178 lines
20 KiB
XML
178 lines
20 KiB
XML
<document id="93CC67EE841722286EE771487E5CB415" ID-CLB-Dataset="26577" ID-DOI="10.11646/zootaxa.4613.3.10" ID-GBIF-Dataset="1a9205b6-b1c8-4f35-814f-15df27982b4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3239825" ID-ZooBank="B193E70D-FC46-44A5-97AA-14DB0AF30AA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559796425776" checkinUser="plazi" docAuthor="Ileperuma Arachchi, Ilesha S. & Benjamin, Suresh P." docDate="2019" docId="CA2C3762FFCCFFE122F6FBCCFE7FF879" docLanguage="en" docName="zootaxa.4613.3.10.pdf" docOrigin="Zootaxa 4613 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tarrocanus Simon 1895" docType="treatment" docVersion="8" lastPageNumber="588" masterDocId="36154F1AFFCDFFE02261FFDEFFF8FF8A" masterDocTitle="The crab spider genus Tarrocanus Simon, 1895 with notes on the genera Alcimochthes Simon, 1895 and Domatha Simon, 1895 (Araneae: Thomisidae)" masterLastPageNumber="593" masterPageNumber="587" pageNumber="588" updateTime="1698733077437" updateUser="ExternalLinkService">
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<mods:title id="1E76C45CC7BB0119CE6735AAC21FE234">The crab spider genus Tarrocanus Simon, 1895 with notes on the genera Alcimochthes Simon, 1895 and Domatha Simon, 1895 (Araneae: Thomisidae)</mods:title>
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<mods:namePart id="8B3E2B546888F25315DF880D05943C00">Ileperuma Arachchi, Ilesha S.</mods:namePart>
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<mods:namePart id="284E2D17A401C2927847F43F52111AEA">Benjamin, Suresh P.</mods:namePart>
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<mods:title id="5B0C03E43A08BFC2D881D688A5CE61B8">Zootaxa</mods:title>
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<mods:part id="82F9C07DC877CAD8AFFEC70EB34EFDBE">
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<mods:date id="95C5D9EFCC3A0BC0CB283A997A448D9B">2019</mods:date>
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<mods:number id="8C49D0BA63894CE6F367621847E93296">4613</mods:number>
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<treatment id="CA2C3762FFCCFFE122F6FBCCFE7FF879" ID-DOI="http://doi.org/10.5281/zenodo.5927738" ID-GBIF-Taxon="157422478" ID-Zenodo-Dep="5927738" LSID="urn:lsid:plazi:treatment:CA2C3762FFCCFFE122F6FBCCFE7FF879" httpUri="http://treatment.plazi.org/id/CA2C3762FFCCFFE122F6FBCCFE7FF879" lastPageNumber="588" pageId="1" pageNumber="588">
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<paragraph id="423A8674FFCCFFE122F6FBCCFDE2FBA6" blockId="1.[151,538,1042,1069]" box="[151,538,1042,1069]" pageId="1" pageNumber="588">
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<heading id="19723118FFCCFFE122F6FBCCFDE2FBA6" bold="true" box="[151,538,1042,1069]" fontSize="11" level="1" pageId="1" pageNumber="588" reason="1">
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<emphasis id="70F15A66FFCCFFE122F6FBCCFDE2FBA6" bold="true" box="[151,538,1042,1069]" pageId="1" pageNumber="588">
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Genus
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<taxonomicName id="8585FDF7FFCCFFE1228FFBCDFDE2FBA6" ID-CoL="645KQ" ID-ENA="635347" authority="Simon, 1895" authorityName="Simon" authorityYear="1895" box="[238,538,1042,1069]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
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<emphasis id="70F15A66FFCCFFE1228FFBCDFE81FBA7" bold="true" box="[238,377,1043,1069]" italics="true" pageId="1" pageNumber="588">Tarrocanus</emphasis>
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<bibRefCitation id="2614FB85FFCCFFE123E0FBCCFDE2FBA6" author="Simon, E." box="[385,538,1042,1068]" pageId="1" pageNumber="588" pagination="761 - 1084" refId="ref3704" refString="Simon, E. (1895) Histoire naturelle des araignees. Vol. 1. Roret, Paris, 324 pp. [pp. 761 - 1084]" type="book chapter" year="1895">Simon, 1895</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A9FD5FFFFCCFFE122F6FB89FCFBFBE5" box="[151,771,1110,1135]" pageId="1" pageNumber="588" type="materials_examined">
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<paragraph id="423A8674FFCCFFE122F6FB89FCFBFBE5" blockId="1.[151,771,1110,1135]" box="[151,771,1110,1135]" pageId="1" pageNumber="588">
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<emphasis id="70F15A66FFCCFFE122F6FB89FEEEFBE4" box="[151,278,1111,1134]" italics="true" pageId="1" pageNumber="588">
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<typeStatus id="9D3E38D6FFCCFFE122F6FB89FF3EFBE4" box="[151,198,1111,1134]" pageId="1" pageNumber="588">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="8585FDF7FFCCFFE12342FB89FD6FFBE5" authority="Simon, 1895: 979" authorityName="Simon" authorityPageNumber="979" authorityYear="1895" box="[291,663,1110,1135]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="capra">
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<emphasis id="70F15A66FFCCFFE12342FB89FE20FBE4" box="[291,472,1111,1134]" italics="true" pageId="1" pageNumber="588">Tarrocanus capra</emphasis>
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<bibRefCitation id="2614FB85FFCCFFE123BFFB88FD9BFBE5" author="Simon, E." box="[478,611,1110,1135]" pageId="1" pageNumber="588" pagination="761 - 1084" refId="ref3704" refString="Simon, E. (1895) Histoire naturelle des araignees. Vol. 1. Roret, Paris, 324 pp. [pp. 761 - 1084]" type="book chapter" year="1895">Simon, 1895</bibRefCitation>
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: 979
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</taxonomicName>
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: fig. 1047.
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</paragraph>
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</subSubSection>
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<subSubSection id="0A9FD5FFFFCCFFE122F6FB42FDF1FA05" pageId="1" pageNumber="588" type="diagnosis">
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<paragraph id="423A8674FFCCFFE122F6FB42FDF1FA05" blockId="1.[151,1437,1180,2035]" pageId="1" pageNumber="588">
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<emphasis id="70F15A66FFCCFFE122F6FB42FEEAFB3D" bold="true" box="[151,274,1180,1207]" pageId="1" pageNumber="588">Diagnosis.</emphasis>
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Species of
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<taxonomicName id="8585FDF7FFCCFFE123F6FB40FDEEFB3D" authorityName="Simon" authorityYear="1895" box="[407,534,1182,1207]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
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<emphasis id="70F15A66FFCCFFE123F6FB40FDEEFB3D" box="[407,534,1182,1207]" italics="true" pageId="1" pageNumber="588">Tarrocanus</emphasis>
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</taxonomicName>
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can be readily distinguished from other thomisid genera by the following characteristics: vertical and broad clypeus, elevated ocular area with slightly curved anterior row, enlarged prominent lateral eye tubercles (arrows in
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<figureCitation id="DABE9AF1FFCCFFE12061FB3BFD5AFB75" box="[512,674,1252,1279]" captionStart="FIGURES 1–9" captionStartId="4.[151,264,1854,1879]" captionTargetBox="[216,1367,197,1824]" captionTargetId="figure@4.[216,1370,197,1824]" captionTargetPageId="4" captionText="FIGURES 1–9. 1 Tarrocanus capra (male), Bowatenna Forest Reserve, dorsal view; 2 Tarrocanus jaffnaensis sp. nov (male), Mundathivu, Jaffna, dorsal view; 3 Domatha vivida (male), type specimen (MNHN 11376), dorsal view; 4 Tarrocanus capra (male), lateral view; 5 Tarrocanus jaffnaensis sp. nov (male), lateral view; 6–7 T.capra, left male palp (6 ventral, 7 retrolateral). 8–9 T. jaffnaensis sp. nov., left male palp (8 ventral, 9 retrolateral).Arrows point to the lateral eye tubercles. Scale bars: 0.2 mm (6–9), 1 mm (1–5)." httpUri="https://zenodo.org/record/3239827/files/figure.png" pageId="1" pageNumber="588">Figs 1, 2, 4, 5</figureCitation>
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), tubercles of PLE larger than that of the other eyes and are either modified with prominent triangular shaped tooth-like projections (in
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<taxonomicName id="8585FDF7FFCCFFE121CFFAD4FBF5FAA9" authorityName="Simon" authorityYear="1895" box="[942,1037,1290,1315]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="capra">
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<emphasis id="70F15A66FFCCFFE121CFFAD4FBF5FAA9" box="[942,1037,1290,1315]" italics="true" pageId="1" pageNumber="588">T. capra</emphasis>
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</taxonomicName>
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) or without such modifications (
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<taxonomicName id="8585FDF7FFCCFFE127E9FAD4FEE8FACD" authority="Ileperuma Arachchi & Benjamin, 2019" authorityName="Ileperuma Arachchi & Benjamin" authorityYear="2019" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="jaffnaensis" status="sp. nov.">
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<emphasis id="70F15A66FFCCFFE127E9FAD4FEE8FACD" italics="true" pageId="1" pageNumber="588">T. jaffnaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6BC2E71DFFCCFFE12379FAF3FE96FACD" box="[280,366,1325,1351]" pageId="1" pageNumber="588" rank="species">
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<emphasis id="70F15A66FFCCFFE12379FAF3FE96FACD" bold="true" box="[280,366,1325,1351]" pageId="1" pageNumber="588">sp. nov</emphasis>
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</taxonomicNameLabel>
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) and horseshoe shaped marking on the centre of prosoma starting from the base of PLE tubercles.
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<taxonomicName id="8585FDF7FFCCFFE122B3FA8CFEA9FAE1" authorityName="Simon" authorityYear="1895" box="[210,337,1362,1387]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
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<emphasis id="70F15A66FFCCFFE122B3FA8CFEA9FAE1" box="[210,337,1362,1387]" italics="true" pageId="1" pageNumber="588">Tarrocanus</emphasis>
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</taxonomicName>
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can be separated from
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<taxonomicName id="8585FDF7FFCCFFE1203FFA8FFD0DFAE1" authorityName="Simon" authorityYear="1885" box="[606,757,1361,1387]" class="Arachnida" family="Thomisidae" genus="Alcimochthes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
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<emphasis id="70F15A66FFCCFFE1203FFA8FFD0DFAE1" box="[606,757,1361,1387]" italics="true" pageId="1" pageNumber="588">Alcimochthes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8585FDF7FFCCFFE1214FFA8FFC6EFAE1" authorityName="Simon" authorityYear="1895" box="[814,918,1361,1387]" class="Arachnida" family="Thomisidae" genus="Domatha" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
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<emphasis id="70F15A66FFCCFFE1214FFA8FFC6EFAE1" box="[814,918,1361,1387]" italics="true" pageId="1" pageNumber="588">Domatha</emphasis>
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</taxonomicName>
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by the strongly recurved anterior margin and posteriorly broader opisthosoma.
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</paragraph>
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</subSubSection>
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<subSubSection id="0A9FD5FFFFCCFFE122A6FA46FC68F891" pageId="1" pageNumber="588" type="description">
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<paragraph id="423A8674FFCCFFE122A6FA46FBE9F97D" blockId="1.[151,1437,1180,2035]" pageId="1" pageNumber="588">
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<emphasis id="70F15A66FFCCFFE122A6FA46FEA1FA39" bold="true" box="[199,345,1432,1459]" pageId="1" pageNumber="588">Description.</emphasis>
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Small spiders, body size 2.5-3.0. Prosoma brown, laterals dark brown, clypeus, ocular area, middle of prosoma light brown with whitish hue, widest at the centre, horseshoe-shaped markings on the centre, outlined by whitish yellow lines (
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<figureCitation id="DABE9AF1FFCCFFE12397FA3FFDA7FA71" box="[502,607,1505,1531]" captionStart="FIGURES 1–9" captionStartId="4.[151,264,1854,1879]" captionTargetBox="[216,1367,197,1824]" captionTargetId="figure@4.[216,1370,197,1824]" captionTargetPageId="4" captionText="FIGURES 1–9. 1 Tarrocanus capra (male), Bowatenna Forest Reserve, dorsal view; 2 Tarrocanus jaffnaensis sp. nov (male), Mundathivu, Jaffna, dorsal view; 3 Domatha vivida (male), type specimen (MNHN 11376), dorsal view; 4 Tarrocanus capra (male), lateral view; 5 Tarrocanus jaffnaensis sp. nov (male), lateral view; 6–7 T.capra, left male palp (6 ventral, 7 retrolateral). 8–9 T. jaffnaensis sp. nov., left male palp (8 ventral, 9 retrolateral).Arrows point to the lateral eye tubercles. Scale bars: 0.2 mm (6–9), 1 mm (1–5)." httpUri="https://zenodo.org/record/3239827/files/figure.png" pageId="1" pageNumber="588">Figs 1, 2</figureCitation>
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), covered with sparsely distributed long setae and sub-erect weak setae. Opisthosoma lighter than prosoma, covered with numerous spots and patches of reddish brown, dark brown and greyish white, anterior half strongly recurved, bell-shaped, covered with sparsely distributed short setae. Eyes with enlarged, yellowish brown, partly jointed tubercles, both eye rows recurved. Eye formula: ALE>PLE>PME>AME, triangular shaped tooth-like projection on PLE (prominent only in
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<taxonomicName id="8585FDF7FFCCFFE1211FF9ACFC22F901" authorityName="Simon" authorityYear="1895" box="[894,986,1650,1675]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="capra">
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<emphasis id="70F15A66FFCCFFE1211FF9ACFC22F901" box="[894,986,1650,1675]" italics="true" pageId="1" pageNumber="588">T. capra</emphasis>
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</taxonomicName>
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). Clypeus vertical and broad, chelicerae yellowish brown, legs pale with parsley distributed spines and setae. Palp: tegulum simple and round with no apophyses, VTA with long stem and hook-shaped, embolus either moderately long and slender or short, curved and stout. RTA rudimentary, ITA may or may not be present. Females are unknown.
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</paragraph>
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<paragraph id="423A8674FFCCFFE122A6F8DEFC68F891" blockId="1.[151,1437,1180,2035]" box="[199,912,1792,1819]" pageId="1" pageNumber="588">
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<emphasis id="70F15A66FFCCFFE122A6F8DEFE9FF891" bold="true" box="[199,359,1792,1819]" pageId="1" pageNumber="588">Composition.</emphasis>
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Two species:
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<taxonomicName id="8585FDF7FFCCFFE12067F8DCFD9AF891" authorityName="Simon" authorityYear="1895" box="[518,610,1794,1819]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="capra">
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<emphasis id="70F15A66FFCCFFE12067F8DCFD9AF891" box="[518,610,1794,1819]" italics="true" pageId="1" pageNumber="588">T. capra</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8585FDF7FFCCFFE120F8F8DCFCD5F891" authority="Ileperuma Arachchi & Benjamin, 2019" authorityName="Ileperuma Arachchi & Benjamin" authorityYear="2019" box="[665,813,1793,1819]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="jaffnaensis" status="sp. nov.">
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<emphasis id="70F15A66FFCCFFE120F8F8DCFCD5F891" box="[665,813,1793,1819]" italics="true" pageId="1" pageNumber="588">T. jaffnaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6BC2E71DFFCCFFE12154F8DFFC68F891" box="[821,912,1793,1819]" pageId="1" pageNumber="588" rank="species">
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<emphasis id="70F15A66FFCCFFE12154F8DFFC71F891" bold="true" box="[821,905,1793,1819]" pageId="1" pageNumber="588">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A9FD5FFFFCCFFE122A6F8FAFCE1F8B5" box="[199,793,1828,1855]" pageId="1" pageNumber="588" type="distribution">
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<paragraph id="423A8674FFCCFFE122A6F8FAFCE1F8B5" blockId="1.[151,1437,1180,2035]" box="[199,793,1828,1855]" pageId="1" pageNumber="588">
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<emphasis id="70F15A66FFCCFFE122A6F8FAFE99F8B5" bold="true" box="[199,353,1828,1855]" pageId="1" pageNumber="588">Distribution.</emphasis>
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Currently known only from
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<collectingCountry id="3A92C6E4FFCCFFE120C2F8FBFCEDF8B5" box="[675,789,1829,1855]" name="Sri Lanka" pageId="1" pageNumber="588">Sri Lanka</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="0A9FD5FFFFCCFFE122A6F897FE7FF879" pageId="1" pageNumber="588" type="discussion">
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<paragraph id="423A8674FFCCFFE122A6F897FE7FF879" blockId="1.[151,1437,1180,2035]" pageId="1" pageNumber="588">
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<emphasis id="70F15A66FFCCFFE122A6F897FEC3F8E9" bold="true" box="[199,315,1865,1891]" pageId="1" pageNumber="588">Remarks.</emphasis>
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Close relationship of
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<taxonomicName id="8585FDF7FFCCFFE12024F894FD3CF8E9" authorityName="Simon" authorityYear="1895" box="[581,708,1866,1891]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70F15A66FFCCFFE12024F894FD3CF8E9" box="[581,708,1866,1891]" italics="true" pageId="1" pageNumber="588">Tarrocanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8585FDF7FFCCFFE12165F897FC63F8E9" authorityName="Simon" authorityYear="1885" box="[772,923,1865,1891]" class="Arachnida" family="Thomisidae" genus="Alcimochthes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70F15A66FFCCFFE12165F897FC63F8E9" box="[772,923,1865,1891]" italics="true" pageId="1" pageNumber="588">Alcimochthes</emphasis>
|
||
</taxonomicName>
|
||
is confirmed by molecular data (Ileperuma Arachchi & Benjamin, in press). The extent of the relatedness of
|
||
<taxonomicName id="8585FDF7FFCCFFE1211AF8B3FC1BF80D" authorityName="Simon" authorityYear="1895" box="[891,995,1901,1927]" class="Arachnida" family="Thomisidae" genus="Domatha" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70F15A66FFCCFFE1211AF8B3FC1BF80D" box="[891,995,1901,1927]" italics="true" pageId="1" pageNumber="588">Domatha</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="8585FDF7FFCCFFE12668F8B0FB70F80D" authorityName="Simon" authorityYear="1895" box="[1033,1160,1902,1927]" class="Arachnida" family="Thomisidae" genus="Tarrocanus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70F15A66FFCCFFE12668F8B0FB70F80D" box="[1033,1160,1902,1927]" italics="true" pageId="1" pageNumber="588">Tarrocanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8585FDF7FFCCFFE126A0F8B3FAA0F80D" authorityName="Simon" authorityYear="1885" box="[1217,1368,1901,1927]" class="Arachnida" family="Thomisidae" genus="Alcimochthes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70F15A66FFCCFFE126A0F8B3FAA0F80D" box="[1217,1368,1901,1927]" italics="true" pageId="1" pageNumber="588">Alcimochthes</emphasis>
|
||
</taxonomicName>
|
||
is yet unknown due to unavailability of molecular data for
|
||
<taxonomicName id="8585FDF7FFCCFFE12086F84FFCB7F821" authorityName="Simon" authorityYear="1895" box="[743,847,1937,1963]" class="Arachnida" family="Thomisidae" genus="Domatha" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="70F15A66FFCCFFE12086F84FFCB7F821" box="[743,847,1937,1963]" italics="true" pageId="1" pageNumber="588">Domatha</emphasis>
|
||
</taxonomicName>
|
||
. A well represented sample, including species of the three genera (both sexes) is essential to evaluate their relationships and to corroborate their generic boundaries in morphological terms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |