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<mods:title id="5633FB1E4B7D1339D7270C37C9D74040">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<emphasis id="779A3362EC005A796BBF1F1AFE38FCE0" bold="true" box="[151,402,834,861]" italics="true" pageId="5" pageNumber="6">Liogenys cartwrighti</emphasis>
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<bibRefCitation id="217F9281EC005A796A171F93FD80FC5F" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[319,554,970,994]" pageId="5" pageNumber="6" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
<emphasis id="779A3362EC005A796A8B1F92FE7CFC5F" box="[419,470,970,994]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
2017: 5
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<emphasis id="779A3362EC005A796BBF1849FE95FB97" bold="true" box="[151,319,1040,1066]" pageId="5" pageNumber="6">Type material</emphasis>
.
<taxonomicName id="82EE94F3EC005A796A661848FD93FB97" authorityName="Frey" authorityYear="1969" box="[334,569,1040,1066]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC005A796A661848FD93FB97" box="[334,569,1040,1066]" italics="true" pageId="5" pageNumber="6">Liogenys cartwrighti</emphasis>
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male
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(
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): [white handwritten] “Sgo. d.
<collectingCounty id="AC3097FCEC005A796F8E1848FB44FB96" box="[1190,1262,1041,1067]" pageId="5" pageNumber="6">Estero</collectingCounty>
/
<collectingCountry id="3DF9AFE0EC005A796E2C1849FA25FB97" box="[1284,1423,1040,1066]" name="Argentina" pageId="5" pageNumber="6">Argentinien</collectingCountry>
”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC005A796938186DFDD1FBF3" authorityName="Guerin-Meneville" authorityYear="1831" box="[528,635,1076,1102]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ cartwrighti / n.sp. /
<collectingMunicipality id="A535750AEC005A796841186CFC0BFBF2" box="[873,929,1077,1103]" pageId="5" pageNumber="6">Type</collectingMunicipality>
/ det.
<determinerName id="B08D03B4EC005A7968CD186DFB9EFBF2" box="[997,1076,1076,1103]" pageId="5" pageNumber="6">G Frey</determinerName>
, 1968”, [red typeset] “TYPUS”, [pink typeset] “
<location id="4031B9ABEC005A796A691801FE17FBCF" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796A691801FE17FBCF" box="[321,445,1112,1138]" country="Argentina" county="Estero" municipality="Type" name="Cartwright" pageId="5" pageNumber="6" stateProvince="Salta">Cartwright</location>
/ Smithson.”, [red, typeset and handwritten] “
<location id="4031B9ABEC005A7968921801FBB7FBCE" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A7968921801FBB7FBCE" box="[954,1053,1112,1139]" country="Argentina" county="Estero" municipality="Type" name="Type No" pageId="5" pageNumber="6" stateProvince="Salta">Type No</location>
/ 74011 / USNM”, [white typeset] “
<location id="4031B9ABEC005A796B891825FF77FB2B" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796B891825FF77FB2B" box="[161,221,1148,1174]" country="Argentina" county="Estero" municipality="Type" name="Loan" pageId="5" pageNumber="6" stateProvince="Salta">Loan</location>
from / USNMNH / 2065060”.
<typeStatus id="9A5551D2EC005A7969661824FD11FB2A" box="[590,699,1149,1175]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
(2): one female
<typeStatus id="9A5551D2EC005A7968A81824FC4AFB2A" box="[896,992,1149,1175]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
(
<collectionCode id="23FF77B5EC005A7968DA1825FBE3FB2B" box="[1010,1097,1148,1174]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="5" pageNumber="6">NHMB</collectionCode>
) [white handwritten] “
<collectingRegion id="872A2192EC005A796E781825FA21FB2B" box="[1360,1419,1148,1174]" country="Argentina" name="Salta" pageId="5" pageNumber="6">Salta</collectingRegion>
/ Dep. Rivadavia / XII.950
<location id="4031B9ABEC005A796A9218F9FD82FB07" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796A9218F9FD82FB07" box="[442,552,1184,1210]" country="Argentina" county="Estero" municipality="Type" name="Daguerre" pageId="5" pageNumber="6" stateProvince="Salta">Daguerre</location>
”, [orange empty], [white typeset] “
<specimenCount id="53E824F9EC005A79689A18F9FC6FFB07" box="[946,965,1184,1210]" pageId="5" pageNumber="6" type="female"></specimenCount>
”, [pink typeset] “Property / USNM”, [red typeset] “
<typeStatus id="9A5551D2EC005A796A28189DFE3EFB63" box="[256,404,1220,1246]" pageId="5" pageNumber="6" type="paratype">PARATYPE</typeStatus>
”, [white typeset and handwritten] “
<taxonomicName id="82EE94F3EC005A796832189DFC2FFB63" authorityName="Guerin-Meneville" authorityYear="1831" box="[794,901,1220,1246]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ cartwrighti / n.sp. / det.
<determinerName id="B08D03B4EC005A796F8A189DFB52FB62" box="[1186,1272,1220,1247]" pageId="5" pageNumber="6">G. Frey</determinerName>
, 1968”, [white typeset] “Museum Frey / M̹nchen”. One female
<typeStatus id="9A5551D2EC005A7969ED18B0FC8FFABE" box="[709,805,1257,1283]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
(
<collectionCode id="23FF77B5EC005A79681C18B1FC21FABF" box="[820,907,1256,1282]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="5" pageNumber="6">NHMB</collectionCode>
) [white handwritten] “
<collectingRegion id="872A2192EC005A796FA118B1FB6EFABF" box="[1161,1220,1256,1282]" country="Argentina" name="Salta" pageId="5" pageNumber="6">Salta</collectingRegion>
/ Dep. Rivadavia”, [white typeset] “
<specimenCount id="53E824F9EC005A796A731955FEC4FA9B" box="[347,366,1292,1318]" pageId="5" pageNumber="6" type="female"></specimenCount>
”, [pink typeset] “Cartwrighti / Smithson.”, [red typeset] “
<typeStatus id="9A5551D2EC005A796F3C1955FB02FA9B" box="[1044,1192,1292,1318]" pageId="5" pageNumber="6" type="paratype">PARATYPE</typeStatus>
”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC005A796A1F1969FE08FAF7" authorityName="Guerin-Meneville" authorityYear="1831" box="[311,418,1328,1354]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ cartwrighti / n.sp. / det.
<determinerName id="B08D03B4EC005A7969EB1969FCB3FAF6" box="[707,793,1328,1355]" pageId="5" pageNumber="6">G. Frey</determinerName>
, 1968”
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC005A796BEF190DFA36F9D7" blockId="5.[151,1437,1039,2039]" pageId="5" pageNumber="6">
<emphasis id="779A3362EC005A796BEF190DFE0EFAD3" bold="true" box="[199,420,1364,1390]" pageId="5" pageNumber="6">Non-type material</emphasis>
(12).
<materialsCitation id="F586E52DEC005A796AC2190DFF53FA2F" ID-GBIF-Occurrence="3047558464" collectionCode="MLPA" collectorName="R. Richter" country="Argentina" location="Jujuy" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Jujuy">
<collectingCountry id="3DF9AFE0EC005A796AC2190DFD38FAD3" box="[490,658,1364,1390]" name="Argentina" pageId="5" pageNumber="6">ARGENTINA</collectingCountry>
:
<collectingRegion id="872A2192EC005A7969B6190DFD75FAD3" box="[670,735,1364,1390]" country="Argentina" name="Jujuy" pageId="5" pageNumber="6">Jujuy</collectingRegion>
: without locality and date,
<collectorName id="E81B8AA6EC005A796F30190CFB27FAD3" box="[1048,1165,1364,1391]" pageId="5" pageNumber="6">R. Richter</collectorName>
<specimenCount id="53E824F9EC005A796FBE190DFB49FAD3" box="[1174,1251,1364,1390]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
and one female (
<collectionCode id="23FF77B5EC005A796BB71921FF5BFA2F" box="[159,241,1400,1426]" pageId="5" pageNumber="6">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A796A2D1920FCFFFA2F" ID-GBIF-Occurrence="3047558417" box="[261,853,1400,1427]" collectingDate="1951-01" collectionCode="DZUP" collectorName="F. Justus Jor." country="Argentina" location="Los Perales" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Jujuy">
<location id="4031B9ABEC005A796A2D1920FE20FA2F" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796A2D1920FE20FA2F" box="[261,394,1400,1427]" country="Argentina" name="Los Perales" pageId="5" pageNumber="6" stateProvince="Jujuy">Los Perales</location>
,
<date id="3150C9B0EC005A796ABE1921FE4BFA2E" box="[406,481,1400,1427]" pageId="5" pageNumber="6" value="1951-01">
<collectingDate id="21143058EC005A796ABE1921FE4BFA2E" box="[406,481,1400,1427]" pageId="5" pageNumber="6" value="1951-01">I.1951</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC005A796AC51920FD2AFA2E" box="[493,640,1401,1427]" pageId="5" pageNumber="6">F. Justus Jor.</collectorName>
,
<specimenCount id="53E824F9EC005A7969A41921FD44FA2F" box="[652,750,1400,1426]" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
(
<collectionCode id="23FF77B5EC005A7969D61921FCE7FA2F" box="[766,845,1400,1426]" country="Brazil" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="6" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A7968491920FECBFA0B" ID-GBIF-Occurrence="3047558370" collectingDate="1986-11-29" collectionCode="IADIZA" collectorName="J. Muzon" country="Argentina" location="Parque Nacional Calilegua" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Jujuy">
<location id="4031B9ABEC005A7968491920FB38FA2F" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A7968491920FB38FA2F" box="[865,1170,1400,1427]" country="Argentina" name="Parque Nacional Calilegua" pageId="5" pageNumber="6" stateProvince="Jujuy">Parque Nacional Calilegua</location>
,
<date id="3150C9B0EC005A796FB61921FA8AFA2F" box="[1182,1312,1400,1426]" pageId="5" pageNumber="6" value="1986-11-29">
<collectingDate id="21143058EC005A796FB61921FA8AFA2F" box="[1182,1312,1400,1426]" pageId="5" pageNumber="6" value="1986-11-29">29.XI.1986</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC005A796E041920FA32FA2F" box="[1324,1432,1400,1427]" pageId="5" pageNumber="6">J. Muzón</collectorName>
,
<specimenCount id="53E824F9EC005A796BBF19C5FF48FA0B" box="[151,226,1436,1462]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC005A796BDA19C5FEF3FA0B" box="[242,345,1436,1462]" pageId="5" pageNumber="6">IADIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A796A4419C4FEA4FA67" ID-GBIF-Occurrence="3047558310" collectingDate="2008-12-10" collectionCode="IADIZA" collectorName="F. C. Ocampo &amp; G. San Blas &amp; Campon" country="Argentina" location="Parque Nacional Calilegua" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Jujuy">
near
<location id="4031B9ABEC005A796A8B19C4FD79FA0B" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796A8B19C4FD79FA0B" box="[419,723,1436,1463]" country="Argentina" name="Parque Nacional Calilegua" pageId="5" pageNumber="6" stateProvince="Jujuy">Parque Nacional Calilegua</location>
,
<date id="3150C9B0EC005A7969F619C5FCC3FA0B" box="[734,873,1436,1462]" pageId="5" pageNumber="6" value="2008-12-10">
<collectingDate id="21143058EC005A7969F619C5FCC3FA0B" box="[734,873,1436,1462]" pageId="5" pageNumber="6" value="2008-12-10">10.XII.2008</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC005A79685D19C5FBA6FA0B" box="[885,1036,1436,1462]" pageId="5" pageNumber="6">F.C. Ocampo</collectorName>
,
<collectorName id="E81B8AA6EC005A796F3F19C5FB35FA0B" box="[1047,1183,1436,1462]" pageId="5" pageNumber="6">G. San Blas</collectorName>
, &amp; F.
<collectorName id="E81B8AA6EC005A796FCB19C5FAEFFA0B" box="[1251,1349,1436,1462]" pageId="5" pageNumber="6">Campón</collectorName>
,
<specimenCount id="53E824F9EC005A796E7919C5FA36FA0B" box="[1361,1436,1436,1462]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC005A796BB71999FEACFA67" box="[159,262,1472,1498]" pageId="5" pageNumber="6">IADIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A796A311999FC31FA67" ID-GBIF-Occurrence="3047558463" box="[281,923,1472,1499]" collectingDate="1953-03" collectionCode="MZSP" collectorName="Martinez" country="Argentina" location="Dique la Cienaga" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Jujuy">
<location id="4031B9ABEC005A796A311999FE4AFA67" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796A311999FE4AFA67" box="[281,480,1472,1498]" country="Argentina" name="Dique la Cienaga" pageId="5" pageNumber="6" stateProvince="Jujuy">Dique la Ciénaga</location>
,
<date id="3150C9B0EC005A796AC31999FDE2FA66" box="[491,584,1472,1499]" pageId="5" pageNumber="6" value="1953-03">
<collectingDate id="21143058EC005A796AC31999FDE2FA66" box="[491,584,1472,1499]" pageId="5" pageNumber="6" value="1953-03">III.1953</collectingDate>
</date>
, A.
<collectorName id="E81B8AA6EC005A79695D1999FD77FA67" box="[629,733,1472,1498]" pageId="5" pageNumber="6">Martínez</collectorName>
,
<specimenCount id="53E824F9EC005A7969C01999FC9EFA67" box="[744,820,1472,1498]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC005A79686B1999FC38FA67" box="[835,914,1472,1498]" country="Brazil" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A79688E1999FEABFA43" ID-GBIF-Occurrence="3047558460" collectingDate="1968-10-11" collectingDateMax="1968-10-12" collectingDateMin="1968-10-11" collectionCode="NHMB" collectorName="L. Pena" country="Argentina" location="Oran" pageId="5" pageNumber="6" specimenCount="2" specimenCount-male="2" stateProvince="Salta">
<collectingRegion id="872A2192EC005A79688E1999FC48FA67" box="[934,994,1472,1498]" country="Argentina" name="Salta" pageId="5" pageNumber="6">Salta</collectingRegion>
:
<location id="4031B9ABEC005A7968C61999FB83FA67" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A7968C61999FB83FA67" box="[1006,1065,1472,1498]" country="Argentina" name="Oran" pageId="5" pageNumber="6" stateProvince="Salta">Orán</location>
,
<date id="3150C9B0EC005A796F1C1999FB7FFA67" box="[1076,1237,1472,1498]" pageId="5" pageNumber="6" value="1968-10-11" valueMax="1968-10-12" valueMin="1968-10-11">
<collectingDate id="21143058EC005A796F1C1999FB7FFA67" box="[1076,1237,1472,1498]" pageId="5" pageNumber="6" value="1968-10-11" valueMax="1968-10-12" valueMin="1968-10-11">1112.X.1968</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC005A796FC91998FA90FA66" box="[1249,1338,1473,1499]" pageId="5" pageNumber="6">L. Peña</collectorName>
,
<specimenCount id="53E824F9EC005A796E6E1999FA36FA67" box="[1350,1436,1472,1498]" pageId="5" pageNumber="6" type="male">2 males</specimenCount>
(
<collectionCode id="23FF77B5EC005A796BB719BDFF53FA43" box="[159,249,1508,1534]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="5" pageNumber="6">NHMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A796A2619BDFEA4F99F" ID-GBIF-Occurrence="3047558339" collectingDate="1995-11-15" collectionCode="IADIZA" collectorName="Flores &amp; Roig" country="Argentina" latitude="-22.0509" location="Dique" longLatPrecision="7" longitude="-63.7101" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Salta">
<location id="4031B9ABEC005A796A2619BDFEF8FA43" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796A2619BDFEF8FA43" box="[270,338,1508,1534]" country="Argentina" latitude="-22.0509" longLatPrecision="7" longitude="-63.7101" name="Dique" pageId="5" pageNumber="6" stateProvince="Salta">Dique</location>
Itiyuro-
<locationDeviation id="54E3F652EC005A796A9A19BDFD02FA42" box="[434,680,1508,1535]" location="Salvador Mazza" pageId="5" pageNumber="6">
<quantity id="82164295EC005A796A9A19BDFE40FA43" box="[434,490,1508,1535]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="km" value="5.0">5 km</quantity>
Salvador Mazza
</locationDeviation>
[
<geoCoordinate id="20DA89B7EC005A79699E19BDFC85FA42" box="[694,815,1508,1535]" degrees="22.0509" direction="south" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="-22.0509">22.0509ºS</geoCoordinate>
,
<geoCoordinate id="20DA89B7EC005A79681419BAFC6AFA43" box="[828,960,1507,1534]" degrees="63.7101" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="-63.7101">63.7101ºW</geoCoordinate>
],
<date id="3150C9B0EC005A7968FA19BDFBFEFA42" box="[978,1108,1508,1535]" pageId="5" pageNumber="6" value="1995-11-15">
<collectingDate id="21143058EC005A7968FA19BDFBFEFA42" box="[978,1108,1508,1535]" pageId="5" pageNumber="6" value="1995-11-15">15.XI.1995</collectingDate>
</date>
, S.
<collectorName id="E81B8AA6EC005A796FA819BDFB6CFA43" box="[1152,1222,1508,1534]" pageId="5" pageNumber="6">Flores</collectorName>
&amp; S.
<collectorName id="E81B8AA6EC005A796E2219BDFAE9FA43" box="[1290,1347,1508,1534]" pageId="5" pageNumber="6">Roig</collectorName>
,
<specimenCount id="53E824F9EC005A796E7819BDFA36FA43" box="[1360,1436,1508,1534]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC005A796BB71A51FEACF99F" box="[159,262,1544,1570]" pageId="5" pageNumber="6">IADIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A796A321A50FB78F9FB" ID-GBIF-Occurrence="3047558306" collectingDate="2008-12-13" collectingDateMax="2008-12-17" collectingDateMin="2008-12-13" collectionCode="IADIZA" collectorName="S Ingeniero Juarez &amp; Ocampo &amp; San Blas &amp; Campon" country="Argentina" latitude="-24.090834" location="Ruta Provincial" longLatPrecision="21" longitude="-61.946945" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Formosa">
<collectingRegion id="872A2192EC005A796A321A50FE2AF99E" box="[282,384,1545,1571]" country="Argentina" name="Formosa" pageId="5" pageNumber="6">Formosa</collectingRegion>
:
<location id="4031B9ABEC005A796AA31A50FD90F99F" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796AA31A50FD90F99F" box="[395,570,1544,1571]" country="Argentina" latitude="-24.090834" longLatPrecision="21" longitude="-61.946945" name="Ruta Provincial" pageId="5" pageNumber="6" stateProvince="Formosa">Ruta Provincial</location>
39, 23 km
<collectorName id="E81B8AA6EC005A7969941A51FC3AF99F" box="[700,912,1544,1570]" pageId="5" pageNumber="6">S Ingeniero Juárez</collectorName>
,
<geoCoordinate id="20DA89B7EC005A7968B41A51FB88F99E" box="[924,1058,1544,1571]" degrees="24" direction="south" minutes="05" orientation="latitude" pageId="5" pageNumber="6" precision="15" seconds="27" value="-24.090834">24º0527S</geoCoordinate>
<geoCoordinate id="20DA89B7EC005A796F011A51FB10F99E" box="[1065,1210,1544,1571]" degrees="61" direction="west" minutes="56" orientation="longitude" pageId="5" pageNumber="6" precision="15" seconds="49" value="-61.946945">61º5649W</geoCoordinate>
,
<date id="3150C9B0EC005A796FEE1A51FAD6F99F" box="[1222,1404,1544,1570]" pageId="5" pageNumber="6" value="2008-12-13" valueMax="2008-12-17" valueMin="2008-12-13">
<collectingDate id="21143058EC005A796FEE1A51FAD6F99F" box="[1222,1404,1544,1570]" pageId="5" pageNumber="6" value="2008-12-13" valueMax="2008-12-17" valueMin="2008-12-13">1317.XII.2008</collectingDate>
</date>
, F.
<collectorName id="E81B8AA6EC005A796BBF1A75FF53F9FB" box="[151,249,1580,1606]" pageId="5" pageNumber="6">Ocampo</collectorName>
, G.
<collectorName id="E81B8AA6EC005A796A011A75FE3AF9FB" box="[297,400,1580,1606]" pageId="5" pageNumber="6">San Blas</collectorName>
, &amp; F.
<collectorName id="E81B8AA6EC005A796AF01A75FD90F9FB" box="[472,570,1580,1606]" pageId="5" pageNumber="6">Campón</collectorName>
, mercury vapor &amp;
<collectingMethod id="9CAF9767EC005A79683B1A75FC52F9FA" box="[787,1016,1580,1607]" pageId="5" pageNumber="6">ultraviolet light trap</collectingMethod>
,
<specimenCount id="53E824F9EC005A796F2E1A75FBF8F9FB" box="[1030,1106,1580,1606]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC005A796F4B1A75FB60F9FB" box="[1123,1226,1580,1606]" pageId="5" pageNumber="6">IADIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A796FF61A75FDFBF9D7" ID-GBIF-Occurrence="3047558470" collectingDate="1957-12" collectionCode="MZSP" collectorName="A. Martinez" country="Argentina" location="Lules" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman">
<collectingRegion id="872A2192EC005A796FF61A75FAE1F9FB" box="[1246,1355,1580,1606]" country="Argentina" name="Tucuman" pageId="5" pageNumber="6">Tucumán</collectingRegion>
:
<location id="4031B9ABEC005A796E7F1A75FA32F9FB" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A796E7F1A75FA32F9FB" box="[1367,1432,1580,1606]" country="Argentina" name="Lules" pageId="5" pageNumber="6" stateProvince="Tucuman">Lules</location>
,
<date id="3150C9B0EC005A796BBF1A09FF55F9D6" box="[151,255,1616,1643]" pageId="5" pageNumber="6" value="1957-12">
<collectingDate id="21143058EC005A796BBF1A09FF55F9D6" box="[151,255,1616,1643]" pageId="5" pageNumber="6" value="1957-12">XII.1957</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC005A796A221A09FE3EF9D7" box="[266,404,1616,1642]" pageId="5" pageNumber="6">A. Martínez</collectorName>
,
<specimenCount id="53E824F9EC005A796AB71A09FE40F9D7" box="[415,490,1616,1642]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC005A796AD11A09FDE2F9D7" box="[505,584,1616,1642]" country="Brazil" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="6" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC005A7969741A09FA32F9D7" ID-GBIF-Occurrence="3047558473" box="[604,1432,1616,1643]" collectingDate="1969-03" collectionCode="CMNC" collectorName="Martinez" country="Argentina" location="Ojo de Agua" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Santiago del Estero">
<collectingRegion id="872A2192EC005A7969741A09FC90F9D6" box="[604,826,1616,1643]" country="Argentina" name="Santiago del Estero" pageId="5" pageNumber="6">Santiago del Estero</collectingRegion>
:
<location id="4031B9ABEC005A79686C1A09FC7EF9D7" LSID="urn:lsid:plazi:treatment:CD475E66EC005A7B6BBF1F1AFCB8FA58:4031B9ABEC005A79686C1A09FC7EF9D7" box="[836,980,1616,1642]" country="Argentina" name="Ojo de Agua" pageId="5" pageNumber="6" stateProvince="Santiago del Estero">Ojo de Agua</location>
,
<date id="3150C9B0EC005A7968C81A09FB97F9D7" box="[992,1085,1616,1642]" pageId="5" pageNumber="6" value="1969-03">
<collectingDate id="21143058EC005A7968C81A09FB97F9D7" box="[992,1085,1616,1642]" pageId="5" pageNumber="6" value="1969-03">III.1969</collectingDate>
</date>
, A.
<collectorName id="E81B8AA6EC005A796F411A09FB7BF9D7" box="[1129,1233,1616,1642]" pageId="5" pageNumber="6">Martínez</collectorName>
,
<specimenCount id="53E824F9EC005A796FF51A09FA82F9D7" box="[1245,1320,1616,1642]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC005A796E1F1A09FA3AF9D7" box="[1335,1424,1616,1642]" pageId="5" pageNumber="6">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC005A796BEF1A2DFDCBF8DB" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="4551EF70EC005A796BEF1A2DFDCBF8DB" blockId="5.[151,1437,1039,2039]" pageId="5" pageNumber="6">
<emphasis id="779A3362EC005A796BEF1A2DFE90F933" bold="true" box="[199,314,1652,1678]" pageId="5" pageNumber="6">Diagnosis</emphasis>
. Body elongate, brown to dark brown, shiny (
<figureCitation id="DDD5F3F5EC005A79686C1A2DFC35F933" box="[836,927,1652,1678]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
); clypeal emargination deep but not reaching the base of the teeth, rounded and wide; lateral margin strongly convex and produced, forming a tooth-like projection; pronotal posterior corners in obtuse angle (
<figureCitation id="DDD5F3F5EC005A7969991AE5FCA3F96B" box="[689,777,1724,1750]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="5" pageNumber="6">Fig. 2B</figureCitation>
); basal apophysis of metacoxa produced; metatibia weakly or not carinate along the inner margin; superior tooth of a tarsal claw slightly longer than the inferior, distance between teeth shorter than the inferior tooth; pygidium with sparse bristles throughout (
<figureCitation id="DDD5F3F5EC005A796F5B1B5DFB1CF8A3" box="[1139,1206,1796,1822]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
DE); ventrites with abundant bristles (
<figureCitation id="DDD5F3F5EC005A796A401B71FE68F8FF" box="[360,450,1832,1858]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="5" pageNumber="6">Fig. 2C</figureCitation>
); in males protarsomere II as long as it is wide; parameres lyre shaped; ventral surface visible in dorsal view, concave (
<figureCitation id="DDD5F3F5EC005A796AD71B15FDFEF8DB" box="[511,596,1868,1894]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="5" pageNumber="6">Fig. 2F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC005A7B6BEF1B29FC87FC74" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="4551EF70EC005A7B6BEF1B29FC87FC74" blockId="5.[151,1437,1039,2039]" lastBlockId="7.[151,1437,151,1509]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<emphasis id="779A3362EC005A796BEF1B29FEC7F837" bold="true" box="[199,365,1904,1930]" pageId="5" pageNumber="6">Redescription</emphasis>
. Length
<quantity id="82164295EC005A796AFB1B29FDC6F836" box="[467,620,1904,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.085" metricValueMax="1.23" metricValueMin="0.94" pageId="5" pageNumber="6" unit="mm" value="10.85" valueMax="12.3" valueMin="9.4">9.412.3 mm</quantity>
; width:
<quantity id="82164295EC005A7969EF1B29FCFBF836" box="[711,849,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.9" metricValueMin="4.4" pageId="5" pageNumber="6" unit="mm" value="5.15" valueMax="5.9" valueMin="4.4">4.45.9 mm</quantity>
. Dark brown.
<emphasis id="779A3362EC005A7968DA1B29FB92F837" box="[1010,1080,1904,1930]" italics="true" pageId="5" pageNumber="6">Head:</emphasis>
distance between eyes twice as wide as one eye; clypeal emargination deep, rounded and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye; lateral margin strongly convex and produced, forming a tooth-like projection; distance between clypeal lateral projection and anterior margin of the eye as long as one eye; distance between clypeal lateral projection and apex of the adjacent anterior tooth longer than the basal width of the anterior tooth; acute angle between outer side of anterior teeth and clypeal lateral projection; distal maxillary palpomere, maximum width twice the apical width; fovea shallow, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and longer than the funicle in males, as long as in females.
<emphasis id="779A3362EC025A7B6FCC1D5AFA94FEA0" box="[1252,1342,259,285]" italics="true" pageId="7" pageNumber="8">Thorax:</emphasis>
anterior margin of pronotum straight; glabrous, punctures fine and very sparse, denser and coarser near the anterior margin; pronotal posterior corners in obtuse angle; hypomere with long bristles, scales near the inner margin; mesepisternum scaly; sides of metaventrite with brisltes and scales; distance between mesocoxae and metacoxae slightly longer than the length of the metacoxa; scutellum ogival, weakly punctate at the sides.
<emphasis id="779A3362EC025A7B68C31DCAFB90FE10" box="[1003,1082,403,429]" italics="true" pageId="7" pageNumber="8">Elytra:</emphasis>
shiny, glabrous, brown or dark brown; elytral suture and elytron unicolored, weakly elevated; all elytral ridges weakly defined.
<emphasis id="779A3362EC025A7B6FCE1DEEFA8EFE6D" box="[1254,1316,439,464]" italics="true" pageId="7" pageNumber="8">Legs:</emphasis>
three protibial teeth, the middle and apical equal in size, basal one slightly shorter; distance between basal and middle teeth slightly longer than between middle and apical; mesofemural surface setose; mesotibia subquadrate or quadrate in cross-section, surface coarsely sculptured; two transverse carinae, the apical one incomplete in males and complete in females; metacoxa scaly, scales long; basal apophysis of metacoxa produced; metatibia not carinate along the inner margin, apical inner surface setose; metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger longer than the diameter of the tibial apex; in males protarsomeres and mesotarsomeres I to IV strongly enlarged; metatarsomeres slightly enlarged; protarsomere II as long as it is wide; metatarsomere I two times shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw slightly longer than the inferior and equal in width, distance between teeth shorter than the inferior tooth.
<emphasis id="779A3362EC025A7B6F361EA2FB3BFCA8" box="[1054,1169,763,789]" italics="true" pageId="7" pageNumber="8">Abdomen:</emphasis>
ventrites with abundant bristles on disc; propygidium slightly visible, with bristles; pygidium more or less convex; subquadrate, as wide as it is long; pygidial width equal in distance or slightly wider than the distance between spiracles of propygidium; pygidial disc with sparse bristles throughout; pygidial apex quadrate.
<emphasis id="779A3362EC025A7B68831F3EFB8FFC3D" box="[939,1061,871,896]" italics="true" pageId="7" pageNumber="8">Parameres</emphasis>
: basal region narrower than both sections of the parameres at its maximum width, parameral split at 2/3, lyre shaped; ventral surface seen from dorsal view, concave; parameres convex in lateral view (
<figureCitation id="DDD5F3F5EC025A7B69EE1FF6FC8AFC74" box="[710,800,943,969]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="7" pageNumber="8">Fig. 2G</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="1191BFF8EC035A7A6BBF1BCEFD51F869" ID-DOI="http://doi.org/10.5281/zenodo.4561266" ID-Zenodo-Dep="4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1943,1967]" targetBox="[240,1385,181,1920]" targetPageId="6">
<paragraph id="4551EF70EC035A7A6BBF1BCEFD51F869" blockId="6.[151,1436,1943,2004]" pageId="6" pageNumber="7">
<emphasis id="779A3362EC035A7A6BBF1BCEFEB9F812" bold="true" box="[151,275,1943,1967]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
<taxonomicName id="82EE94F3EC035A7A6A311BC1FDCAF80D" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[281,608,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC035A7A6A311BC1FE47F80D" box="[281,493,1944,1968]" italics="true" pageId="6" pageNumber="7">Liogenys cartwrighti</emphasis>
<bibRefCitation id="217F9281EC035A7A6ADB1BC1FDCAF80D" author="Frey, G." box="[499,608,1944,1968]" pageId="6" pageNumber="7" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral.
</paragraph>
</caption>
<subSubSection id="0DF4BCFBEC025A7B6BEF1F8AFA85FBAC" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="4551EF70EC025A7B6BEF1F8AFD55FC50" blockId="7.[151,1437,151,1509]" box="[199,767,978,1005]" pageId="7" pageNumber="8">
<materialsCitation id="F586E52DEC025A7B6BEF1F8AFD51FC50" ID-GBIF-Occurrence="3047558372" box="[199,763,978,1005]" country="Argentina" location="Santiago del Estero" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Santiago del Estero" typeStatus="holotype">
<emphasis id="779A3362EC025A7B6BEF1F8AFECAFC51" bold="true" box="[199,352,978,1005]" pageId="7" pageNumber="8">
<typeStatus id="9A5551D2EC025A7B6BEF1F8AFEA8FC50" box="[199,258,979,1005]" pageId="7" pageNumber="8">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="3DF9AFE0EC025A7B6A441F8AFDB8FC50" box="[364,530,979,1005]" name="Argentina" pageId="7" pageNumber="8">ARGENTINA</collectingCountry>
,
<collectingRegion id="872A2192EC025A7B69361F8AFD51FC50" box="[542,763,979,1005]" country="Argentina" name="Santiago del Estero" pageId="7" pageNumber="8">Santiago del Estero</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC025A7B6BEF1FAFFA85FBAC" blockId="7.[151,1437,151,1509]" box="[199,1327,1014,1041]" pageId="7" pageNumber="8">
<emphasis id="779A3362EC025A7B6BEF1FAFFE54FBAD" bold="true" box="[199,510,1014,1040]" pageId="7" pageNumber="8">Geographical distribution</emphasis>
.
<materialsCitation id="F586E52DEC025A7B69241FAEFA81FBAC" ID-GBIF-Occurrence="3047558412" box="[524,1323,1014,1041]" country="Argentina" location="Jujuy" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Jujuy">
<collectingCountry id="3DF9AFE0EC025A7B69241FAEFD04FBAC" box="[524,686,1015,1041]" name="Argentina" pageId="7" pageNumber="8">ARGENTINA</collectingCountry>
(
<emphasis id="779A3362EC025A7B69961FAFFCA8FBAD" bold="true" box="[702,770,1014,1040]" pageId="7" pageNumber="8">
<collectingRegion id="872A2192EC025A7B69961FAFFCA8FBAD" box="[702,770,1014,1040]" country="Argentina" name="Jujuy" pageId="7" pageNumber="8">Jujuy</collectingRegion>
</emphasis>
,
<collectingRegion id="872A2192EC025A7B68381FAEFCE1FBAC" box="[784,843,1015,1041]" country="Argentina" name="Salta" pageId="7" pageNumber="8">Salta</collectingRegion>
,
<emphasis id="779A3362EC025A7B687E1FAEFC6AFBAC" bold="true" box="[854,960,1015,1041]" pageId="7" pageNumber="8">
<collectingRegion id="872A2192EC025A7B687E1FAEFC6AFBAC" box="[854,960,1015,1041]" country="Argentina" name="Formosa" pageId="7" pageNumber="8">Formosa</collectingRegion>
</emphasis>
,
<collectingRegion id="872A2192EC025A7B68E51FAEFB93FBAC" box="[973,1081,1015,1041]" country="Argentina" name="Tucuman" pageId="7" pageNumber="8">Tucumán</collectingRegion>
,
<collectingRegion id="872A2192EC025A7B6F6C1FAEFA89FBAC" box="[1092,1315,1015,1041]" country="Argentina" name="Santiago del Estero" pageId="7" pageNumber="8">Santiago del Estero</collectingRegion>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC025A7B6BEF1842FCB8FA58" pageId="7" pageNumber="8" type="discussion">
<paragraph id="4551EF70EC025A7B6BEF1842FE53FA20" blockId="7.[151,1437,151,1509]" pageId="7" pageNumber="8">
<emphasis id="779A3362EC025A7B6BEF1842FE91FB88" bold="true" box="[199,315,1051,1077]" pageId="7" pageNumber="8">Remarks.</emphasis>
<taxonomicName id="82EE94F3EC025A7B6A6B1842FD85FB88" authorityName="Frey" authorityYear="1969" box="[323,559,1051,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC025A7B6A6B1842FD85FB88" box="[323,559,1051,1077]" italics="true" pageId="7" pageNumber="8">Liogenys cartwrighti</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="82EE94F3EC025A7B69ED1842FBBAFB88" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[709,1040,1050,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="denticeps">
<emphasis id="779A3362EC025A7B69ED1842FCE4FB88" box="[709,846,1051,1077]" italics="true" pageId="7" pageNumber="8">L. denticeps</emphasis>
<bibRefCitation id="217F9281EC025A7B687E1842FBBAFB88" author="Blanchard, E." box="[854,1040,1050,1077]" pageId="7" pageNumber="8" pagination="129 - 240" refId="ref35099" refString="Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240." type="book chapter" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
(
<figureCitation id="DDD5F3F5EC025A7B6F091842FBCEFB88" box="[1057,1124,1050,1077]" captionStart="FIGURE 5" captionStartId="14.[151,250,1933,1957]" captionTargetBox="[151,1435,181,1910]" captionTargetId="figure-21@14.[151,1435,181,1910]" captionTargetPageId="14" captionText="FIGURE 5. Liogenys denticeps Blanchard, 1851 female lectotype views: C) dorsal, F) labels, I) pygidium; L. denticulata Moser, 1918 male lectotype: A) dorsal, D) labels, G) pygidium, J) lateral, and parameres views: K) dorsal and L) lateral; L. denticulata female paralectotype views: B) dorsal, E) labels." figureDoi="http://doi.org/10.5281/zenodo.4561274" httpUri="https://zenodo.org/record/4561274/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
) in the body size and color and in the clypeus and pygidium in shape.
<taxonomicName id="82EE94F3EC025A7B69AD1866FCDBFBE4" authorityName="Frey" authorityYear="1969" box="[645,881,1087,1113]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC025A7B69AD1866FCDBFBE4" box="[645,881,1087,1113]" italics="true" pageId="7" pageNumber="8">Liogenys cartwrighti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="82EE94F3EC025A7B6F231866FB3EFBE4" authorityName="Blanchard" authorityYear="1851" box="[1035,1172,1087,1113]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="denticeps">
<emphasis id="779A3362EC025A7B6F231866FB3EFBE4" box="[1035,1172,1087,1113]" italics="true" pageId="7" pageNumber="8">L. denticeps</emphasis>
</taxonomicName>
(in parenthesis) by the obtuse (rouded) pronotal posterior corners; pronotum and elytra finely punctate (coarsely); pygidium with bristles throughout (glabrous); males with protarsomeres and mesotarsomeres strongly enlarged (slightly wider than the metatarsomeres); and in the different shape of the parameres (
<figureCitation id="DDD5F3F5EC025A7B687218F2FC77FB78" box="[858,989,1195,1221]" captionStart="FIGURE 2" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[240,1385,181,1920]" captionTargetId="figure-21@6.[203,1385,181,1920]" captionTargetPageId="6" captionText="FIGURE 2. Liogenys cartwrighti Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) female pygidium, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561266" httpUri="https://zenodo.org/record/4561266/files/figure.png" pageId="7" pageNumber="8">Figs. 2F, G</figureCitation>
;
<figureCitation id="DDD5F3F5EC025A7B68C318F3FB98FB78" box="[1003,1074,1194,1221]" captionStart="FIGURE 5" captionStartId="14.[151,250,1933,1957]" captionTargetBox="[151,1435,181,1910]" captionTargetId="figure-21@14.[151,1435,181,1910]" captionTargetPageId="14" captionText="FIGURE 5. Liogenys denticeps Blanchard, 1851 female lectotype views: C) dorsal, F) labels, I) pygidium; L. denticulata Moser, 1918 male lectotype: A) dorsal, D) labels, G) pygidium, J) lateral, and parameres views: K) dorsal and L) lateral; L. denticulata female paralectotype views: B) dorsal, E) labels." figureDoi="http://doi.org/10.5281/zenodo.4561274" httpUri="https://zenodo.org/record/4561274/files/figure.png" pageId="7" pageNumber="8">5K, L</figureCitation>
). After examining the specimen at NHMB, from which
<bibRefCitation id="217F9281EC025A7B6A8B1896FD83FB57" author="Frey, G." box="[419,553,1231,1258]" pageId="7" pageNumber="8" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
had drawn the genitalia of
<taxonomicName id="82EE94F3EC025A7B684C1896FB3DFB54" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[868,1175,1231,1257]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="779A3362EC025A7B684C1896FBABFB54" box="[868,1025,1231,1257]" italics="true" pageId="7" pageNumber="8">L. denticulata</emphasis>
<bibRefCitation id="217F9281EC025A7B6F211896FB3DFB54" author="Moser, J." box="[1033,1175,1231,1257]" pageId="7" pageNumber="8" pagination="95 - 167" refId="ref36796" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">Moser, 1918</bibRefCitation>
</taxonomicName>
(
<emphasis id="779A3362EC025A7B6F811896FB4DFB55" box="[1193,1255,1231,1256]" italics="true" pageId="7" pageNumber="8">sensu</emphasis>
<bibRefCitation id="217F9281EC025A7B6FC71896FACDFB57" author="Frey, G." box="[1263,1383,1231,1258]" pageId="7" pageNumber="8" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey 1969</bibRefCitation>
), we realized that it is a misidentified specimen of
<taxonomicName id="82EE94F3EC025A7B69BA18AAFC84FAB0" authorityName="Frey" authorityYear="1969" box="[658,814,1267,1293]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC025A7B69BA18AAFC84FAB0" box="[658,814,1267,1293]" italics="true" pageId="7" pageNumber="8">L. cartwrighti</emphasis>
</taxonomicName>
.
<bibRefCitation id="217F9281EC025A7B681318AAFC6AFAB3" author="Frey, G." box="[827,960,1267,1294]" pageId="7" pageNumber="8" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
mentioned “
<collectingRegion id="872A2192EC025A7B6F6618AAFB21FAB0" box="[1102,1163,1267,1293]" country="Argentina" name="Salta" pageId="7" pageNumber="8">Salta</collectingRegion>
, Rivadavia” [
<collectingCountry id="3DF9AFE0EC025A7B6E0818ABFA3CFAB0" box="[1312,1430,1266,1293]" name="Argentina" pageId="7" pageNumber="8">Argentina</collectingCountry>
] as the type locality of the complete series, and that the type [
<typeStatus id="9A5551D2EC025A7B6867194FFC1DFA8C" box="[847,951,1302,1329]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
] is deposited at the USNM. The
<typeStatus id="9A5551D2EC025A7B6E19194EFA36FA8C" box="[1329,1436,1303,1329]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
examined at NHMB match with this locality, but the
<typeStatus id="9A5551D2EC025A7B69F51962FC95FAE8" box="[733,831,1339,1365]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
at the USNM bears “
<collectingRegion id="872A2192EC025A7B6F0F1962FAAEFAE8" box="[1063,1284,1339,1365]" country="Argentina" name="Santiago del Estero" pageId="7" pageNumber="8">Santiago del Estero</collectingRegion>
” [
<collectingCountry id="3DF9AFE0EC025A7B6E321963FA39FAE8" box="[1306,1427,1338,1365]" name="Argentina" pageId="7" pageNumber="8">Argentina</collectingCountry>
]. As this type specimen bears all the original labels of Frey, we confirm that
<collectingRegion id="872A2192EC025A7B68C11906FB6FFAC4" box="[1001,1221,1375,1401]" country="Argentina" name="Santiago del Estero" pageId="7" pageNumber="8">Santiago del Estero</collectingRegion>
is the type locality of
<taxonomicName id="82EE94F3EC025A7B6B9E19DAFEFBFA20" authorityName="Frey" authorityYear="1969" box="[182,337,1411,1437]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC025A7B6B9E19DAFEFBFA20" box="[182,337,1411,1437]" italics="true" pageId="7" pageNumber="8">L. cartwrighti</emphasis>
</taxonomicName>
, and not
<collectingRegion id="872A2192EC025A7B6A9119DAFE5FFA20" box="[441,501,1411,1437]" country="Argentina" name="Salta" pageId="7" pageNumber="8">Salta</collectingRegion>
.
</paragraph>
<paragraph id="4551EF70EC025A7B6BEF19FEFCB8FA58" blockId="7.[151,1437,151,1509]" pageId="7" pageNumber="8">
Besides the Chacoan region,
<taxonomicName id="82EE94F3EC025A7B692519FEFD02FA7C" authorityName="Frey" authorityYear="1969" box="[525,680,1447,1473]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cartwrighti">
<emphasis id="779A3362EC025A7B692519FEFD02FA7C" box="[525,680,1447,1473]" italics="true" pageId="7" pageNumber="8">L. cartwrighti</emphasis>
</taxonomicName>
can be also found at the Yungas Province, in sympatry with
<taxonomicName id="82EE94F3EC025A7B6E6519FEFF72FA59" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="denticeps">
<emphasis id="779A3362EC025A7B6E6519FEFF72FA59" italics="true" pageId="7" pageNumber="8">L. denticeps</emphasis>
</taxonomicName>
and their similar species
<taxonomicName id="82EE94F3EC025A7B6ADE1992FCA7FA58" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[502,781,1483,1509]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="latitarsis">
<emphasis id="779A3362EC025A7B6ADE1992FDDDFA58" box="[502,631,1483,1509]" italics="true" pageId="7" pageNumber="8">L. latitarsis</emphasis>
<bibRefCitation id="217F9281EC025A7B69561992FCA7FA58" author="Moser, J." box="[638,781,1483,1509]" pageId="7" pageNumber="8" pagination="95 - 167" refId="ref36796" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">Moser, 1918</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>