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<document id="A8926E0ECEDB312C2DD8B44927E3F360" ID-CLB-Dataset="68733" ID-DOI="10.5852/ejt.2022.823.1811" ID-GBIF-Dataset="5cb7c978-e62d-4576-b32d-2469745af89a" ID-ISSN="2118-9773" ID-Zenodo-Dep="6628057" ID-ZooBank="D5AC626B-4A10-4CD9-B77B-6D7EFE8D180E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1654780041203" checkinUser="felipe" docAuthor="Korn, Dieter &amp; Bockwinkel, Jürgen" docDate="2022" docId="C9186647421A8941FDF25E15FD1EC19B" docLanguage="en" docName="ejt-823_10-47.pdf" docOrigin="European Journal of Taxonomy 823" docSource="http://zoobank.org/d5ac626b-4a10-4cd9-b77b-6d7efe8d180e" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Acanthoclymenia Hyatt 1900" docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="35211E3F4207895FFF965B4FFFC4C04C" masterDocTitle="The early gephuroceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea)" masterLastPageNumber="47" masterPageNumber="10" pageNumber="39" updateTime="1699339239020" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="691F5782241F27971049CF96029AE158">The early gephuroceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea)</mods:title>
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<mods:namePart id="D6235F2FA9368EF6581751B7B6CB9189">Korn, Dieter</mods:namePart>
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<mods:affiliation id="DA8A290AD7570E39B3098B933532D9F7">Museum für Naturkunde, Leibniz-Institut für Evolutions-und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="B6E7D29D9852D6468CAD8624AF80397B" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier>
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<mods:namePart id="850D078F6B22C17157A4F29EA6812E1D">Bockwinkel, Jürgen</mods:namePart>
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<mods:affiliation id="D5DFEA7559660196F8F5931E9A3856AE">Dechant-Fein-Strasse 22, 51375 Leverkusen, Germany.</mods:affiliation>
<mods:nameIdentifier id="9F66277BD0685E77AC36619C3B99129A" type="email">jbockwinkel@t-online.de</mods:nameIdentifier>
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<mods:title id="E007E1C09359E5FC7443229F31177D21">European Journal of Taxonomy</mods:title>
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<mods:date id="A12B56355CBF055A5A709D5DDCA68400">2022</mods:date>
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<mods:number id="FF89713A4FA2ADE66B3B782442E12D7F">2022-06-08</mods:number>
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<mods:number id="90DDDF625F2C60A0D108F1AD6BA27F2E">823</mods:number>
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<treatment id="C9186647421A8941FDF25E15FD1EC19B" ID-DOI="http://doi.org/10.5281/zenodo.6629246" ID-GBIF-Taxon="195832115" ID-Zenodo-Dep="6629246" LSID="urn:lsid:plazi:treatment:C9186647421A8941FDF25E15FD1EC19B" httpUri="http://treatment.plazi.org/id/C9186647421A8941FDF25E15FD1EC19B" lastPageId="30" lastPageNumber="40" pageId="29" pageNumber="39">
<subSubSection id="09AB84DA421A8942FDF25E15FBDFC538" box="[612,1051,1369,1396]" pageId="29" pageNumber="39" type="nomenclature">
<paragraph id="410ED751421A8942FDF25E15FBDFC538" blockId="29.[612,1051,1369,1396]" box="[612,1051,1369,1396]" pageId="29" pageNumber="39">
<heading id="1A46603D421A8942FDF25E15FBDFC538" box="[612,1051,1369,1396]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="39" reason="2">
Genus
<taxonomicName id="86B1ACD2421A8942FD235E16FBDFC538" ID-CoL="M5B" authority="Hyatt, 1900" authorityName="Hyatt" authorityYear="1900" box="[693,1051,1369,1396]" class="Cephalopoda" family="Acanthoclymeniidae" genus="Acanthoclymenia" kingdom="Animalia" order="Ammonoidea" pageId="29" pageNumber="39" phylum="Mollusca" rank="genus">
<emphasis id="73C50B43421A8942FD235E16FC4DC538" bold="true" box="[693,905,1369,1396]" italics="true" pageId="29" pageNumber="39">Acanthoclymenia</emphasis>
Hyatt, 1900
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="09AB84DA421A8942FF755ED3FB9BC5AA" pageId="29" pageNumber="39" type="type_taxon">
<paragraph id="410ED751421A8942FF755ED3FEB9C5F9" blockId="29.[227,381,1435,1462]" box="[227,381,1435,1462]" pageId="29" pageNumber="39">
<heading id="1A46603D421A8942FF755ED3FEB9C5F9" bold="true" box="[227,381,1435,1462]" fontSize="11" level="3" pageId="29" pageNumber="39" reason="3">
<emphasis id="73C50B43421A8942FF755ED3FEB9C5F9" bold="true" box="[227,381,1435,1462]" pageId="29" pageNumber="39">
<typeStatus id="9E0A69F3421A8942FF755ED3FEE4C5FA" box="[227,288,1436,1462]" pageId="29" pageNumber="39">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="410ED751421A8942FF755E84FB9BC5AA" blockId="29.[227,1119,1483,1510]" box="[227,1119,1483,1510]" pageId="29" pageNumber="39">
<taxonomicName id="86B1ACD2421A8942FF755E84FC96C5A9" authority="Neapolitana Clarke, 1892" authorityName="Neapolitana Clarke" authorityYear="1892" box="[227,850,1483,1509]" class="Magnoliopsida" family="Rutaceae" genus="Clymenia" kingdom="Plantae" order="Sapindales" pageId="29" pageNumber="39" phylum="Tracheophyta" rank="subGenus" subGenus="Cyrtoclymenia">
<emphasis id="73C50B43421A8942FF755E84FE97C5A9" box="[227,339,1483,1509]" italics="true" pageId="29" pageNumber="39">Clymenia</emphasis>
(
<emphasis id="73C50B43421A8942FEF25E84FDD5C5A9" box="[356,529,1483,1509]" italics="true" pageId="29" pageNumber="39">Cyrtoclymenia</emphasis>
)
<emphasis id="73C50B43421A8942FDB45E84FD77C5A9" box="[546,691,1483,1509]" italics="true" pageId="29" pageNumber="39">Neapolitana</emphasis>
Clarke, 1892
</taxonomicName>
(original designation).
</paragraph>
</subSubSection>
<subSubSection id="09AB84DA421A8942FF755D43FA90C6D2" pageId="29" pageNumber="39" type="diagnosis">
<paragraph id="410ED751421A8942FF755D43FE98C66A" blockId="29.[227,348,1548,1574]" box="[227,348,1548,1574]" pageId="29" pageNumber="39">
<heading id="1A46603D421A8942FF755D43FE98C66A" bold="true" box="[227,348,1548,1574]" fontSize="11" level="3" pageId="29" pageNumber="39" reason="3">
<emphasis id="73C50B43421A8942FF755D43FE98C66A" bold="true" box="[227,348,1548,1574]" pageId="29" pageNumber="39">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="410ED751421A8942FF755D73FA90C6D2" blockId="29.[227,1436,1596,1700]" pageId="29" pageNumber="39">
Genus of the family
<taxonomicName id="86B1ACD2421A8942FE4B5D73FD14C61A" authorityName="Schindewolf" authorityYear="1955" box="[477,720,1596,1622]" class="Cephalopoda" family="Acanthoclymeniidae" kingdom="Animalia" order="Ammonoidea" pageId="29" pageNumber="39" phylum="Mollusca" rank="family">Acanthoclymeniidae</taxonomicName>
with extremely discoidal, subinvolute to subevolute conch; venter flattened or concave. Suture line with relatively narrow wide lobe, moderately high median saddle, V-shaped E
<subScript id="DD35D514421A8942FE555DD9FE08C6E8" attach="left" box="[451,460,1686,1700]" fontSize="6" pageId="29" pageNumber="39">2</subScript>
lobe, shallow and rounded lateral lobe. Suture line formula (E
<subScript id="DD35D514421A8942FB3E5DD9FB75C6E8" attach="left" box="[1192,1201,1686,1700]" fontSize="6" pageId="29" pageNumber="39">2</subScript>
E
<subScript id="DD35D514421A8942FB5C5DD9FB17C6E8" attach="left" box="[1226,1235,1686,1700]" fontSize="6" pageId="29" pageNumber="39">1</subScript>
E
<subScript id="DD35D514421A8942FB7A5DD9FB31C6E8" attach="left" box="[1260,1269,1686,1700]" fontSize="6" pageId="29" pageNumber="39">2</subScript>
) L: U I.
</paragraph>
</subSubSection>
<subSubSection id="09AB84DA421A8942FF755D8AFC15C71A" pageId="29" pageNumber="39" type="reference_group">
<paragraph id="410ED751421A8942FF755D8AFE6BC693" blockId="29.[227,431,1733,1759]" box="[227,431,1733,1759]" pageId="29" pageNumber="39">
<heading id="1A46603D421A8942FF755D8AFE6BC693" bold="true" box="[227,431,1733,1759]" fontSize="11" level="3" pageId="29" pageNumber="39" reason="3">
<emphasis id="73C50B43421A8942FF755D8AFE6BC693" bold="true" box="[227,431,1733,1759]" pageId="29" pageNumber="39">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="410ED751421A8942FF755DBBFC15C71A" blockId="29.[227,1436,1780,1878]" pageId="29" pageNumber="39">
<taxonomicName id="86B1ACD2421A8942FF755DBBFC9BC742" authority="Neapolitana Clarke, 1892" authorityName="Neapolitana Clarke" authorityYear="1892" box="[227,863,1780,1807]" class="Magnoliopsida" family="Rutaceae" genus="Clymenia" kingdom="Plantae" order="Sapindales" pageId="29" pageNumber="39" phylum="Tracheophyta" rank="subGenus" subGenus="Cyrtoclymenia">
<emphasis id="73C50B43421A8942FF755DBBFE97C742" box="[227,339,1780,1806]" italics="true" pageId="29" pageNumber="39">Clymenia</emphasis>
(
<emphasis id="73C50B43421A8942FEF15DBBFDD0C742" box="[359,532,1780,1806]" italics="true" pageId="29" pageNumber="39">Cyrtoclymenia</emphasis>
)
<emphasis id="73C50B43421A8942FDBE5DBBFD7DC742" box="[552,697,1780,1806]" italics="true" pageId="29" pageNumber="39">Neapolitana</emphasis>
Clarke, 1892
</taxonomicName>
, New York;
<taxonomicName id="86B1ACD2421A8942FC6C5DBAFE73C77F" authority="Sandberger &amp; Sandberger, 1851" authorityName="Sandberger &amp; Sandberger" authorityYear="1851" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" order="Ammonoidea" pageId="29" pageNumber="39" phylum="Mollusca" rank="species" species="forcipifer">
<emphasis id="73C50B43421A8942FC6C5DBAFB29C742" box="[1018,1261,1780,1807]" italics="true" pageId="29" pageNumber="39">Goniatites forcipifer</emphasis>
Sandberger &amp; Sandberger, 1851
</taxonomicName>
, Rhenish Mountains;
<taxonomicName id="86B1ACD2421A8942FD535C57FB46C77E" authority="Clarke, 1879" authorityName="Clarke" authorityYear="1879" box="[709,1154,1816,1842]" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" order="Ammonoidea" pageId="29" pageNumber="39" phylum="Mollusca" rank="species" species="genundewa">
<emphasis id="73C50B43421A8942FD535C57FC13C77E" box="[709,983,1816,1842]" italics="true" pageId="29" pageNumber="39">Goniatites Genundewa</emphasis>
Clarke, 1879
</taxonomicName>
, New York;
<taxonomicName id="86B1ACD2421A8942FAB45C57FD1CC71A" authority="Sandberger &amp; Sandberger, 1851" authorityName="Sandberger &amp; Sandberger" authorityYear="1851" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" order="Ammonoidea" pageId="29" pageNumber="39" phylum="Mollusca" rank="species" species="planorbis">
<emphasis id="73C50B43421A8942FAB45C57FE93C71A" italics="true" pageId="29" pageNumber="39">Goniatites Planorbis</emphasis>
Sandberger &amp; Sandberger, 1851
</taxonomicName>
, Rhenish Mountains.
</paragraph>
</subSubSection>
<subSubSection id="09AB84DA421A8941FF755C31FD1EC19B" lastPageId="30" lastPageNumber="40" pageId="29" pageNumber="39" type="discussion">
<paragraph id="410ED751421A8942FF755C31FE91C7D4" blockId="29.[227,341,1918,1944]" box="[227,341,1918,1944]" pageId="29" pageNumber="39">
<heading id="1A46603D421A8942FF755C31FE91C7D4" bold="true" box="[227,341,1918,1944]" fontSize="11" level="3" pageId="29" pageNumber="39" reason="3">
<emphasis id="73C50B43421A8942FF755C31FE91C7D4" bold="true" box="[227,341,1918,1944]" pageId="29" pageNumber="39">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="410ED751421A8942FF755CE2FC47C843" blockId="29.[227,1436,1965,2063]" pageId="29" pageNumber="39">
In the description of the
<typeStatus id="9E0A69F3421A8942FD9E5CE2FDFFC78B" box="[520,571,1965,1991]" pageId="29" pageNumber="39">type</typeStatus>
species as well as its assignment to a genus, it was erroneously assumed that it was a clymeniid; this error was corrected by
<bibRefCitation id="2520AAA0421A8942FCD65C9EFC25C7A7" author="House M. R." box="[832,993,2000,2027]" pageId="29" pageNumber="39" pagination="472 - 476" refId="ref17890" refString="House M. R. 1961. Acanthoclymenia, the supposed earliest Devonian clymenid, is a Manticoceras. Palaeontology 3: 472 - 476." type="journal article" year="1961">House (1961)</bibRefCitation>
and the species was classified among the gephuroceratids, although in the genus
<taxonomicName id="86B1ACD2421A8942FD4D5CBAFCB8C842" box="[731,892,2037,2062]" class="Cephalopoda" family="Gephuroceratidae" genus="Manticoceras" kingdom="Animalia" order="Ammonoidea" pageId="29" pageNumber="39" phylum="Mollusca" rank="genus">
<emphasis id="73C50B43421A8942FD4D5CBAFCB8C842" box="[731,892,2037,2062]" italics="true" pageId="29" pageNumber="39">Manticoceras</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="410ED75142198941FF755A60FD1EC19B" blockId="30.[226,1437,303,471]" pageId="30" pageNumber="40">
<taxonomicName id="86B1ACD242198941FF755A60FE6BC105" authorityName="Hyatt" authorityYear="1900" box="[227,431,303,329]" class="Cephalopoda" family="Acanthoclymeniidae" genus="Acanthoclymenia" kingdom="Animalia" order="Ammonoidea" pageId="30" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="73C50B4342198941FF755A60FE6BC105" box="[227,431,303,329]" italics="true" pageId="30" pageNumber="40">Acanthoclymenia</emphasis>
</taxonomicName>
is considered the stratigraphically earliest and morphologically simplest genus of the superfamily
<taxonomicName id="86B1ACD242198941FEEE5A1DFCEEC121" authority="(Korn &amp; Klug 2002)" baseAuthorityName="Korn &amp; Klug" baseAuthorityYear="2002" box="[376,810,338,365]" pageId="30" pageNumber="40" rank="superFamily" superFamily="Beloceratoidea">
Beloceratoidea (
<bibRefCitation id="2520AAA042198941FDAE5A1CFCE6C121" author="Korn D. &amp; Klug C." box="[568,802,338,365]" pageId="30" pageNumber="40" refId="ref18280" refString="Korn D. &amp; Klug C. 2002. Ammoneae Devonicae. Backhuys, Leiden." type="book" year="2002">Korn &amp; Klug 2002</bibRefCitation>
)
</taxonomicName>
. It still has a suture line of the early gephuroceratids (for example
<taxonomicName id="86B1ACD242198941FEE95A38FE3AC1DC" authorityName="Matern" authorityYear="1929" box="[383,510,375,400]" class="Cephalopoda" family="Ponticeratidae" genus="Ponticeras" kingdom="Animalia" order="Ammonoidea" pageId="30" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="73C50B4342198941FEE95A38FE3AC1DC" box="[383,510,375,400]" italics="true" pageId="30" pageNumber="40">Ponticeras</emphasis>
</taxonomicName>
), consisting of a divided external lobe and a lateral, umbilical and internal lobe each. The genus is distinguished from the ponticeratids by the concave venter; a character that persists in subsequent forms of the
<taxonomicName id="86B1ACD242198941FDB45AF2FD12C19B" baseAuthorityName="Korn &amp; Klug" baseAuthorityYear="2002" box="[546,726,445,471]" pageId="30" pageNumber="40" rank="superFamily" superFamily="Beloceratoidea">Beloceratoidea</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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