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<document id="AB1086B820277FEAB96B2627AD3A4DC3" ID-CLB-Dataset="54696" ID-DOI="10.11646/phytotaxa.597.1.4" ID-GBIF-Dataset="212b203f-b7ba-4244-9bdd-ced58149cb64" ID-ISSN="1179-3163" ID-Zenodo-Dep="7918770" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1683700350056" checkinUser="plazi" docAuthor="Yang, Jian-Jun, Parris, Barbara, Knapp, Ralf, Sundue, Michael, Zhang, Li-Bing &amp; Zhou, Xin-Mao" docDate="2023" docId="CA2C67019F44D93754C94B51C14FFEFF" docLanguage="en" docName="phytotaxa.597.1.4.pdf" docOrigin="Phytotaxa 597 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.597.1.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Calligrammitis Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp 2023, gen. nov." docType="treatment" docVersion="6" lastPageNumber="31" masterDocId="36151F799F46D93454744E29C351FFB7" masterDocTitle="Circumscription of the grammitid fern genus Oreogrammitis (Polypodiaceae) with the description of three new genera: Calligrammitis, Devolia, and Glabrigrammitis" masterLastPageNumber="36" masterPageNumber="28" pageNumber="30" updateTime="1699504062938" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D39865C695D212CAAE2953587FE11498">Circumscription of the grammitid fern genus Oreogrammitis (Polypodiaceae) with the description of three new genera: Calligrammitis, Devolia, and Glabrigrammitis</mods:title>
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<mods:namePart id="919A6FA11A8A1CE3C5CE5A8B1E8943C9">Yang, Jian-Jun</mods:namePart>
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<mods:affiliation id="66323CD800CFBBCD439106088DF4127F">School of Ecology and Environmental Science, Yunnan University, Kunming, Yunnan 650091, China &amp; yjj 2514 @ mail. ynu. edu. cn; https: // orcid. org / 0000 - 0002 - 4994 - 6345</mods:affiliation>
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<mods:namePart id="496B3C0976549150C59F8A93B47E3006">Parris, Barbara</mods:namePart>
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<mods:affiliation id="968E730F1863647ABBF80C1D0BD17ADC">Fern Research Foundation, 21 James Kemp Place, Kerikeri, Bay of Islands 0230, New Zealand &amp; barbara 2 parris @ gmail. com; https: // orcid. org / 0000 - 0002 - 0478 - 627 X</mods:affiliation>
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<mods:namePart id="2F3141A9771CB3CCC440AB274BA7F6E5">Knapp, Ralf</mods:namePart>
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<mods:affiliation id="D97408F6CD965BB66EB6E75CC3B185D4">Correspondent of the Muséum national dHistoire naturelle (MNHN, Paris, France), Steigestrasse 78, 69412 Eberbach, Germany &amp; ralf. knapp @ gmail. com; https: // orcid. org / 0000 - 0002 - 0147 - 5687</mods:affiliation>
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<mods:namePart id="F873AE3B921983A0F435BF6895AEBCBA">Sundue, Michael</mods:namePart>
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<mods:affiliation id="35C2F8C120AF90BD588C9A176397CFB2">The Pringle Herbarium, Department of Plant Biology, The University of Vermont, 27 Colchester Ave., Burlington, VT 05405, USA &amp; sundue @ gmail. com; https: // orcid. org / 0000 - 0003 - 1568 - 150 X</mods:affiliation>
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<mods:namePart id="D5DB3B06520F5DB87DDBE423A38478DA">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="4C38E017B687F13589929E3951295B05">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, MO 63110, USA.; Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu 610041, China &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<mods:affiliation id="53C907258FCE2D28E7A1A73BFB2FCDAE">School of Ecology and Environmental Science, Yunnan University, Kunming, Yunnan 650091, China &amp; xinmao. zhou @ ynu. edu. cn; https: // orcid. org / 0000 - 0003 - 3555 - 7784</mods:affiliation>
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<subSubSection id="0A9F859C9F44D93654C94B51C7F0FA25" box="[189,1185,1400,1427]" pageId="2" pageNumber="30" type="nomenclature">
<paragraph id="423AD6179F44D93654C94B51C7F0FA25" blockId="2.[136,1452,1364,2039]" box="[189,1185,1400,1427]" pageId="2" pageNumber="30">
<taxonomicName id="8585AD949F44D93654C94B51C775FA24" ID-CoL="BLRXX" authority="Parris, Sundue, Li Bing Zhang, X.M.Zhou &amp; Ralf Knapp" authorityName="Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp" authorityYear="2023" box="[189,1060,1400,1427]" class="Polypodiopsida" family="Polypodiaceae" genus="Calligrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="70F10A059F44D93654C94B51C220FA25" bold="true" box="[189,369,1400,1426]" pageId="2" pageNumber="30">Calligrammitis</emphasis>
Parris, Sundue, Li Bing Zhang, X.M.Zhou &amp; Ralf Knapp
</taxonomicName>
,
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<taxonomicNameLabel id="6BC2B77E9F44D93650414B51C7F0FA25" box="[1077,1185,1400,1426]" pageId="2" pageNumber="30" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="423AD6179F44D93650D84B50C128FA6D" blockId="2.[136,1452,1364,2039]" pageId="2" pageNumber="30">
<typeStatus id="9D3E68B59F44D93650D84B50C7B9FA24" box="[1196,1256,1401,1427]" pageId="2" pageNumber="30">Type</typeStatus>
:
<taxonomicName id="8585AD949F44D93650834B51C02AFA00" ID-CoL="6X3V9" authority="(Alderw.) Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp" authorityName="Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp" authorityYear="2007" baseAuthorityName="Alderwerelt" baseAuthorityYear="1918" class="Polypodiopsida" family="Polypodiaceae" genus="Radiogrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="species" species="beddomeana">
<emphasis id="70F10A059F44D93650834B51C6FAFA25" box="[1271,1451,1400,1426]" italics="true" pageId="2" pageNumber="30">C. beddomeana</emphasis>
(Alderw.) Parris, Sundue, Li Bing Zhang, X.M.Zhou &amp; Ralf Knapp
</taxonomicName>
<taxonomicName id="8585AD949F44D93657EC4BB5C26AFA6D" ID-CoL="6W5B6" authority="Alderw., Bull. Jard. Bot. Buitenzorg" class="Polypodiopsida" family="Polypodiaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="species" species="beddomeanum">
<emphasis id="70F10A059F44D93657EC4BB5C798FA01" box="[920,1225,1436,1462]" italics="true" pageId="2" pageNumber="30">Polypodium beddomeanum</emphasis>
Alderw., Bull. Jard. Bot. Buitenzorg
</taxonomicName>
sér. 2, 28: 39. 1918 (
<figureCitation id="DABECA929F44D93656534BE9C13DFA6D" box="[551,620,1472,1498]" captionStart="FIGURE 2" captionStartId="3.[136,234,1921,1943]" captionTargetBox="[264,1323,353,1896]" captionTargetId="figure-95@3.[264,1323,353,1896]" captionTargetPageId="3" captionText="FIGURE 2. Calligrammitis beddomeana (Alderw.) Parris, Sundue, Li Bing Zhang, X.M.Zhou &amp; Ralf Knapp.—A, B. Habit and habitats.—C. Lower portion of plant.—D. Adaxial view of lamina.—E. Portion of lamina margin showing hairs.—F. Young shoot of leaf showing hairs.—G. Stipes.—H. Abaxial view of laminae showing sori.—I. Abaxial view of portion of lamina showing sori (photo credit: Ralf Knapp)." figureDoi="http://doi.org/10.5281/zenodo.7918774" httpUri="https://zenodo.org/record/7918774/files/figure.png" pageId="2" pageNumber="30">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0A9F859C9F44D93654C94BCDC388F995" pageId="2" pageNumber="30" type="etymology">
<paragraph id="423AD6179F44D93654C94BCDC388F995" blockId="2.[136,1452,1364,2039]" pageId="2" pageNumber="30">
<emphasis id="70F10A059F44D93654C94BCDC269FA48" bold="true" box="[189,312,1508,1535]" italics="true" pageId="2" pageNumber="30">Etymology</emphasis>
—Greek,
<emphasis id="70F10A059F44D93655DD4BCDC28AFA49" box="[425,475,1508,1534]" italics="true" pageId="2" pageNumber="30">calli</emphasis>
= beautiful, and
<taxonomicName id="8585AD949F44D93656D14BCDC04CFA49" box="[677,797,1508,1534]" class="Polypodiopsida" family="Polypodiaceae" genus="Grammitis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D93656D14BCDC04CFA49" box="[677,797,1508,1534]" italics="true" pageId="2" pageNumber="30">Grammitis</emphasis>
</taxonomicName>
, a genus of ferns in which both species were previously placed.
</paragraph>
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<paragraph id="423AD6179F44D93654C94805C05CF961" blockId="2.[136,1452,1364,2039]" pageId="2" pageNumber="30">
<emphasis id="70F10A059F44D93654C94805C2A6F9F1" box="[189,503,1580,1607]" italics="true" pageId="2" pageNumber="30">
<emphasis id="70F10A059F44D93654C94805C261F9F1" bold="true" box="[189,304,1580,1606]" italics="true" pageId="2" pageNumber="30">Diagnosis</emphasis>
<taxonomicName id="8585AD949F44D93655394805C2A6F9F1" authorityName="Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp" authorityYear="2023" box="[333,503,1580,1606]" class="Polypodiopsida" family="Polypodiaceae" genus="Calligrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">Calligrammitis</taxonomicName>
</emphasis>
differs from other genera of Grammitidoideae by the following combination of characters: rhizomes radial, rhizome scales absent, stipes present, fronds not articulated to rhizome, laminae simple, without sclerotic margin, sometimes slightly crenulate, setae present, branched hairs with setae as branches present, basally stellate hairs present, veins simple, 1-forked or pinnately branched, hydathodes sometimes present at vein endings on adaxial surface of lamina, sporangia glabrous.
</paragraph>
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<subSubSection id="0A9F859C9F44D93654C948C9C754F841" pageId="2" pageNumber="30" type="discussion">
<paragraph id="423AD6179F44D93654C948C9C754F841" blockId="2.[136,1452,1364,2039]" pageId="2" pageNumber="30">
Species of
<taxonomicName id="8585AD949F44D936554948C9C2B6F94D" authorityName="Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp" authorityYear="2023" box="[317,487,1760,1786]" class="Polypodiopsida" family="Polypodiaceae" genus="Calligrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D936554948C9C2B6F94D" box="[317,487,1760,1786]" italics="true" pageId="2" pageNumber="30">Calligrammitis</emphasis>
</taxonomicName>
were placed in
<taxonomicName id="8585AD949F44D93656D548C9C0A5F94D" authority="(Parris 2007)" baseAuthorityName="Parris" baseAuthorityYear="2007" box="[673,1012,1760,1786]" class="Polypodiopsida" family="Polypodiaceae" genus="Radiogrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D93656D548C9C004F94D" box="[673,853,1760,1786]" italics="true" pageId="2" pageNumber="30">Radiogrammitis</emphasis>
(
<bibRefCitation id="2614ABE69F44D936571248C9C0BCF94D" author="Parris, B. S." box="[870,1005,1760,1786]" pageId="2" pageNumber="30" pagination="233 - 274" refId="ref4973" refString="Parris, B. S. (2007) Five new genera and three new species of Grammitidaceae (Filicales) and the re-establishment of Oreogrammitis. Gardens' Bulletin Singapore 58: 233 - 274." type="journal article" year="2007">Parris 2007</bibRefCitation>
)
</taxonomicName>
and
<emphasis id="70F10A059F44D936505248C9C77FF94D" box="[1062,1070,1760,1786]" italics="true" pageId="2" pageNumber="30">/</emphasis>
or
<taxonomicName id="8585AD949F44D936503A48C9C7A9F94D" authorityName="sensu Parris" authorityYear="2007" box="[1102,1272,1760,1786]" class="Polypodiopsida" family="Polypodiaceae" genus="Oreogrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D936503A48C9C7A9F94D" box="[1102,1272,1760,1786]" italics="true" pageId="2" pageNumber="30">Oreogrammitis</emphasis>
</taxonomicName>
(
<bibRefCitation id="2614ABE69F44D936517D48C9C392F8A9" author="Knapp, R. &amp; Hsu, T. C." pageId="2" pageNumber="30" refId="ref4640" refString="Knapp, R. &amp; Hsu, T. C. (2017) Ferns and Fern Allies of Taiwan: Second Supplement. KBCC Press, Gaoshu." type="book" year="2017">Knapp &amp; Hsu 2017</bibRefCitation>
,
<bibRefCitation id="2614ABE69F44D93654A4492CC2DBF8A9" author="Kuo, L. - Y. &amp; Hsu, T. - C. &amp; Chao, Y. - S. &amp; Liou, T. - W. &amp; Chang, H. - M. &amp; Chen, C. - W. &amp; Huang, Y. - M. &amp; Li, F. - W. &amp; Huang, Y. - F. &amp; Shao, W. &amp; Lu, P. - F. &amp; Chen, C. - W. &amp; Chang, Y. - H. &amp; Chiou, W. - L." box="[208,394,1796,1823]" pageId="2" pageNumber="30" pagination="367 - 395" refId="ref4669" refString="Kuo, L. - Y., Hsu, T. - C., Chao, Y. - S., Liou, T. - W., Chang, H. - M., Chen, C. - W., Huang, Y. - M., Li, F. - W., Huang, Y. - F., Shao, W., Lu, P. - F., Chen, C. - W., Chang, Y. - H. &amp; Chiou, W. - L. (2019) Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania 64: 367 - 395. https: // doi. org / 10.6165 / tai. 2019.64.367" type="journal article" year="2019">
Kuo
<emphasis id="70F10A059F44D936557E492CC215F8A9" box="[266,324,1796,1822]" italics="true" pageId="2" pageNumber="30">et al.</emphasis>
2019
</bibRefCitation>
). More recently
<bibRefCitation id="2614ABE69F44D936563C492DC00BF8A9" author="Parris, B. S. &amp; Sundue, M. A." box="[584,858,1796,1822]" pageId="2" pageNumber="30" pagination="45 - 54" refId="ref5200" refString="Parris, B. S. &amp; Sundue, M. A. (2020) The fern genus Oreogrammitis (Grammitidoideae: Polypodiaceae) re-defined. Phytotaxa 436: 45 - 54. https: // doi. org / 10.11646 / phytotaxa. 436.1.4" type="journal article" year="2020">Parris &amp; Sundue (2020)</bibRefCitation>
synonymized
<taxonomicName id="8585AD949F44D9365076492DC7E7F8A9" baseAuthorityName="Parris" baseAuthorityYear="2007" box="[1026,1206,1796,1822]" class="Polypodiopsida" family="Polypodiaceae" genus="Radiogrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D9365076492DC7E7F8A9" box="[1026,1206,1796,1822]" italics="true" pageId="2" pageNumber="30">Radiogrammitis</emphasis>
</taxonomicName>
with
<taxonomicName id="8585AD949F44D936508E492DC6F5F8A9" authorityName="sensu Parris" authorityYear="2007" box="[1274,1444,1796,1822]" class="Polypodiopsida" family="Polypodiaceae" genus="Oreogrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D936508E492DC6F5F8A9" box="[1274,1444,1796,1822]" italics="true" pageId="2" pageNumber="30">Oreogrammitis</emphasis>
</taxonomicName>
.
<taxonomicName id="8585AD949F44D93654FC4901C263F8F5" authorityName="Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp" authorityYear="2023" box="[136,306,1832,1858]" class="Polypodiopsida" family="Polypodiaceae" genus="Calligrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D93654FC4901C263F8F5" box="[136,306,1832,1858]" italics="true" pageId="2" pageNumber="30">Calligrammitis</emphasis>
</taxonomicName>
differs from similar species of
<taxonomicName id="8585AD949F44D93656D44901C01BF8F5" authorityName="sensu Parris" authorityYear="2007" box="[672,842,1832,1858]" class="Polypodiopsida" family="Polypodiaceae" genus="Oreogrammitis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D93656D44901C01BF8F5" box="[672,842,1832,1858]" italics="true" pageId="2" pageNumber="30">Oreogrammitis</emphasis>
</taxonomicName>
that lack rhizome scales and sporangial setae by the presence of basally stellate hairs. It is closely related to
<taxonomicName id="8585AD949F44D936569F4964C009F8D1" box="[747,856,1869,1894]" class="Polypodiopsida" family="Polypodiaceae" genus="Prosaptia" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D936569F4964C009F8D1" box="[747,856,1869,1894]" italics="true" pageId="2" pageNumber="30">Prosaptia</emphasis>
</taxonomicName>
, which also has branched hairs with setae as branches and lacks sporangial setae, but can be easily distinguished from it by having radial rhizomes without rhizome scales, and superficial sori on the abaxial surface of the lamina, while
<taxonomicName id="8585AD949F44D936576E49BCC0D6F819" box="[794,903,1941,1966]" class="Polypodiopsida" family="Polypodiaceae" genus="Prosaptia" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="70F10A059F44D936576E49BCC0D6F819" box="[794,903,1941,1966]" italics="true" pageId="2" pageNumber="30">Prosaptia</emphasis>
</taxonomicName>
has dorsiventral rhizomes with stipes articulated to prominent phyllopodia and clathrate ciliate rhizome scales, with sori sunken in marginal pouches or in pits on the abaxial surface of the lamina or superficial on the abaxial surface of the lamina.
</paragraph>
</subSubSection>
<subSubSection id="0A9F859C9F45D93754C94EB7C616FEB7" pageId="3" pageNumber="31" type="description">
<paragraph id="423AD6179F45D93754C94EB7C616FEB7" blockId="3.[136,1452,158,329]" pageId="3" pageNumber="31">
<emphasis id="70F10A059F45D93754C94EB7C212FF0F" bold="true" box="[189,323,158,184]" italics="true" pageId="3" pageNumber="31">Description</emphasis>
—Lamina narrowly lanceolate to narrowly oblanceolate, or narrowly oblanceolate to linear-oblanceolate, texture membranous to coriaceous, setae usually of two different lengths; veins visible or not in transmitted light. Sori ± circular to broadly elliptic, superficial, one or more rows on each side of the midrib.
</paragraph>
</subSubSection>
<subSubSection id="0A9F859C9F45D93754C94F22C14FFEFF" pageId="3" pageNumber="31" type="discussion">
<paragraph id="423AD6179F45D93754C94F22C14FFEFF" blockId="3.[136,1452,158,329]" pageId="3" pageNumber="31">
<emphasis id="70F10A059F45D93754C94F22C2ADFE93" box="[189,508,266,293]" italics="true" pageId="3" pageNumber="31">
<emphasis id="70F10A059F45D93754C94F22C264FE92" bold="true" box="[189,309,267,293]" italics="true" pageId="3" pageNumber="31">Taxonomy</emphasis>
<taxonomicName id="8585AD949F45D93755264F23C2ADFE93" authorityName="Parris, Sundue, Li Bing Zhang, X. M. Zhou &amp; Ralf Knapp" authorityYear="2023" box="[338,508,266,292]" class="Polypodiopsida" family="Polypodiaceae" genus="Calligrammitis" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="31" phylum="Tracheophyta" rank="genus">Calligrammitis</taxonomicName>
</emphasis>
contains two species in
<collectingCountry id="3A9296879F45D937577B4F23C0E1FE93" box="[783,944,266,292]" name="Taiwan Island" pageId="3" pageNumber="31">Taiwan Island</collectingCountry>
, Southeast Asia, New
<collectingCountry id="3A9296879F45D93750D94F23C650FE93" box="[1197,1281,266,292]" name="Guinea-Bissau" pageId="3" pageNumber="31">Guinea</collectingCountry>
, and
<collectingCountry id="3A9296879F45D93751494F23C6FAFE93" box="[1341,1451,266,292]" name="Sri Lanka" pageId="3" pageNumber="31">Sri Lanka</collectingCountry>
(
<bibRefCitation id="2614ABE69F45D93754E44F07C205FEFE" author="Parris, B. S. &amp; Chen, C. - W. &amp; Hsu, T. C. &amp; Lu, T. N. &amp; Nguyen, Q. D. &amp; Luu, H. T." box="[144,340,302,329]" pageId="3" pageNumber="31" refId="ref5123" refString="Parris, B. S., Chen, C. - W., Hsu, T. C., Lu, T. N., Nguyen, Q. D. &amp; Luu, H. T. (2015) New species and records of grammitid ferns (Polypodiaceae) for Vietnam. Phytotaxa 226: 039 - 050. https: // doi. org / 10.11646 / phytotaxa. 226.1.4" type="journal volume" year="2015">
Parris
<emphasis id="70F10A059F45D93754AE4F06C243FEFF" box="[218,274,302,328]" italics="true" pageId="3" pageNumber="31">et al.</emphasis>
2015
</bibRefCitation>
,
<bibRefCitation id="2614ABE69F45D937552B4F06C143FEFF" author="Kuo, L. - Y. &amp; Hsu, T. - C. &amp; Chao, Y. - S. &amp; Liou, T. - W. &amp; Chang, H. - M. &amp; Chen, C. - W. &amp; Huang, Y. - M. &amp; Li, F. - W. &amp; Huang, Y. - F. &amp; Shao, W. &amp; Lu, P. - F. &amp; Chen, C. - W. &amp; Chang, Y. - H. &amp; Chiou, W. - L." box="[351,530,302,329]" pageId="3" pageNumber="31" pagination="367 - 395" refId="ref4669" refString="Kuo, L. - Y., Hsu, T. - C., Chao, Y. - S., Liou, T. - W., Chang, H. - M., Chen, C. - W., Huang, Y. - M., Li, F. - W., Huang, Y. - F., Shao, W., Lu, P. - F., Chen, C. - W., Chang, Y. - H. &amp; Chiou, W. - L. (2019) Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania 64: 367 - 395. https: // doi. org / 10.6165 / tai. 2019.64.367" type="journal article" year="2019">
Kuo
<emphasis id="70F10A059F45D93755E24F06C29FFEFF" box="[406,462,302,328]" italics="true" pageId="3" pageNumber="31">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>