9281EC065A7F6E0C1824FA36FB2F" author="Moser, J." box="[1316,1436,1149,1170]" pageId="3" pageNumber="4" pagination="133 - 182" refId="ref36844" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF18C2FA36FB12" blockId="3.[151,1437,151,1288]" box="[151,1436,1178,1200]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF18C2FA36FB12" box="[151,1436,1178,1200]" pageId="3" pageNumber="4">
- Base of pygidium twice as wide as the apex (
<figureCitation id="DDD5F3F5EC065A7F69BB18C3FD49FB0D" box="[659,739,1178,1200]" captionStart="FIGURE 7" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@17.[203,1385,181,1920]" captionTargetPageId="17" captionText="FIGURE 7. Liogenys tarsalis Moser, 1921 female views: A) dorsal, C) head and pronotum, E) pygidium; compared with L. forcipata Frey, 1970 female views: B) dorsal, D) head and pronotum, F) pygidium." figureDoi="http://doi.org/10.5281/zenodo.4561280" httpUri="https://zenodo.org/record/4561280/files/figure.png" pageId="3" pageNumber="4">Figs. 7F</figureCitation>
,
<figureCitation id="DDD5F3F5EC065A7F69C418C3FCB3FB0D" box="[748,793,1178,1200]" captionStart="FIGURE 13" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 13. Liogenys neoforcipata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) base of metatibia, white arrow indicates basal apophysis and metatrochanter, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561296" httpUri="https://zenodo.org/record/4561296/files/figure.png" pageId="3" pageNumber="4">13D</figureCitation>
); in males, protarsi and mesotarsi wider than the metatarsi.... 19
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FE1AF7D5EC065A7F6BBF18E1FA36FAB5" pageId="3" pageNumber="4">
<paragraph id="4551EF70EC065A7F6BBF18E1FA37FB56" blockId="3.[151,1437,151,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF18E1FA37FB56" pageId="3" pageNumber="4">
19. Base of metacoxae and trochanter setose (
<figureCitation id="DDD5F3F5EC065A7F695D18EEFD61FB73" box="[629,715,1207,1230]" captionStart="FIGURE 13" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 13. Liogenys neoforcipata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) base of metatibia, white arrow indicates basal apophysis and metatrochanter, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561296" httpUri="https://zenodo.org/record/4561296/files/figure.png" pageId="3" pageNumber="4">Fig. 13E</figureCitation>
); in males, parameres glabrous (
<figureCitation id="DDD5F3F5EC065A7F6F2A18EEFBFFFB73" box="[1026,1109,1207,1230]" captionStart="FIGURE 13" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 13. Liogenys neoforcipata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) base of metatibia, white arrow indicates basal apophysis and metatrochanter, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561296" httpUri="https://zenodo.org/record/4561296/files/figure.png" pageId="3" pageNumber="4">Fig. 13F</figureCitation>
)..........................................................................................
<taxonomicName id="82EE94F3EC065A7F68CE188CFAB7FB56" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[998,1309,1237,1259]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="neoforcipata" status="sp. nov.">
<emphasis id="779A3362EC065A7F68CE188CFB17FB57" box="[998,1213,1237,1258]" italics="true" pageId="3" pageNumber="4">Liogenys neoforcipata</emphasis>
Cherman
</taxonomicName>
,
<emphasis id="779A3362EC065A7F6E0E188CFA37FB56" bold="true" box="[1318,1437,1237,1259]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="6CA98E19EC065A7F6E0E188CFA37FB56" box="[1318,1437,1237,1259]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="4551EF70EC065A7F6BBF18AAFA36FAB5" blockId="3.[151,1437,151,1288]" box="[151,1436,1266,1288]" pageId="3" pageNumber="4">
<keyLead id="FE1F4C45EC065A7F6BBF18AAFA36FAB5" box="[151,1436,1266,1288]" pageId="3" pageNumber="4">
- Base of metacoxae and trochanter almost glabrous (
<figureCitation id="DDD5F3F5EC065A7F69FB18ABFC8BFAB5" box="[723,801,1266,1288]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="3" pageNumber="4">Fig. 6D</figureCitation>
); in males, parameres setose (
<figureCitation id="DDD5F3F5EC065A7F6F1718ABFB2DFAB5" box="[1087,1159,1266,1288]" captionStart="FIGURE 6" captionStartId="16.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@16.[203,1385,181,1920]" captionTargetPageId="16" captionText="FIGURE 6. Liogenys forcipata Frey, 1970 male views: A) dorsal, B) head and pronotum, C) lateral, D) base of metatibia, white arrow indicates basal apophysis and metatrochanter, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561276" httpUri="https://zenodo.org/record/4561276/files/figure.png" pageId="3" pageNumber="4">Fig. 6F</figureCitation>
)...
<taxonomicName id="82EE94F3EC065A7F6F9E18AAFA36FAB5" authority="Frey" authorityName="Frey" authorityYear="1970" box="[1206,1436,1266,1288]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis id="779A3362EC065A7F6F9E18AAFAC0FABA" box="[1206,1386,1266,1288]" italics="true" pageId="3" pageNumber="4">Liogenys forcipata</emphasis>
Frey
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>