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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.573.7016" ID-GBIF-Dataset="28121693-576c-493f-b725-b93a41ebb514" ID-PMC="PMC4829927" ID-Pensoft-Pub="1313-2970-573-85" ID-PubMed="27110168" ID-ZBK="2AE04FDB4A0440ABB854FF4461C1C634" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-573-85" ModsDocOrigin="ZooKeys 573" ModsDocTitle="Further contributions to the Aleocharinae (Coleoptera, Staphylinidae) fauna of New Brunswick and Canada including descriptions of 27 new species" checkinTime="1458839696031" checkinUser="pensoft" docAuthor="Webster, Reginald P., Klimaszewski, Jan, Bourdon, Caroline, Sweeney, Jon D., Hughes, Cory C. &amp; Labrecque, Myriam" docDate="2016" docId="C6DE08777A6BE3AA524216A9A2851C32" docLanguage="en" docName="ZooKeys 573: 85-216" docOrigin="ZooKeys 573" docSource="http://dx.doi.org/10.3897/zookeys.573.7016" docTitle="Phloeopora gilbertae Klimaszewski &amp; Webster, sp. n." docType="treatment" docUuid="59A46EAB-8630-4A35-AA56-945C45A2CCD1" docUuidSource="ZooBank" docVersion="5" lastPageNumber="189" masterDocId="3E711C649E54FFC9FA58FFFCFF9C2A71" masterDocTitle="Further contributions to the Aleocharinae (Coleoptera, Staphylinidae) fauna of New Brunswick and Canada including descriptions of 27 new species" masterLastPageNumber="216" masterPageNumber="85" pageNumber="187" updateTime="1668162850657" updateUser="ExternalLinkService">
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<mods:title>Further contributions to the Aleocharinae (Coleoptera, Staphylinidae) fauna of New Brunswick and Canada including descriptions of 27 new species</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Webster, Reginald P.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Klimaszewski, Jan</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bourdon, Caroline</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sweeney, Jon D.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hughes, Cory C.</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Labrecque, Myriam</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>573</mods:number>
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<mods:start>85</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.573.7016</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.573.7016</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-573-85</mods:identifier>
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<mods:identifier type="ZooBank">2AE04FDB4A0440ABB854FF4461C1C634</mods:identifier>
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<treatment ID-GBIF-Taxon="127860381" LSID="urn:lsid:zoobank.org:act:59A46EAB-8630-4A35-AA56-945C45A2CCD1" httpUri="http://treatment.plazi.org/id/C6DE08777A6BE3AA524216A9A2851C32" lastPageId="104" lastPageNumber="189" pageId="102" pageNumber="187">
<subSubSection pageId="102" pageNumber="187" type="multiple">
<paragraph pageId="102" pageNumber="187">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
</subSubSection>
<subSubSection pageId="102" pageNumber="187" type="nomenclature">
<paragraph pageId="102" pageNumber="187">
<taxonomicName LSID="http://zoobank.org/59A46EAB-8630-4A35-AA56-945C45A2CCD1" authority="Klimaszewski &amp; Webster" class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora gilbertae" order="Coleoptera" pageId="102" pageNumber="187" phylum="Arthropoda" rank="species" species="gilbertae">Phloeopora gilbertae Klimaszewski &amp; Webster</taxonomicName>
<taxonomicNameLabel pageId="102" pageNumber="187">sp. n.</taxonomicNameLabel>
Figs 494-500
</paragraph>
</subSubSection>
<subSubSection lastPageId="103" lastPageNumber="188" pageId="102" pageNumber="187" type="holotype (male)">
<paragraph pageId="102" pageNumber="187">Holotype (male).</paragraph>
<paragraph lastPageId="103" lastPageNumber="188" pageId="102" pageNumber="187">
Canada, New Brunswick, Sunbury Co., Gilbert Island,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.877">45.8770°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.2954">66.2954°W</geoCoordinate>
. 18-28.V.2012, C. Alderson &amp; V. Webster // Hardwood forest, Lindgren
<pageBreakToken pageId="103" pageNumber="188" start="start">funnel</pageBreakToken>
trap in canopy of
<taxonomicName class="Magnoliopsida" family="Juglandaceae" genus="Juglans" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juglans cinerea" order="Fagales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="cinerea">Juglans cinerea</taxonomicName>
(LFC). Paratypes: Canada, New Brunswick, Gloucester Co., Bathurst, Daly Point Nature Preserve,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.6392">47.6392°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-65.6098">65.6098°W</geoCoordinate>
, 13-28.V.2015, C. Alderson &amp; V. Webster // Mixed forest, purple Lindgren funnel trap in canopy (1, AFC); same data but 25.VI-9.VII.2015, purple Lindgren funnel trap in canopy (1, AFC). Northumberland Co., ca. 1.5 km NW of Sevogle,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.0939">47.0939°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-65.8387">65.8387°W</geoCoordinate>
, 11-26.VI.2013, C. Alderson &amp; V. Webster //
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus tremuloides" order="Malpighiales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="tremuloides">Populus tremuloides</taxonomicName>
stand with a few conifers, Lindgren funnel trap in canopy of
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus tremuloides" order="Malpighiales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="tremuloides">Populus tremuloides</taxonomicName>
(1 sex undetermined, RWC); ca. 2.5 km NW of Sevogle,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.0876">47.0876°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-65.8613">65.8613°W</geoCoordinate>
, 26.VI-8.VII.2013, C. Alderson &amp; V. Webster //
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus banksiana" order="Pinales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="banksiana">Pinus banksiana</taxonomicName>
forest, Lindgren funnel trap (1 ♀, RWC). Queens Co., Cranberry Lake P.N.A.,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="46.1125">46.1125°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-65.6075">65.6075°W</geoCoordinate>
, 25.V-7.VI.2011, M. Roy &amp; V. Webster // Red oak forest, Lindgren funnel trap (1 ♂, RWC). Restigouche Co., Jacquet River Gorge P.N.A.,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.8257">47.8257°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.0764">66.0764°W</geoCoordinate>
, 29.V-10.VI.2014, C. Alderson &amp; V. Webster // Old
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus balsamifera" order="Malpighiales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="balsamifera">Populus balsamifera</taxonomicName>
stand near river, Lindgren funnel traps under trees (2 sex undetermined, AFC; 1 ♀, RWC). Sunbury Co., Gilbert Island,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.877">45.8770°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.2954">66.2954°W</geoCoordinate>
, 18-28.V.2012, C. Alderson, C. Hughes, &amp; V. Webster // Hardwood forest, Lindgren funnel trap 1 m high under
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia americana" order="Malvales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="americana">Tilia americana</taxonomicName>
(1 ♂, RWC); same data except 28.V-12.VI.2012 // Lindgren funnel trap in canopy of
<taxonomicName class="Magnoliopsida" family="Juglandaceae" genus="Juglans" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juglans cinerea" order="Fagales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="cinerea">Juglans cinerea</taxonomicName>
(1 sex undetermined, LFC; 2 ♂, 1 ♀, RWC); same data except 29.VI-11.VII.2012 (1 ♂, LFC; 1 ♂, RWC); same data except 20.VI-5.VII.2013 // Lindgren funnel trap in canopy of
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus tremuloides" order="Malpighiales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="species" species="tremuloides">Populus tremuloides</taxonomicName>
(1 ♂, RWC). York Co., 16 km W of Tracy, off Rt. 645,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.6855">45.6855°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.8847">66.8847°W</geoCoordinate>
, 18.V-2.VI.2010, R. Webster &amp; C. MacKay, coll. // Old red pine forest, Lindgren funnel trap (1 ♀, LFC); Fredericton, Odell Park,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.9539">45.9539°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.6666">66.6666°W</geoCoordinate>
, 2-15.V.2013, C. Alderson &amp; V. Webster // Hardwood stand, Lindgren funnel trap 1 m high under trees (1 ♂, RWC). Northwest Territories, vic. Inuvik, 5 km SE townsite,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="68.32881">68.32881°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-133.63556">133.63556°W</geoCoordinate>
, 17.VII-3.VIII.2001 mixed
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
-
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="103" pageNumber="188" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
forest // UHR ethanol funnel trap 1c, M. Gavel et al., collectors (1 ♂, LFC).
</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="188" type="etymology">
<paragraph pageId="103" pageNumber="188">Etymology.</paragraph>
<paragraph pageId="103" pageNumber="188">
This species is dedicated to
<normalizedToken originalValue="Amélie">Amelie</normalizedToken>
Gilbert (LFC), who dissected thousands of specimens of
<taxonomicName lsidName="" pageId="103" pageNumber="188" rank="subfamily" subfamily="Aleocharinae">Aleocharinae</taxonomicName>
for our projects.
</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="188" type="description">
<paragraph pageId="103" pageNumber="188">Description.</paragraph>
<paragraph pageId="103" pageNumber="188">
Body length 2.2-2.4 mm, narrowly elongate, subparallel, black except tarsi and antennae reddish brown (Fig. 494); integument strongly glossy, moderately punctate and pubescent, pubescence short and adhering to body; head subequal to pronotum in size, pubescence directed outward and posterolaterad from midline of disk; eyes large, slightly shorter than postocular area; antennal articles incrassate and articles
<normalizedToken originalValue="VX">V-X</normalizedToken>
moderately to strongly transverse; pronotum trapezoidal in shape, broadest subapically, narrower than elytra, pubescence directed almost straight posteriad; elytra elongate, with distinct narrowly rounded shoulders, pubescence directed straight and obliquely posteriad; abdomen subparallel with three basal tergites deeply impressed basally. Male. Median lobe of aedeagus with tubus long, narrow and curved ventrally (Fig. 495); tergite VIII truncate apically (Fig. 496); sternite VIII with apical margin obtusely triangularly produced in middle one-sixth (Fig. 497). Female. Tergite VIII broadly arcuate apically (Fig. 498); sternite VIIII with apical margin broadly triangularly produced, apex rounded (Fig. 499); spermatheca with capsule spherical, stem forming loose coil in apical half, sinuous and narrower basad (Fig. 500).
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="189" type="distribution">
<paragraph pageId="104" pageNumber="189">
<pageBreakToken pageId="104" pageNumber="189" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="104" pageNumber="189">This species is known from NB and the NT and is likely transcontinental in Canada.</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="189" type="natural history">
<paragraph pageId="104" pageNumber="189">Natural history.</paragraph>
<paragraph pageId="104" pageNumber="189">
All adults of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora gilbertae" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="gilbertae">Phloeopora gilbertae</taxonomicName>
from NB were captured in Lindgren funnel traps, most in hardwood forests. Specimens were captured in the canopy of a
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus tremuloides" order="Malpighiales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="species" species="tremuloides">Populus tremuloides</taxonomicName>
Michx. (trembling aspen) stand, in the canopy of
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus tremuloides" order="Malpighiales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="species" species="tremuloides">Populus tremuloides</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Juglandaceae" genus="Juglans" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juglans cinerea" order="Fagales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="species" species="cinerea">Juglans cinerea</taxonomicName>
L. (butternut) and under
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia americana" order="Malvales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="species" species="americana">Tilia americana</taxonomicName>
L. (American basswood) in a hardwood forest on an island in a river, in Lindgren traps under
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus balsamifera" order="Malpighiales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="species" species="balsamifera">Populus balsamifera</taxonomicName>
L. in a
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus balsamifera" order="Malpighiales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="species" species="balsamifera">Populus balsamifera</taxonomicName>
stand near a river, in a red oak stand with
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
, a hardwood stand, and in a
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus banksiana" order="Pinales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="species" species="banksiana">Pinus banksiana</taxonomicName>
forest. The specimen from the Northwest Territories was collected in a Lindgren funnel trap in a mixed
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
-
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="104" pageNumber="189" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
forest. Adults were collected during May, June, July, and August. Other members of this genus live in subcortical habitats (
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft Publishers, Sofia and Moscow" pageId="107" pageNumber="192" title="Aleocharine beetles (Coleoptera, Staphylinidae) of the province of Newfoundland and Labrador, Canada." year="2011">Klimaszewski et al. 2011</bibRefCitation>
,
<bibRefCitation author="Webster, RP" editor="Klimaszewski, J" journalOrPublisher="ZooKeys" pageId="108" pageNumber="193" pagination="83 - 118" title="New Staphylinidae (Coleoptera) records with new collection data from New Brunswick, and an addition to the fauna of Quebec, Canada: Aleocharinae." url="10.3897/zookeys.186.2655" volume="186" volumeTitle="Biosystematics and Ecology of Canadian Staphylinidae (Coleoptera) II." year="2012">Webster et al. 2012</bibRefCitation>
), and we presume this species lives in similar habitats.
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="189" type="comments">
<paragraph pageId="104" pageNumber="189">Comments.</paragraph>
<paragraph pageId="104" pageNumber="189">
This species may be separated from its Nearctic congeners by its body proportions, the uniformly black body color except for the appendages, and by the shape of the median lobe of the aedeagus in lateral view, and the spermatheca. It differs from the other two eastern Canadian species,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora oregona" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="oregona">Phloeopora oregona</taxonomicName>
Casey and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora canadensis" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="canadensis">Phloeopora canadensis</taxonomicName>
Klimaszewski and Langor by the black body (brown with darker head in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora canadensis" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="canadensis">Phloeopora canadensis</taxonomicName>
and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora oregona" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="oregona">Phloeopora oregona</taxonomicName>
), the elongate rather than transverse elytra, the apical part of the median lobe of the tubus straight in lateral view (strongly produced in the other two species), the spermathecal stem forming a shorter loop, and male tergite VIII with the apex more abruptly produced than that of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora canadensis" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="canadensis">Phloeopora canadensis</taxonomicName>
. For illustrations of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora canadensis" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="canadensis">Phloeopora canadensis</taxonomicName>
, see
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft Publishers, Sofia and Moscow" pageId="107" pageNumber="192" title="Aleocharine beetles (Coleoptera, Staphylinidae) of the province of Newfoundland and Labrador, Canada." year="2011">Klimaszewski et al. 2011</bibRefCitation>
.
</paragraph>
<caption pageId="104" pageNumber="189">
<paragraph pageId="104" pageNumber="189">
Figures 494-500.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Phloeopora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phloeopora gilbertae" order="Coleoptera" pageId="104" pageNumber="189" phylum="Arthropoda" rank="species" species="gilbertae">Phloeopora gilbertae</taxonomicName>
Klimaszewski &amp; Webster: 494 habitus in dorsal view 495 median lobe of aedeagus in lateral view 496 male tergite VIII 497 male sternite VIII 498 female tergite VIII 499 female sternite VIII 500 spermatheca. Scale bar of habitus = 1 mm; remaining scale bars = 0.2 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>