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<mods:title id="3D150E4AC634A192A02C1415D6804AE4">Revision of Dugastella nitida n. comb. (Crustacea: Decapoda: Atyidae), a freshwater shrimp from the Oligocene of Aix-en-Provence</mods:title>
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<mods:namePart id="7AE53C5855040299731927BB54CE04AE">Mazancourt, Valentin de</mods:namePart>
<mods:affiliation id="3A137D9B767D4A39C6F8851E915B7E8D">Unité Biologie des Organismes et Écosystèmes Aquatiques (UMR 8067 BOREA), CNRS, MNHN, Sorbonne Université, Université de Caen Normandie, IRD, Université des Antilles and Muséum national dHistoire naturelle, Département Adaptations du Vivant, case postale 26, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) valentin. demazancourt _ ext @ mnhn. fr (corresponding author)</mods:affiliation>
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<mods:namePart id="F132A78D74E8B97248EB9882B64FD80B">Audo, Denis</mods:namePart>
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<paragraph id="4174895A89433A32C2FEF8CBFADCA3B8" blockId="5.[961,1307,1916,1942]" box="[961,1307,1916,1942]" pageId="5" pageNumber="970">
<heading id="1A3C3E3689433A32C2FEF8CBFADCA3B8" box="[961,1307,1916,1942]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="970" reason="2">
Genus
<taxonomicName id="86CBF2D989433A32C531F8CBFADCA3B8" ID-CoL="8VJQC" authority="Bouvier, 1912" authorityName="Bouvier" authorityYear="1912" box="[1038,1307,1916,1942]" class="Malacostraca" family="Atyidae" genus="Dugastella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="970" phylum="Arthropoda" rank="genus">
<emphasis id="73BF554889433A32C531F8CBFBBCA3B8" box="[1038,1147,1916,1942]" italics="true" pageId="5" pageNumber="970">Dugastella</emphasis>
<bibRefCitation id="255AF4AB89433A32C5BDF8CBFADCA3B8" author="BOUVIER E. L." box="[1154,1307,1916,1942]" pageId="5" pageNumber="970" pagination="993 - 998" refId="ref4276" refString="BOUVIER E. L. 1912. - Dugastella marocana, crevette primitive nouvelle de la famille des Atyides. Comptes rendus hebdomadaires des Seances de l'Academie des sciences 155: 993 - 998. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 31089 / f 1065. item" type="journal article" year="1912">Bouvier, 1912</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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. —
<taxonomicName id="86CBF2D989433A32C2DFF800FAFBA3E1" authority="Bouvier, 1912" authorityName="Bouvier" authorityYear="1912" box="[992,1340,1975,1999]" class="Malacostraca" family="Atyidae" genus="Dugastella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="970" phylum="Arthropoda" rank="species" species="marocana">
<emphasis id="73BF554889433A32C2DFF800FB6FA3E1" box="[992,1192,1975,1999]" italics="true" pageId="5" pageNumber="970">Dugastella marocana</emphasis>
<bibRefCitation id="255AF4AB89433A32C590F800FAFBA3E1" author="BOUVIER E. L." box="[1199,1340,1975,1999]" pageId="5" pageNumber="970" pagination="993 - 998" refId="ref4276" refString="BOUVIER E. L. 1912. - Dugastella marocana, crevette primitive nouvelle de la famille des Atyides. Comptes rendus hebdomadaires des Seances de l'Academie des sciences 155: 993 - 998. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 31089 / f 1065. item" type="journal article" year="1912">Bouvier, 1912</bibRefCitation>
</taxonomicName>
(
<figureCitation id="D9F095DF89433A32C472F800FA40A3E1" box="[1357,1415,1975,1999]" captionStart="FIG" captionStartId="10.[132,143,1148,1165]" captionTargetBox="[132,1455,215,1105]" captionTargetId="figure-462@10.[698,1267,213,749]" captionTargetPageId="10" captionText="FIG. 8. — Specimens of Dugastella marocana Bouvier, 1912, MNHN-IU-2016-11857: A, B, whole body; C, D, detailed views of the rostrum; C, corresponds to specimen figured in A; D, corresponds to specimen figured in B. Scale bars: A, B, 3 mm; C, D, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14287354" httpUri="https://zenodo.org/record/14287354/files/figure.png" pageId="5" pageNumber="970">Fig. 8</figureCitation>
) by monotypy.
</paragraph>
</subSubSection>
<caption id="15B4D9D289403A31C1BBF9EEFDE6A2B9" ID-DOI="http://doi.org/10.5281/zenodo.14287346" ID-Zenodo-Dep="14287346" httpUri="https://zenodo.org/record/14287346/files/figure.png" pageId="6" pageNumber="971" startId="6.[132,143,1625,1642]" targetBox="[185,1437,194,1597]" targetPageId="6" targetType="figure">
<paragraph id="4174895A89403A31C1BBF9EEFDE6A2B9" blockId="6.[132,1457,1625,1687]" pageId="6" pageNumber="971">
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. 4. — Detailed views and interpretative drawings of mouthparts of
<taxonomicName id="86CBF2D989403A31C3FBF9EEFBE2A244" authority="(A. Milne-Edwards, 1879)" authorityName="Mazancourt &amp; Audo" authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1879" box="[708,1061,1625,1642]" class="Malacostraca" family="Atyidae" genus="Dugastella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="971" phylum="Arthropoda" rank="species" species="nitida" status="comb. nov.">
<emphasis id="73BF554889403A31C3FBF9EEFC8AA244" box="[708,845,1625,1642]" italics="true" pageId="6" pageNumber="971">Dugastella nitida</emphasis>
(A.
<bibRefCitation id="255AF4AB89403A31C253F9EEFBE7A244" author="MILNE-EDWARDS A." box="[876,1056,1625,1642]" pageId="6" pageNumber="971" pagination="77 - 78" refId="ref5218" refString="MILNE-EDWARDS A. 1879. - Description d'un Crustace fossile provenant des marnes d'Aix (Caridina nitida). Bulletin de la Societe philomathique de Paris 7 (3): 77 - 78. https: // www. biodiversitylibrary. org / page / 31660123" type="journal article" year="1879">Milne-Edwards,1879</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="688CE83389403A31C513F9EEFBBEA244" box="[1068,1145,1625,1642]" pageId="6" pageNumber="971" rank="species">n. comb.</taxonomicNameLabel>
:
<emphasis id="73BF554889403A31C541F9EEFB4CA244" bold="true" box="[1150,1163,1625,1642]" pageId="6" pageNumber="971">A</emphasis>
,
<emphasis id="73BF554889403A31C5AAF9EEFB65A244" bold="true" box="[1173,1186,1625,1642]" pageId="6" pageNumber="971">B</emphasis>
, mandible of MNHN.F.A90842a;
<emphasis id="73BF554889403A31C1BBF9D8FF55A2AE" bold="true" box="[132,146,1647,1664]" pageId="6" pageNumber="971">C</emphasis>
,
<emphasis id="73BF554889403A31C1A1F9D8FF6BA2AE" bold="true" box="[158,172,1647,1664]" pageId="6" pageNumber="971">D</emphasis>
, mandible of MNHN.F.A59324a;
<emphasis id="73BF554889403A31C0FAF9D8FE16A2AE" bold="true" box="[453,465,1647,1664]" pageId="6" pageNumber="971">E</emphasis>
,
<emphasis id="73BF554889403A31C0E2F9D8FE2FA2AE" bold="true" box="[477,488,1647,1664]" pageId="6" pageNumber="971">F</emphasis>
, first maxilla of MNHN.F.A90842a. Abbreviations:
<emphasis id="73BF554889403A31C2ACF9D8FC6AA2AE" bold="true" box="[915,941,1647,1664]" pageId="6" pageNumber="971">i.p.</emphasis>
, incisor process;
<emphasis id="73BF554889403A31C57AF9D8FBABA2AE" bold="true" box="[1093,1132,1647,1664]" pageId="6" pageNumber="971">m.p.</emphasis>
, molar process;
<emphasis id="73BF554889403A31C5C3F9D8FADBA2AE" bold="true" box="[1276,1308,1647,1664]" pageId="6" pageNumber="971">p.e.</emphasis>
, proximal endite;
<emphasis id="73BF554889403A31C1BBF931FF62A2B9" bold="true" box="[132,165,1670,1687]" pageId="6" pageNumber="971">d.e.</emphasis>
, distal endite;
<emphasis id="73BF554889403A31C020F931FE87A2B9" bold="true" box="[287,320,1670,1687]" pageId="6" pageNumber="971">plp.</emphasis>
, palp. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="09D1DAD189403A31C1BBF8B6FEF9A361" pageId="6" pageNumber="971" type="taxon_list">
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. —
<taxonomicName id="86CBF2D989403A31C052F8B7FDF3A336" authorityName="Bouvier" authorityYear="1912" box="[365,564,1792,1816]" class="Malacostraca" family="Atyidae" genus="Dugastella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="971" phylum="Arthropoda" rank="species" species="marocana">
<emphasis id="73BF554889403A31C052F8B7FDF3A336" box="[365,564,1792,1816]" italics="true" pageId="6" pageNumber="971">Dugastella marocana</emphasis>
</taxonomicName>
,
<taxonomicName id="86CBF2D989403A31C37DF8B7FEBAA31A" authority="(Ferrer Galdiano, 1924)" baseAuthorityName="Ferrer Galdiano" baseAuthorityYear="1924" class="Malacostraca" family="Atyidae" genus="Dugastella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="971" phylum="Arthropoda" rank="species" species="valentina">
<emphasis id="73BF554889403A31C37DF8B7FCC0A336" box="[578,775,1792,1816]" italics="true" pageId="6" pageNumber="971">Dugastella valentina</emphasis>
(
<bibRefCitation id="255AF4AB89403A31C1B4F8ABFEB2A31A" author="FERRER GALDIANO M." box="[139,373,1819,1844]" pageId="6" pageNumber="971" pagination="210 - 213" refId="ref4769" refString="FERRER GALDIANO M. 1924. - Una nueva especie del genero Atyaephira (Decap., Atyidae). Boletin de La Real Sociedad Espanola 24: 210 - 213." type="journal article" year="1924">Ferrer Galdiano, 1924</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="86CBF2D989403A31C0B3F8ACFF06A361" authority="(A. Milne-Edwards, 1879)" authorityName="Mazancourt &amp; Audo" authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1879" class="Malacostraca" family="Atyidae" genus="Dugastella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="971" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="73BF554889403A31C0B3F8ACFDF2A31D" box="[396,565,1819,1843]" italics="true" pageId="6" pageNumber="971">Dugastella nitida</emphasis>
(A.
<bibRefCitation id="255AF4AB89403A31C35BF8ACFF7DA361" author="MILNE-EDWARDS A." pageId="6" pageNumber="971" pagination="77 - 78" refId="ref5218" refString="MILNE-EDWARDS A. 1879. - Description d'un Crustace fossile provenant des marnes d'Aix (Caridina nitida). Bulletin de la Societe philomathique de Paris 7 (3): 77 - 78. https: // www. biodiversitylibrary. org / page / 31660123" type="journal article" year="1879">Milne-Edwards, 1879</bibRefCitation>
)
</taxonomicName>
†† n. comb.
</paragraph>
</subSubSection>
<subSubSection id="09D1DAD189403A31C1BBF8D0FA75A3C5" pageId="6" pageNumber="971" type="diagnosis">
<paragraph id="4174895A89403A31C1BBF8D0FA75A3C5" blockId="6.[130,776,1895,2027]" lastBlockId="6.[811,1458,1792,2027]" pageId="6" pageNumber="971">
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(
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<bibRefCitation id="255AF4AB89403A31C004F8D0FDC0A351" author="SKET B. &amp; ZAKsEK V." box="[315,519,1895,1919]" pageId="6" pageNumber="971" pagination="786 - 818" refId="ref5793" refString="SKET B. &amp; ZAKsEK V. 2009. - European cave shrimp species (Decapoda: Caridea: Atyidae), redefined after a phylogenetic study; redefinition of some taxa, a new genus and four new Troglocaris species. Zoological Journal of the Linnean Society 155: 786 - 818. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00473. x" type="book chapter" year="2009">Sket &amp; Zakšek 2009</bibRefCitation>
). — Paratyine genus with supraorbital and suborbital teeth, and with a pterygostomial carapace border jutting out (pterygostomial spine); dorsal rostral denticulation not continued posteriorly on carapace; maxilliped I exopodal lobe gradually narrowed distally, with terminal flagellum as long as width of lobe; pereiopods I-III or more with exopodites; mature male pereiopods III-IV not distally widened, dactylus not pectinate; pereiopod V dactylus comb-like; male pleopod I endopodite short, ribbon-shaped, with parallel side borders, part of distal border extended into comparatively wide appendix interna, which is approximately twice as long as it is wide; male pleopod II with sausage-shaped appendix masculina, which is set in the distal half with long straight spines (longer than appendix masculina diameter), moderately longer than appendix interna. All known species are epigean, with fully developed corneas.
</paragraph>
</subSubSection>
</treatment>
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