168 lines
20 KiB
XML
168 lines
20 KiB
XML
<document id="46CBCFFBF3F6B179F2AF60A9E8BF6D88" ID-DOI="10.11646/zootaxa.3925.2.2" ID-GBIF-Dataset="66d99618-4396-47bd-ae66-89bf89404d4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="232614" ID-ZooBank="1C440345-E190-41BF-977B-DCA5DFE705A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461001372614" checkinUser="plazi" docAuthor="Tóth, Balázs & Ronkay, László" docDate="2015" docId="CC285D33FFBD8B2CEDDAFB8E48DC8802" docLanguage="en" docName="zt03925p201.pdf" docOrigin="Zootaxa 3925 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naarda gigaloba Tóth & Ronkay, 2015, sp. n." docType="treatment" docUuid="37A320ED-2CFD-4CCA-84C2-755CEB2643A7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="192" masterDocId="3011254BFFB18B21ED4DFFC648338C7F" masterDocTitle="Revision of the Palaearctic and Oriental species of the genus Naarda Walker, 1866 (Lepidoptera: Erebidae, Hypeninae). Part 5. Description of 13 new species from Asia" masterLastPageNumber="201" masterPageNumber="179" pageNumber="191" updateTime="1698564773577" updateUser="plazi">
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<mods:mods id="C0334F22BF1E908ADF51A779AC546A34" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0F7206BDD70386CE49D5465DB3E75853">
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<mods:title id="D10C632922BD6F8D9E58BB7CFE497DFE">Revision of the Palaearctic and Oriental species of the genus Naarda Walker, 1866 (Lepidoptera: Erebidae, Hypeninae). Part 5. Description of 13 new species from Asia</mods:title>
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</mods:titleInfo>
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<mods:name id="E48A336D9EA089803CD1037B356B8D54" type="personal">
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<mods:roleTerm id="22FB207C699F289934AC537111C5A5E0">Author</mods:roleTerm>
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<mods:namePart id="4EF1ADA2986BF9F74BD7D600ACE07733">Tóth, Balázs</mods:namePart>
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</mods:name>
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<mods:name id="C15DA8A88C527CB06785A0C49E74495A" type="personal">
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<mods:roleTerm id="595EB02B4B61146212D68DB89CC6DF8B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58E79AFCF559B3A93F07C4BA07A226E0">Ronkay, László</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="9099FA4345400DCD18773BA37870DC4D">text</mods:typeOfResource>
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<mods:titleInfo id="A5DFBC27572DC610134941DBA7037D71">
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<mods:title id="8494C4315E96C5C1A49E7058C4027A3C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="27FA0E580ACCA7CD389D854B053013EB">
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<mods:date id="2997C674D7F5B47C210B358EF473F32B">2015</mods:date>
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<mods:detail id="1ADF9D5FD951E1B39B4C61B34EE13144" type="volume">
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<mods:number id="B0CD9DB165326A9D885ACD1DE142ADEA">3925</mods:number>
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</mods:detail>
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<mods:detail id="10533AE7D1400E08A580D5800F22524C" type="issue">
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<mods:number id="ED54FC73C1BDB89428250037B8A9E7AD">2</mods:number>
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<mods:start id="100322806FBA72CCDC0BD62EEF87F010">179</mods:start>
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<mods:classification id="9A8F434E11181F2A4824514A87C10E56">journal article</mods:classification>
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<mods:identifier id="9B781E7B7751352022533CF9C6ACED3B" type="DOI">10.11646/zootaxa.3925.2.2</mods:identifier>
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<mods:identifier id="AFABB85E68B02B00CB9E51CAEBF47368" type="GBIF-Dataset">66d99618-4396-47bd-ae66-89bf89404d4b</mods:identifier>
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<mods:identifier id="A587CE34CDEE2EB645315B90866D5075" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E518C07C3083D28DA09770005D7AF1A0" type="Zenodo-Dep">232614</mods:identifier>
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</mods:mods>
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<treatment id="CC285D33FFBD8B2CEDDAFB8E48DC8802" ID-DOI="http://doi.org/10.5281/zenodo.6110983" ID-GBIF-Taxon="119598208" ID-Zenodo-Dep="6110983" LSID="urn:lsid:zoobank.org:act:37A320ED-2CFD-4CCA-84C2-755CEB2643A7" httpUri="http://treatment.plazi.org/id/CC285D33FFBD8B2CEDDAFB8E48DC8802" lastPageId="13" lastPageNumber="192" pageId="12" pageNumber="191">
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<subSubSection id="0C9BBFAEFFBD8B2DEDDAFB8E49EC88FC" pageId="12" pageNumber="191" type="nomenclature">
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<paragraph id="443EEC25FFBD8B2DEDDAFB8E4998881D" blockId="12.[151,479,1096,1156]" box="[151,427,1096,1122]" pageId="12" pageNumber="191">
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<heading id="1F765B49FFBD8B2DEDDAFB8E4998881D" bold="true" box="[151,427,1096,1122]" fontSize="11" level="1" pageId="12" pageNumber="191" reason="1">
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<emphasis id="76F53037FFBD8B2DEDDAFB8E4998881D" bold="true" box="[151,427,1096,1122]" pageId="12" pageNumber="191">
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<taxonomicName id="838197A6FFBD8B2DEDDAFB8E4952881D" box="[151,353,1096,1122]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="gigaloba" status="sp. nov.">
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<emphasis id="76F53037FFBD8B2DEDDAFB8E4952881D" bold="true" box="[151,353,1096,1122]" italics="true" pageId="12" pageNumber="191">Naarda gigaloba</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6DC68D4CFFBD8B2DEC24FB8E4998881D" box="[361,427,1096,1122]" pageId="12" pageNumber="191" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="443EEC25FFBD8B2DEDDAFBAD49EC88FC" blockId="12.[151,479,1096,1156]" box="[151,479,1131,1156]" pageId="12" pageNumber="191">male: fig. 48, female: fig. 49.</paragraph>
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</subSubSection>
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<subSubSection id="0C9BBFAEFFBD8B2DEDDAFB754C948948" pageId="12" pageNumber="191" type="materials_examined">
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<paragraph id="443EEC25FFBD8B2DEDDAFB7549AC8890" blockId="12.[151,1437,1203,2025]" pageId="12" pageNumber="191">
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<emphasis id="76F53037FFBD8B2DEDDAFB7549F388B3" bold="true" box="[151,448,1203,1228]" pageId="12" pageNumber="191">
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<typeStatus id="9B3A5287FFBD8B2DEDDAFB7548E188B3" box="[151,210,1203,1228]" pageId="12" pageNumber="191">Type</typeStatus>
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material.
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<typeStatus id="9B3A5287FFBD8B2DEC1EFB7549F388B3" box="[339,448,1203,1228]" pageId="12" pageNumber="191" type="holotype">Holotype</typeStatus>
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</emphasis>
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. ♂: “SEPTEMBE[R]; MASKELIYA;
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<collectingCountry id="3C96ACB5FFBD8B2DEECCFB754BC588B4" box="[897,1014,1203,1227]" name="Sri Lanka" pageId="12" pageNumber="191">CEYLON</collectingCountry>
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; 1907.112 J.P.” slide No. BM Noct. 21798♂ (coll.
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<collectionCode id="229074E0FFBD8B2DEC77FB1149A08890" box="[314,403,1239,1263]" pageId="12" pageNumber="191">BMNH</collectionCode>
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).
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</paragraph>
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<paragraph id="443EEC25FFBD8B2DED8AFB3D4C948948" blockId="12.[151,1437,1203,2025]" pageId="12" pageNumber="191">
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<emphasis id="76F53037FFBD8B2DED8AFB3D4907896B" bold="true" box="[199,308,1275,1300]" pageId="12" pageNumber="191">
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<typeStatus id="9B3A5287FFBD8B2DED8AFB3D4907896B" box="[199,308,1275,1300]" pageId="12" pageNumber="191" type="paratype">Paratype</typeStatus>
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</emphasis>
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. ♀: “2986; Haputale;
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<collectingCountry id="3C96ACB5FFBD8B2DEF0AFB3D4A8F896C" box="[583,700,1275,1299]" name="Sri Lanka" pageId="12" pageNumber="191">CEYLON</collectingCountry>
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; OCTOBER; ex coll. G.C. Alston; Nearly simple, very short ciliation; Joicey Bequest. Brit. Mus. 1934–120”, slide No. BM Noct. 21799♀ (coll.
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<collectionCode id="229074E0FFBD8B2DE90EFAD94CAF8948" box="[1091,1180,1311,1335]" pageId="12" pageNumber="191">BMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="0C9BBFAEFFBD8B2DED8AFA854D2C8A28" pageId="12" pageNumber="191" type="diagnosis">
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<paragraph id="443EEC25FFBD8B2DED8AFA854D2C8A28" blockId="12.[151,1437,1203,2025]" pageId="12" pageNumber="191">
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<emphasis id="76F53037FFBD8B2DED8AFA8549098923" bold="true" box="[199,314,1347,1372]" pageId="12" pageNumber="191">Diagnosis</emphasis>
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. The relationship of
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<taxonomicName id="838197A6FFBD8B2DEF7DFA824A848924" box="[560,695,1347,1371]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="gigaloba">
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<emphasis id="76F53037FFBD8B2DEF7DFA824A848924" box="[560,695,1347,1371]" italics="true" pageId="12" pageNumber="191">N. gigaloba</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="838197A6FFBD8B2DEFB1FA824C4A8924" authority="Holloway, 2008" authorityName="Holloway" authorityYear="2008" box="[764,1145,1347,1372]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="kinabaluensis">
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<emphasis id="76F53037FFBD8B2DEFB1FA824B898924" box="[764,954,1347,1371]" italics="true" pageId="12" pageNumber="191">N. kinabaluensis</emphasis>
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<bibRefCitation id="201091D4FFBD8B2DEE89FA854C4A8924" author="Holloway" box="[964,1145,1347,1372]" pageId="12" pageNumber="191" refString="Holloway, J. D. (2008) The moths of Borneo: family Noctuidae, subfamilies Rivulinae, Phytometrinae, Herminiinae, Hypeninae, Hypenodinae. Malayan Nature Journal, 60 (1 - 4), 1 - 268." type="journal article" year="2008">Holloway, 2008</bibRefCitation>
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</taxonomicName>
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is supported by the long labial palps, the bipectinate male antenna with long rami, the shape of the forewing, the fragmented subterminal line, the similar reniform stigma and the similar structure of ductus bursae.
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<taxonomicName id="838197A6FFBD8B2DEE91FA4D4CA989DC" box="[988,1178,1419,1443]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="gigaloba">
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<emphasis id="76F53037FFBD8B2DEE91FA4D4CA989DC" box="[988,1178,1419,1443]" italics="true" pageId="12" pageNumber="191">Naarda gigaloba</emphasis>
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</taxonomicName>
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differs externally from
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<taxonomicName id="838197A6FFBD8B2DEDDAFA76496589B8" box="[151,342,1455,1479]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="kinabaluensis">
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<emphasis id="76F53037FFBD8B2DEDDAFA76496589B8" box="[151,342,1455,1479]" italics="true" pageId="12" pageNumber="191">N. kinabaluensis</emphasis>
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</taxonomicName>
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by the lack of brown shade in the ground colour of the wings, the lighter hindwing and the presence of the discal mark on the hindwing. The male genitalia of the new species have, in comparison with
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<taxonomicName id="838197A6FFBD8B2DE8CFFA1249028A70" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="kinabaluensis">
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<emphasis id="76F53037FFBD8B2DE8CFFA1249028A70" italics="true" pageId="12" pageNumber="191">N. kinabaluensis</emphasis>
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</taxonomicName>
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, much narrower uncus, narrower sacculus and much more pointed tip of valva and the huge lobe of
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<taxonomicName id="838197A6FFBD8B2DEDDAF9DA492C8A4C" box="[151,287,1563,1587]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="gigaloba">
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<emphasis id="76F53037FFBD8B2DEDDAF9DA492C8A4C" box="[151,287,1563,1587]" italics="true" pageId="12" pageNumber="191">N. gigaloba</emphasis>
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</taxonomicName>
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is absent from its close relative. In the female genitalia, the sterigma is present in
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<taxonomicName id="838197A6FFBD8B2DE9AAF9DA4D5C8A4C" box="[1255,1391,1563,1587]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="gigaloba">
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<emphasis id="76F53037FFBD8B2DE9AAF9DA4D5C8A4C" box="[1255,1391,1563,1587]" italics="true" pageId="12" pageNumber="191">N. gigaloba</emphasis>
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</taxonomicName>
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but absent from
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<taxonomicName id="838197A6FFBD8B2DEC6EF98649EE8A28" box="[291,477,1599,1623]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="kinabaluensis">
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<emphasis id="76F53037FFBD8B2DEC6EF98649EE8A28" box="[291,477,1599,1623]" italics="true" pageId="12" pageNumber="191">N. kinabaluensis</emphasis>
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</taxonomicName>
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, and the corpus bursae is larger in the new species than in the latter taxon.
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</paragraph>
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</subSubSection>
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<subSubSection id="0C9BBFAEFFBD8B2CED8AF9A54BE88FDA" lastPageId="13" lastPageNumber="192" pageId="12" pageNumber="191" type="description">
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<paragraph id="443EEC25FFBD8B2CED8AF9A54BE88FDA" blockId="12.[151,1437,1203,2025]" lastBlockId="13.[151,1437,151,1149]" lastPageId="13" lastPageNumber="192" pageId="12" pageNumber="191">
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<emphasis id="76F53037FFBD8B2DED8AF9A549628A03" bold="true" box="[199,337,1635,1660]" pageId="12" pageNumber="191">Description</emphasis>
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. Wingspan
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<quantity id="837941C0FFBD8B2DECAFF9A54A0A8A04" box="[482,569,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="191" unit="mm" value="21.0">21 mm</quantity>
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, length of forewing
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<quantity id="837941C0FFBD8B2DEE7FF9A54B4A8A04" box="[818,889,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="191" unit="mm" value="9.0">9 mm</quantity>
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(right forewing of female specimen missing). Antennae bipectinate in male, longest rami six times longer than diameter of flagellum, cilia twice as long as diameter of flagellum, present only on ventral third of rami; female having filiform and ciliate antennae with sparse cilia 2/3 as long as diameter of flagellum, setae apparently absent—the antennae of the female specimen are mouldy. Labial palps similar in both sexes, long: five times longer than diameter of eye; 3rd segment narrow, conspicuous, directed upwards, its apical half whitish, its tip pointed in male, rounded in female; 2nd segment broad, knife-shaped, abruptly widening then edges parallel. Scale-hood of vertex broad-based and moderately long in both sexes, rounded triangular in male, trapezoidal in female. Characteristic wing pattern features: forewing grey, slightly darker in female than in male; subterminal line fragmented to dots between veins in male, outer half of dots ash-grey, inner half black; this line continuous, slightly sinuous, grey in female; postmedial line in both sexes narrow, slightly darker than ground colour and parallel with subterminal (or its remnants); medial and antemedial lines as narrow as, but slightly more indistinct than postmedial; reniform stigma ovoid, dirty deep ochreous, with narrow black ring, a conspicuous black spot in its lower half (similarly to
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<taxonomicName id="838197A6FFBC8B2CEE81FF7B4CCA8CAB" authority="Sugi, 1982" authorityName="Sugi" authorityYear="1982" box="[972,1273,188,213]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="192" phylum="Arthropoda" rank="species" species="punctirena">
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<emphasis id="76F53037FFBC8B2CEE81FF7B4C568CAB" box="[972,1125,189,212]" italics="true" pageId="13" pageNumber="192">N. punctirena</emphasis>
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(Sugi, 1982)
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</taxonomicName>
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, and in female an additional small black dot in top third; orbicular stigma tiny, having the same colour as reniform in both sexes. Costa of forewing straight in male therefore the wing shape not differing from that of female. Hindwing off-white at costal third, remaining parts gradually approaching the greyish ground colour of forewing towards outer and inner edges; transverse lines indistinct, subterminal being marked with ash-grey dots in male; dark discal spot of underside translucent to upperside in both sexes.
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<emphasis id="76F53037FFBC8B2CEFF3FEA94B5A8DF7" bold="true" box="[702,873,367,392]" pageId="13" pageNumber="192">Male genitalia</emphasis>
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(fig. 13). Uncus moderately long, slightly curved, slightly dilated towards apex but apical quarter abruptly tapering to small, hooked structure, ventral edge of uncus with rounded emergence on its mid-third. Scaphium straight, as long as uncus. Tegumen as long as vinculum; saccus narrow, short, rounded. Juxta rounded, ca as long as wide, dorso-medially with rounded notch so the shape of juxta being similar to an apple. Valva broad-based, basal half slightly, apical half abruptly tapering, distal quarter of valva composed by the fused structure of sacculus, cucullus and costa. This section strongly sclerotised, dorsal edge curved, convex, with big teeth, ventral edge straight, apex extended to narrow, short and straight section (without teeth) so the shape of this whole fused structure reminiscent to the head of a duck in profile view. Sacculus narrow and long. Dorsal section of valva dominated by an enormous, long, rounded, dorsally directed, foot-shaped lobe; its rounded apical part having several long setae. This lobe extending from the base of valva beyond basal half of valva; originating probably by the fusion of transtilla and editum. Aedeagus thick, straight, vesica (uneverted) scobinate.
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<emphasis id="76F53037FFBC8B2CECD1FD3A4A508F6A" bold="true" box="[412,611,764,789]" pageId="13" pageNumber="192">Female genitalia</emphasis>
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(fig. 25). Ovipositor lobes angular. Apophyses posteriores 5/4 times as long as apophyses posteriores. Sternum A7 simple, not bilobed. Sterigma present: its posterior third section narrow, with slightly bilobed and granulose tip, remaining section of sterigma very broad, triangular, with some small, longitudinal creases. Antrum situated close to anterior edge of sterigma. Ductus bursae broad, medium long, somewhat spindle-like; corpus bursae big, ovoid, smooth, without signum.
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</paragraph>
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</subSubSection>
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<subSubSection id="0C9BBFAEFFBC8B2CED8AFC6949EE8FB7" box="[199,477,943,968]" pageId="13" pageNumber="192" type="distribution">
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<paragraph id="443EEC25FFBC8B2CED8AFC6949EE8FB7" blockId="13.[151,1437,151,1149]" box="[199,477,943,968]" pageId="13" pageNumber="192">
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<emphasis id="76F53037FFBC8B2CED8AFC69496A8FB7" bold="true" box="[199,345,943,968]" pageId="13" pageNumber="192">Distribution</emphasis>
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.
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<collectingCountry id="3C96ACB5FFBC8B2CEC2AFC6949EA8FB7" box="[359,473,943,968]" name="Sri Lanka" pageId="13" pageNumber="192">Sri Lanka</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="0C9BBFAEFFBC8B2CED8AFC1248DC8802" pageId="13" pageNumber="192" type="etymology">
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<paragraph id="443EEC25FFBC8B2CED8AFC124993886F" blockId="13.[151,1437,151,1149]" pageId="13" pageNumber="192">
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<emphasis id="76F53037FFBC8B2CED8AFC1249758F92" bold="true" box="[199,326,980,1005]" pageId="13" pageNumber="192">Etymology</emphasis>
|
||
. The specific name refers to the huge lobe on the valva in the male genitalia. The size of the lobe is the largest in the genus.
|
||
</paragraph>
|
||
<paragraph id="443EEC25FFBC8B2CED8AFBDA48DC8802" blockId="13.[151,1437,151,1149]" pageId="13" pageNumber="192">
|
||
<emphasis id="76F53037FFBC8B2CED8AFBDA4986884A" bold="true" box="[199,437,1052,1077]" pageId="13" pageNumber="192">Taxonomic remarks</emphasis>
|
||
. The shape of uncus is very simple, similar to those of
|
||
<taxonomicName id="838197A6FFBC8B2CE968FBDB4D78884A" authority="Walker, 1866" authorityName="Walker" authorityYear="1866" box="[1061,1355,1052,1077]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="192" phylum="Arthropoda" rank="species" species="bisignata">
|
||
<emphasis id="76F53037FFBC8B2CE968FBDB4C9D884B" box="[1061,1198,1052,1076]" italics="true" pageId="13" pageNumber="192">N. bisignata</emphasis>
|
||
<bibRefCitation id="201091D4FFBC8B2CE9FBFBDA4D78884A" author="Walker" box="[1206,1355,1052,1077]" pageId="13" pageNumber="192" refString="Walker, F. (1866) List of specimens of the lepidopterous insects of the Collection of the British Museum 35. Edward Newman, London, 506 pp." type="book" year="1866">Walker, 1866</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="838197A6FFBC8B2CE8CFFBDB49228827" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="192" phylum="Arthropoda" rank="species" species="blepharota">
|
||
<emphasis id="76F53037FFBC8B2CE8CFFBDB49228827" italics="true" pageId="13" pageNumber="192">N. blepharota</emphasis>
|
||
</taxonomicName>
|
||
. The shape of the apical fused structure of valva and the shape of the sterigma are apomorphies of this lineage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |