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<document id="18C90A4C8F1489457A0648DAB01B8599" ID-CLB-Dataset="45647" ID-DOI="10.11646/zootaxa.3802.4.6" ID-GBIF-Dataset="9a1086b4-468f-472e-9dc6-17b9e73d1e8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="249896" ID-ZooBank="4A83AAAE-6F0B-4173-A066-DBBCFBC3BDD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460427674668" checkinUser="plazi" docAuthor="Macgown, Joe A., Boudinot, Brendon, Deyrup, Mark &amp; Sorger, D. Magdalena" docDate="2014" docId="C66787A55B27FF839AE0F9E6FEF0FBF0" docLanguage="en" docName="zt03802p552.pdf" docOrigin="Zootaxa 3802 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Odontomachus clarus Roger 1861" docType="treatment" docVersion="17" lastPageNumber="533" masterDocId="3A5EFFDD5B29FF919A77FFBAFFEDFFC4" masterDocTitle="A review of the Nearctic Odontomachus (Hymenoptera: Formicidae: Ponerinae) with a treatment of the males" masterLastPageNumber="552" masterPageNumber="515" pageNumber="529" updateTime="1732493224424" updateUser="ExternalLinkService">
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<mods:title id="103DA8D4606ACB2ADC88E478750DE631">A review of the Nearctic Odontomachus (Hymenoptera: Formicidae: Ponerinae) with a treatment of the males</mods:title>
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<mods:namePart id="D5430EA02391B18F4DCB02043C0FD992">Macgown, Joe A.</mods:namePart>
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<mods:namePart id="B9CD022E5BECAB71015D8393D5297A32">Deyrup, Mark</mods:namePart>
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<mods:namePart id="961C62BFCD5032CB207117E5D1D5CF27">Sorger, D. Magdalena</mods:namePart>
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<mods:date id="37FFB3B63F041DB6E8AA6EACA7A76D16">2014</mods:date>
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<treatment id="C66787A55B27FF839AE0F9E6FEF0FBF0" ID-DOI="http://doi.org/10.5281/zenodo.6129410" ID-GBIF-Taxon="119429107" ID-Zenodo-Dep="6129410" LSID="urn:lsid:plazi:treatment:C66787A55B27FF839AE0F9E6FEF0FBF0" httpUri="http://treatment.plazi.org/id/C66787A55B27FF839AE0F9E6FEF0FBF0" lastPageId="18" lastPageNumber="533" pageId="14" pageNumber="529" scope_family="Formicidae" scope_order="Hymenoptera" scope_subFamily="Ponerinae">
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<heading id="153981DF5B27FF9F9AE0F9E6FDD1F9B2" bold="true" box="[151,572,1628,1654]" fontSize="11" level="1" pageId="14" pageNumber="529" reason="1">
<taxonomicName id="89CE4D305B27FF9F9AE0F9E6FDD1F9B2" ID-CoL="48QBK" authority="Roger, 1861" authorityName="Roger" authorityYear="1861" box="[151,572,1628,1654]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9AE0F9E6FDD1F9B2" bold="true" box="[151,572,1628,1654]" pageId="14" pageNumber="529">
<emphasis id="7CBAEAA15B27FF9F9AE0F9E6FE4CF9B2" bold="true" box="[151,417,1628,1654]" italics="true" pageId="14" pageNumber="529">Odontomachus clarus</emphasis>
<bibRefCitation id="2A5F4B425B27FF9F9BDFF9E6FDD1F9B2" author="Roger" box="[424,572,1628,1654]" pageId="14" pageNumber="529" refString="Roger, J. (1861) Die Ponera - artigen Ameisen (Schluss). Berliner Entomologische Zeitschrift, 5, 1 - 54." type="journal article" year="1861">Roger, 1861</bibRefCitation>
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</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F9C4FD8FF953" blockId="14.[151,610,1628,1687]" box="[151,610,1662,1687]" pageId="14" pageNumber="529">
(
<figureCitation id="D6F52A365B27FF9F9AE8F9C4FF03F953" box="[159,238,1662,1687]" captionStart="FIGURE 2" captionStartId="6.[151,249,1213,1235]" captionTargetBox="[152,1434,306,1191]" captionTargetId="figure@6.[152,1435,300,1192]" captionTargetPageId="6" captionText="FIGURE 2. Worker petioles, profile view: (A) O. ruginodis and (B) O. clarus." httpUri="https://zenodo.org/record/249898/files/figure.png" pageId="14" pageNumber="529">Figs. 2</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9A8DF9C5FEE7F953" box="[250,266,1663,1687]" captionStart="FIGURE 4" captionStartId="7.[151,249,1425,1447]" captionTargetBox="[151,1435,486,1403]" captionTargetId="figure@7.[151,1435,486,1403]" captionTargetPageId="7" captionText="FIGURE 4. First gastral tergites of workers, dorsal view: (A) O. brunneus and (B) O. clarus." httpUri="https://zenodo.org/record/249900/files/figure.png" pageId="14" pageNumber="529">4</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B61F9C4FEC8F953" box="[278,293,1662,1687]" captionStart="FIGURE 5" captionStartId="8.[151,249,1008,1030]" captionTargetBox="[211,1380,183,992]" captionTargetId="figure@8.[211,1380,183,993]" captionTargetPageId="8" captionText="FIGURE 5. Worker basalar lobes: (A) O. relictus, basalar lobe ovate, with striae (inset at bottom right with simple diagram showing strial pattern of basalar lobe) and (B) O. clarus, basalar lobe spheroid, mostly smooth (inset with simple diagram showing general lack of sculpture of basalar lobe)." httpUri="https://zenodo.org/record/249901/files/figure.png" pageId="14" pageNumber="529">5</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B45F9C4FEAFF953" box="[306,322,1662,1687]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="14" pageNumber="529">6</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B39F9C5FEB3F953" box="[334,350,1663,1687]" captionStart="FIGURE 9" captionStartId="10.[151,249,823,845]" captionTargetBox="[151,1435,324,784]" captionTargetId="figure@10.[151,1435,324,784]" captionTargetPageId="10" captionText="FIGURE 9. Male propodea (Pp), oblique view: (A) O. clarus (rectangular inset shows striate sculpture, arrow indicates lack of obvious propodeal carina) and (B) O. ruginodis (rectangular inset shows reticulated sculpture, arrows point to propodeal carina)." httpUri="https://zenodo.org/record/249905/files/figure.png" pageId="14" pageNumber="529">9</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B1DF9C5FE65F953" box="[362,392,1663,1687]" captionStart="FIGURE 10" captionStartId="10.[151,250,1657,1679]" captionTargetBox="[152,1435,1182,1627]" captionTargetId="figure@10.[152,1435,1182,1627]" captionTargetPageId="10" captionText="FIGURE 10. Male petiolar sterna (ps), profile view: (A) O. desertorum, left arrow indicates subrectangular subpetiolar process (spp), middle arrow indicates the angular posterior process of the petiolar sternum, and right arrow points to the depressed region behind the posterior angled process; and (B) O. clarus, left arrow indicates triangular subpetiolar process, middle arrow points to tubercles with setae, and right arrow indicates the lack of angular posterior process and accompanying shallow depression." httpUri="https://zenodo.org/record/249906/files/figure.png" pageId="14" pageNumber="529">10</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9BE3F9C5FE5FF953" box="[404,434,1663,1687]" captionStart="FIGURE 14" captionStartId="15.[151,249,1651,1673]" captionTargetBox="[151,1436,195,1629]" captionTargetId="figure@15.[151,1436,195,1629]" captionTargetPageId="15" captionText="FIGURE 14. Odontomachus clarus, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in lateral view, (E) mesosoma in dorsal view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence." httpUri="https://zenodo.org/record/249909/files/figure.png" pageId="14" pageNumber="529">14</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9BC9F9C4FE31F953" box="[446,476,1662,1687]" captionStart="FIGURE 15" captionStartId="16.[151,249,1908,1930]" captionTargetBox="[151,1436,409,1886]" captionTargetId="figure@16.[151,1436,409,1886]" captionTargetPageId="16" captionText="FIGURE 15. Odontomachus clarus, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view." httpUri="https://zenodo.org/record/249910/files/figure.png" pageId="14" pageNumber="529">15</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9B9FF9C5FDEBF953" box="[488,518,1663,1687]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="14" pageNumber="529">24</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F9865F9C4FDDCF953" box="[530,561,1662,1687]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A – F) Paramere and volsella, medial view; (G – L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="14" pageNumber="529">25</figureCitation>
,
<figureCitation id="D6F52A365B27FF9F984BF9C4FDB4F953" box="[572,601,1662,1687]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="14" pageNumber="529">26</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="6EDE119D5B27FF9F9AE0F97CFC51F8DE" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9AE0F97CFDF1F918" ID-CoL="48QBK" authority="Roger, 1861: 26" authorityName="Roger" authorityPageNumber="26" authorityYear="1861" box="[151,540,1734,1756]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9AE0F97CFE98F918" box="[151,373,1734,1756]" italics="true" pageId="14" pageNumber="529">Odontomachus clarus</emphasis>
<treatmentCitation id="CF6F10A25B27FF9F9B0CF97DFDF1F918" author="Roger" box="[379,540,1734,1756]" page="26" pageId="14" pageNumber="529" year="1861">
<bibRefCitation id="2A5F4B425B27FF9F9B0CF97DFE17F918" author="Roger" box="[379,506,1734,1756]" pageId="14" pageNumber="529" refString="Roger, J. (1861) Die Ponera - artigen Ameisen (Schluss). Berliner Entomologische Zeitschrift, 5, 1 - 54." type="journal article" year="1861">Roger, 1861</bibRefCitation>
: 26
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(worker) Texas,
<collectingCountry id="36D976235B27FF9F98BFF97CFD13F918" box="[712,766,1734,1756]" name="United States of America" pageId="14" pageNumber="529">USA</collectingCountry>
;
<treatmentCitation id="CF6F10A25B27FF9F9970F97DFC57F918" author="Patton" box="[775,954,1734,1756]" page="618" pageId="14" pageNumber="529" year="1894">
<bibRefCitation id="2A5F4B425B27FF9F9970F97DFC67F918" author="Patton" box="[775,906,1734,1756]" pageId="14" pageNumber="529" refString="Patton, W. H. (1894) Habits of the leaping-ant of southern Georgia. American Naturalist, 28, 618 - 619." type="journal article" year="1894">Patton, 1894</bibRefCitation>
: 618
</treatmentCitation>
(male);
<treatmentCitation id="CF6F10A25B27FF9F9E78F97CFB38F918" author="Wheeler" box="[1039,1237,1734,1756]" page="407" pageId="14" pageNumber="529" year="1908">
<bibRefCitation id="2A5F4B425B27FF9F9E78F97CFB4BF918" author="Wheeler" box="[1039,1190,1734,1756]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1908) The ants of Texas, New Mexico and Arizona. (Part 1.). Bulletin of the American Museum of Natural History, 24, 399 - 485." type="journal article" year="1908">Wheeler, 1908</bibRefCitation>
: 407
</treatmentCitation>
(queen);
<treatmentCitation id="CF6F10A25B27FF9F9F42F97CFE66F93F" author="Petralia" page="381" pageId="14" pageNumber="529" year="1980">
<bibRefCitation id="2A5F4B425B27FF9F9F42F97CFEBBF93F" author="Petralia" pageId="14" pageNumber="529" refString="Petralia, R. S. &amp; Vinson, S. B. (1980 [&quot; 1979 &quot;]) Comparative anatomy of the ventral region of ant larvae, and its relation to feeding behavior. Psyche, 86, 375 - 394. [Cambridge]" type="journal article" year="1980">Petralia &amp; Vinson, 1980</bibRefCitation>
: 381
</treatmentCitation>
(larva). Subspecies of
<taxonomicName id="89CE4D305B27FF9F98F0F95CFC1AF93F" ID-CoL="48QBS" authority="Emery, 1911: 115" authorityName="Emery" authorityPageNumber="115" authorityYear="1911" box="[647,1015,1765,1787]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B27FF9F98F0F95CFCC8F93F" box="[647,805,1765,1787]" italics="true" pageId="14" pageNumber="529">O. haematodus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9940F95CFC1AF93F" author="Emery" box="[823,1015,1765,1787]" page="115" pageId="14" pageNumber="529" year="1911">
<bibRefCitation id="2A5F4B425B27FF9F9940F95CFC2DF93F" author="Emery" box="[823,960,1765,1787]" pageId="14" pageNumber="529" refString="Emery, C. (1911) Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum, 118, 1 - 125." type="journal article" year="1911">Emery, 1911</bibRefCitation>
: 115
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B27FF9F9E71F95FFB23F93F" author="Wheeler" box="[1030,1230,1765,1787]" page="40" pageId="14" pageNumber="529" year="1914">
<bibRefCitation id="2A5F4B425B27FF9F9E71F95FFB4FF93F" author="Wheeler" box="[1030,1186,1765,1787]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1914) Ants collected by W. M. Mann in the state of Hidalgo, Mexico. Journal of the New York Entomological Society, 22, 37 - 61." type="journal article" year="1914">Wheeler, 1914</bibRefCitation>
: 40
</treatmentCitation>
;
<treatmentCitation id="CF6F10A25B27FF9F9EAAF95FFA74F93F" author="Smith" box="[1245,1433,1765,1787]" page="129" pageId="14" pageNumber="529" year="1939">
<bibRefCitation id="2A5F4B425B27FF9F9EAAF95FFA8CF93F" author="Smith" box="[1245,1377,1765,1787]" pageId="14" pageNumber="529" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">Smith, 1939</bibRefCitation>
: 129
</treatmentCitation>
. Revived status as species:
<treatmentCitation id="CF6F10A25B27FF9F9BA4F8BEFD1EF8DE" author="Taylor" box="[467,755,1796,1818]" page="142" pageId="14" pageNumber="529" year="1962">
<bibRefCitation id="2A5F4B425B27FF9F9BA4F8BEFD2DF8DE" author="Taylor" box="[467,704,1796,1818]" pageId="14" pageNumber="529" refString="Taylor, R. W. &amp; Wilson, E. O. (1962 [&quot; 1961 &quot;]) Ants from three remote oceanic islands. Psyche 68, 137 - 144. http: // dx. doi. org / 10.1155 / 1961 / 87982" type="journal article" year="1962">Taylor &amp; Wilson, 1962</bibRefCitation>
: 142
</treatmentCitation>
;
<treatmentCitation id="CF6F10A25B27FF9F988AF8BFFC55F8DE" author="Brown" box="[765,952,1796,1818]" page="136" pageId="14" pageNumber="529" year="1976">
<bibRefCitation id="2A5F4B425B27FF9F988AF8BFFC68F8DE" author="Brown" box="[765,901,1796,1818]" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 136
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F898FA71F8FC" blockId="14.[151,1436,1734,1848]" box="[151,1436,1826,1848]" pageId="14" pageNumber="529">
<treatmentCitationGroup id="6EDE119D5B27FF9F9AE0F898FA71F8FC" box="[151,1436,1826,1848]" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9AE0F898FDABF8FC" ID-CoL="CMB53" authority="Buckley, 1867: 355" authorityName="Buckley" authorityPageNumber="355" authorityYear="1867" box="[151,582,1826,1848]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="7CBAEAA15B27FF9F9AE0F898FE97F8FC" box="[151,378,1826,1848]" italics="true" pageId="14" pageNumber="529">Odontomachus texana</emphasis>
Buckley, 1867: 355
</taxonomicName>
. Junior synonym of
<taxonomicName id="89CE4D305B27FF9F9965F899FBA1F8FC" ID-CoL="48QBW" authority="Emery, 1895: 268" authorityName="Emery" authorityPageNumber="268" authorityYear="1895" box="[786,1100,1826,1848]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="7CBAEAA15B27FF9F9965F899FC67F8FC" box="[786,906,1826,1848]" italics="true" pageId="14" pageNumber="529">O. insularis</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F99E1F899FBA1F8FC" author="Emery" box="[918,1100,1826,1848]" page="268" pageId="14" pageNumber="529" year="1895">
<bibRefCitation id="2A5F4B425B27FF9F99E1F899FBF7F8FC" author="Emery" box="[918,1050,1826,1848]" pageId="14" pageNumber="529" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery, 1895</bibRefCitation>
: 268
</treatmentCitation>
</taxonomicName>
; of
<taxonomicName id="89CE4D305B27FF9F9E06F899FA75F8FC" ID-CoL="48QBK" authority="Wheeler, 1902: 26" authorityName="Wheeler" authorityPageNumber="26" authorityYear="1902" box="[1137,1432,1826,1848]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9E06F899FB22F8FC" box="[1137,1231,1826,1848]" italics="true" pageId="14" pageNumber="529">O. clarus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9EABF898FA75F8FC" author="Wheeler" box="[1244,1432,1826,1848]" page="26" pageId="14" pageNumber="529" year="1902">
<bibRefCitation id="2A5F4B425B27FF9F9EABF898FA9EF8FC" author="Wheeler" box="[1244,1395,1826,1848]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1902) A consideration of S. B. Buckley's &quot; North American Formicidae. &quot; Transactions of the Texas Academy of Sciences, 4, 17 - 31." type="journal article" year="1902">Wheeler, 1902</bibRefCitation>
: 26
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F8DAFED9F850" blockId="14.[151,1437,1888,2002]" pageId="14" pageNumber="529">
<treatmentCitationGroup id="6EDE119D5B27FF9F9AE0F8DAFED9F850" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9AE0F8DAFE5EF8B2" ID-CoL="48QBS" box="[151,435,1888,1910]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B27FF9F9AE0F8DAFE5EF8B2" box="[151,435,1888,1910]" italics="true" pageId="14" pageNumber="529">Odontomachus haematodus</emphasis>
</taxonomicName>
subsp.
<taxonomicName id="89CE4D305B27FF9F9873F8DAFCD3F8B2" authority="Wheeler, 1915: 391" authorityName="Wheeler" authorityPageNumber="391" authorityYear="1915" box="[516,830,1888,1910]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="coninodis">
<emphasis id="7CBAEAA15B27FF9F9873F8DAFD8AF8B2" box="[516,615,1888,1910]" italics="true" pageId="14" pageNumber="529">coninodis</emphasis>
<treatmentCitation id="CF6F10A25B27FF9F9818F8DAFCD3F8B2" author="Wheeler" box="[623,830,1888,1910]" page="391" pageId="14" pageNumber="529" year="1915">
<bibRefCitation id="2A5F4B425B27FF9F9818F8DAFCE4F8B2" author="Wheeler" box="[623,777,1888,1910]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler, 1915</bibRefCitation>
: 391
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B27FF9F993CF8DBFBE7F8B2" author="Brown" box="[843,1034,1888,1910]" page="103" pageId="14" pageNumber="529" year="1976">
<bibRefCitation id="2A5F4B425B27FF9F993CF8DBFC39F8B2" author="Brown" box="[843,980,1888,1910]" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
. Junior synonym of
<taxonomicName id="89CE4D305B27FF9F9E96F8DBFEDDF850" ID-CoL="48QBK" authority="Brown, 1976: 103" authorityName="Brown" authorityPageNumber="103" authorityYear="1976" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9E96F8DBFAACF8B2" box="[1249,1345,1888,1910]" italics="true" pageId="14" pageNumber="529">O. clarus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9F26F8DBFEDDF850" author="Brown" page="103" pageId="14" pageNumber="529" year="1976">
<bibRefCitation id="2A5F4B425B27FF9F9F26F8DBFF10F850" author="Brown" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B27FF9F9AE0F827FF19F816" blockId="14.[151,1437,1888,2002]" pageId="14" pageNumber="529">
<treatmentCitationGroup id="6EDE119D5B27FF9F9AE0F827FF19F816" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9AE0F827FE5BF877" box="[151,438,1949,1971]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haemataoda">
<emphasis id="7CBAEAA15B27FF9F9AE0F827FE5BF877" box="[151,438,1949,1971]" italics="true" pageId="14" pageNumber="529">Odontomachus haemataoda</emphasis>
</taxonomicName>
subsp.
<emphasis id="7CBAEAA15B27FF9F987FF827FD69F877" box="[520,644,1949,1971]" italics="true" pageId="14" pageNumber="529">clarionensis</emphasis>
<treatmentCitation id="CF6F10A25B27FF9F98FBF827FCB6F877" author="Wheeler" box="[652,859,1949,1971]" page="141" pageId="14" pageNumber="529" year="1934">
<bibRefCitation id="2A5F4B425B27FF9F98FBF827FCCBF877" author="Wheeler" box="[652,806,1949,1971]" pageId="14" pageNumber="529" refString="Wheeler, W. M. (1934) Ants from the islands off the west coast of Lower California and Mexico. Pan-Pacific Entomologist, 10, 132 - 144." type="journal article" year="1934">Wheeler, 1934</bibRefCitation>
: 141
</treatmentCitation>
. Junior synonym of
<taxonomicName id="89CE4D305B27FF9F9E44F824FF1DF816" ID-CoL="48QBK" authority="Taylor &amp; Wilson, 1962: 142" authorityName="Taylor &amp; Wilson" authorityPageNumber="142" authorityYear="1962" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B27FF9F9E44F824FB79F877" box="[1075,1172,1949,1971]" italics="true" pageId="14" pageNumber="529">O. clarus</emphasis>
:
<treatmentCitation id="CF6F10A25B27FF9F9ED3F827FF1DF816" author="Taylor" page="142" pageId="14" pageNumber="529" year="1962">
<bibRefCitation id="2A5F4B425B27FF9F9ED3F827FA74F877" author="Taylor" box="[1188,1433,1949,1971]" pageId="14" pageNumber="529" refString="Taylor, R. W. &amp; Wilson, E. O. (1962 [&quot; 1961 &quot;]) Ants from three remote oceanic islands. Psyche 68, 137 - 144. http: // dx. doi. org / 10.1155 / 1961 / 87982" type="journal article" year="1962">Taylor &amp; Wilson, 1962</bibRefCitation>
: 142
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="1AB1663B5B26FF9E9AE0F9C9FEFEF902" httpUri="https://zenodo.org/record/249909/files/figure.png" pageId="15" pageNumber="530" targetBox="[151,1436,195,1629]" targetPageId="15">
<paragraph id="4E7136B35B26FF9E9AE0F9C9FEFEF902" blockId="15.[151,1436,1651,1734]" pageId="15" pageNumber="530">
<emphasis id="7CBAEAA15B26FF9E9AE0F9C9FEF2F94D" bold="true" box="[151,287,1651,1673]" pageId="15" pageNumber="530">FIGURE 14.</emphasis>
<taxonomicName id="89CE4D305B26FF9E9B51F9C9FDE8F94D" box="[294,517,1651,1673]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E9B51F9C9FDE8F94D" box="[294,517,1651,1673]" italics="true" pageId="15" pageNumber="530">Odontomachus clarus</emphasis>
</taxonomicName>
, worker: (A) head in full-face view, (B) lateral habitus, (C) dorsal habitus, (D) mesosoma in lateral view, (E) mesosoma in dorsal view, (F) petiole in lateral view, and (G) first gastral tergite in dorsal view showing pubescence.
</paragraph>
</caption>
<subSubSection id="06D465385B26FF819AE0F957FB0BFEA1" lastPageId="16" lastPageNumber="531" pageId="15" pageNumber="530" type="diagnosis">
<paragraph id="4E7136B35B26FF819AE0F957FB0BFEA1" blockId="15.[151,1436,1773,2014]" lastBlockId="16.[151,1436,151,357]" lastPageId="16" lastPageNumber="531" pageId="15" pageNumber="530">
<emphasis id="7CBAEAA15B26FF9E9AE0F957FEFCF8C2" bold="true" box="[151,273,1773,1798]" pageId="15" pageNumber="530">Diagnosis.</emphasis>
Workers of
<taxonomicName id="89CE4D305B26FF9E9BE8F954FDE5F8C1" box="[415,520,1773,1797]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E9BE8F954FDE5F8C1" box="[415,520,1773,1797]" italics="true" pageId="15" pageNumber="530">O. clarus</emphasis>
</taxonomicName>
are most similar to
<taxonomicName id="89CE4D305B26FF9E989AF954FC89F8C1" box="[749,868,1773,1797]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E989AF954FC89F8C1" box="[749,868,1773,1797]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B26FF9E99EBF954FBD2F8C1" box="[924,1087,1773,1797]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B26FF9E99EBF954FBD2F8C1" box="[924,1087,1773,1797]" italics="true" pageId="15" pageNumber="530">O. desertorum</emphasis>
</taxonomicName>
, but are distinguished from
<taxonomicName id="89CE4D305B26FF9E9FF6F954FF07F8ED" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E9FF6F954FF07F8ED" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
by their smooth basilar lobes (striate in
<taxonomicName id="89CE4D305B26FF9E98DCF8A8FCCDF8ED" box="[683,800,1809,1833]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E98DCF8A8FCCDF8ED" box="[683,800,1809,1833]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
), and from
<taxonomicName id="89CE4D305B26FF9E99D4F8A8FBA9F8ED" box="[931,1092,1809,1833]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B26FF9E99D4F8A8FBA9F8ED" box="[931,1092,1809,1833]" italics="true" pageId="15" pageNumber="530">O. desertorum</emphasis>
</taxonomicName>
by the following: Size smaller (HL &lt;
<quantity id="89369B565B26FF9E9A9FF88CFEAFF889" box="[232,322,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="530" unit="mm" value="2.2">2.2 mm</quantity>
, WL &lt;
<quantity id="89369B565B26FF9E9BE8F88CFE16F88A" box="[415,507,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="530" unit="mm" value="2.8">2.8 mm</quantity>
); dorsomedian petiolar process nub-like or well-differentiated; propleurae almost entirely smooth and polished-looking; pronotal cervical lobe longer than broad; and antennal scapes relatively shorter (SL/HW = 0.99–1.07 vs. 1.11–1.15).
<taxonomicName id="89CE4D305B26FF9E98D4F8C7FC70F851" box="[675,925,1917,1941]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E98D4F8C7FC70F851" box="[675,925,1917,1941]" italics="true" pageId="15" pageNumber="530">Odontomachus clarus</emphasis>
</taxonomicName>
is further differentiated from
<taxonomicName id="89CE4D305B26FF9E9E8AF8C4FA9BF851" box="[1277,1398,1917,1941]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E9E8AF8C4FA9BF851" box="[1277,1398,1917,1941]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
by geography:
<taxonomicName id="89CE4D305B26FF9E9B56F818FE62F87D" box="[289,399,1953,1977]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B26FF9E9B56F818FE62F87D" box="[289,399,1953,1977]" italics="true" pageId="15" pageNumber="530">O. clarus</emphasis>
</taxonomicName>
only occurs west of the Mississippi river; whereas,
<taxonomicName id="89CE4D305B26FF9E9E73F818FB92F87D" box="[1028,1151,1953,1977]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="7CBAEAA15B26FF9E9E73F818FB92F87D" box="[1028,1151,1953,1977]" italics="true" pageId="15" pageNumber="530">O. relictus</emphasis>
</taxonomicName>
only occurs east of the Mississippi river. Males are uniquely identifiable among the Nearctic fauna by, at minimum, the following combination of characters: 1) dark brown coloration; 2) ocelli small, weakly bulging past posterior head margin; 3) mesoscutum longitudinally striate (striae may be posterolaterally divergent); 4) posterior margin of petiolar sternum lacking an angular process, but with paired setae on tubercles; and 5) propodeum finely striate. The ninth abdominal sternum of
<taxonomicName id="89CE4D305B39FF819BEAFEBEFDE4FED8" box="[413,521,260,284]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B39FF819BEAFEBEFDE4FED8" box="[413,521,260,284]" italics="true" pageId="16" pageNumber="531">O. clarus</emphasis>
</taxonomicName>
is unique, having a short, broad apical portion of abdominal sternum IX (
<figureCitation id="D6F52A365B39FF819F18FEBEFF5AFE84" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="16" pageNumber="531">Fig. 24</figureCitation>
B). The genitalia are otherwise most similar to those of
<taxonomicName id="89CE4D305B39FF819935FE92FC0BFE84" box="[834,998,296,320]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B39FF819935FE92FC0BFE84" box="[834,998,296,320]" italics="true" pageId="16" pageNumber="531">O. desertorum</emphasis>
</taxonomicName>
, differing primarily in having a more strongly sclerotized digitus and anteroventral process shorter, sharply triangular, and not upturned.
</paragraph>
</subSubSection>
<caption id="1AB1663B5B39FF819AE0F8CEFB92F86C" httpUri="https://zenodo.org/record/249910/files/figure.png" pageId="16" pageNumber="531" targetBox="[151,1436,409,1886]" targetPageId="16">
<paragraph id="4E7136B35B39FF819AE0F8CEFB92F86C" blockId="16.[151,1436,1908,1960]" pageId="16" pageNumber="531">
<emphasis id="7CBAEAA15B39FF819AE0F8CEFECCF84E" bold="true" box="[151,289,1908,1930]" pageId="16" pageNumber="531">FIGURE 15.</emphasis>
<taxonomicName id="89CE4D305B39FF819B5EF8CEFDE4F84E" box="[297,521,1908,1930]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B39FF819B5EF8CEFDE4F84E" box="[297,521,1908,1930]" italics="true" pageId="16" pageNumber="531">Odontomachus clarus</emphasis>
</taxonomicName>
, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<subSubSection id="06D465385B38FF809AB0FF2DFDD4FABC" pageId="17" pageNumber="532" type="description">
<paragraph id="4E7136B35B38FF809AB0FF2DFAE2FD63" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">
<emphasis id="7CBAEAA15B38FF809AB0FF2DFEB4FF74" bold="true" box="[199,345,151,176]" pageId="17" pageNumber="532">Description.</emphasis>
Worker: HL 1.92–2.32, HW 1.57–1.90, SL 1.68–1.90, EL 0.31–0.36, ML 1.06–1.30, WL 2.43–2.83, PTH 0.81–1.10, PTL 0.42–0.48 (n=7). Entire body generally shiny except where obscured by dense pubescence; head, mesosoma, and petiole reddish-brown, legs yellowish-brown to orangish-brown, and gaster blackish. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning from frontal lobes and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head lacking sculpture; with numerous, fine, appressed pubescence and scattered elongate, erect setae present dorsally. Pronotum with circular, slightly concentric striae that become transverse near rear margin; appressed pubescence abundant; 5–6 elongate, erect setae present. Mesonotum and propodeum with deep transverse striae; propleuron, mesopleuron, and basalar lobe lacking sculpture; abundant pubescence present dorsally. Metasternum lacking paired elongate, spiniform processes between hind coxae. Petiole widest at base, either gradually tapering apically to a short spiniform spine or stoutly conical (see
<figureCitation id="D6F52A365B38FF809964FE45FCB2FDDC" box="[787,863,511,536]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="17" pageNumber="532">Figs. 6</figureCitation>
D, E and 14F); mostly lacking striae with only faint striae present near base; subpetiolar process rounded triangular; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly shiny beneath pubescence, lacking striae or other strong sculpture, but with fine coriaceous sculpture (seen at high magnification); slightly coarse, appressed pubescence somewhat sparse, hairs separated by at least 1/2 of their length; scattered erect, elongate setae present.
</paragraph>
<paragraph id="4E7136B35B38FF809AB0FD0EFA8AFAF4" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">Male: HL 0.91–0.97, HW 1.18–1.30, SL 0.17–0.25, EL 0.62–0.64, EW 0.35–0.37, OL 0.12–0.14, OES 0.25–0.26, PTH 0.66–0.73, WL 2.39–2.65, PTL 0.40–0.59, FWL 4.56–4.97 (n=3). Body generally shiny except where obscured by dense pubescence; head and mesosoma brownish-black, petiole and gaster dark reddish-brown to brownish-black, legs and antennae reddish-brown. Head and body with abundant semi erect, whitish pubescence. Eyes extremely large, maximum diameter of each eye at least 70% of the length of the head in fullface view. Ocelli small, the length of each ocellus about half the distance between lateral ocellus and eye margin; in full-face view, lateral ocelli do not noticeably protrude beyond posterior border of head. Mesosoma: pronotum lacking sculpture; mesoscutum with fine transversely arcuate striae anteriorly, striae becoming longitudinal posteriorly; mesoscutellum raised and convex, with longitudinal striae; propodeum and mesopleuron with weak to slightly strong longitudinal striae. Petiole bluntly rounded apically, with apically acute to apically rounded triangular subpetiolar process present anteriorly; densely pubescent anteriorly and laterally, with reduced pubescence posteriorly; petiolar sternum lacking angular process near posterior margin, but with paired setae on small tubercles. Abdominal sternum IX disc subrectangular, breadth almost twice length; posterior lobe length less than twice maximum width, basal half barely narrowed. Telomeral apex narrowly rounded; telomere length distinctly greater than height; valviceps ventral apex strongly produced and very narrow; apical margin of valviceps linear until abruptly curving posteriorly at apicoventral process; anteroventral process of valviceps sharply triangular, and not upturned; vertical portion of dorsolateral carina and lateral margin of subapical lamina curving into one another; apicodorsal lobe of valviceps narrow and dorsally elongate; subapical lamina broad.</paragraph>
<paragraph id="4E7136B35B38FF809AB0FA86FDD4FABC" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">Queen: No specimens measured, but similar to workers in color and general appearance except slightly larger, with mesosoma developed for wings.</paragraph>
</subSubSection>
<subSubSection id="06D465385B38FF809AB0FA3EFF0CF98B" pageId="17" pageNumber="532" type="distribution">
<paragraph id="4E7136B35B38FF809AB0FA3EFF0CF98B" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="532">
<emphasis id="7CBAEAA15B38FF809AB0FA3EFDD1FA59" bold="true" box="[199,572,1412,1437]" pageId="17" pageNumber="532">
Distribution (
<figureCitation id="D6F52A365B38FF809B19FA3EFE27FA59" box="[366,458,1412,1437]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="17" pageNumber="532">Fig. 26</figureCitation>
).
<collectingCountry id="36D976235B38FF809B92FA3EFDD1FA59" box="[485,572,1412,1437]" name="Mexico" pageId="17" pageNumber="532">Mexico</collectingCountry>
</emphasis>
. Chihuahua, Coahuila, Hidalgo, Nuevo Leon, and Clarión Island.
<emphasis id="7CBAEAA15B38FF809F2AFA3EFA71FA59" bold="true" box="[1373,1436,1412,1437]" pageId="17" pageNumber="532">
<collectingCountry id="36D976235B38FF809F2AFA3EFA7AFA59" box="[1373,1431,1412,1437]" name="United States of America" pageId="17" pageNumber="532">USA</collectingCountry>
.
</emphasis>
Arizona: Cochise, Pima, Santa Cruz, and Yavapai Counties. New
<collectingCountry id="36D976235B38FF8099D6FA1DFC1AFA04" box="[929,1015,1447,1472]" name="Mexico" pageId="17" pageNumber="532">Mexico</collectingCountry>
. McKinley County. Texas: Bastrop, Bexar, Bosque, Brewster, Brown, Denton, Jeff Davis, and Travis Counties. Louisiana: Natchitoches Parish. Distributional information from AntWeb (2013), Anonymous (2013),
<bibRefCitation id="2A5F4B425B38FF8099DCFA4AFBA6F9CC" author="Brown" box="[939,1099,1519,1544]" pageId="17" pageNumber="532" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
,
<bibRefCitation id="2A5F4B425B38FF809E2FFA55FAD1F9CC" box="[1112,1340,1519,1544]" pageId="17" pageNumber="532" refString="Adams, B. J., Chen, X. &amp; Hooper-Bui, L. M. (2010) Odontomachus clarus Roger (Hymenoptera: Formicidae) reported in Kisatchie National Forest, Louisiana. Midsouth Entomologist, 3, 104 - 105." type="journal article">
Adams
<emphasis id="7CBAEAA15B38FF809ED9FA4BFB32F9CC" box="[1198,1247,1520,1544]" italics="true" pageId="17" pageNumber="532">et al</emphasis>
. (2010)
</bibRefCitation>
,
<bibRefCitation id="2A5F4B425B38FF809F3FFA4AFE88F9E8" author="Deyrup" pageId="17" pageNumber="532" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
, and examination of specimens from ABS, JTLC, LACM, LSAM, MEM, UCDC, UGA, and UTEP.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B38FF839AB0F9E6FEF0FBF0" lastPageId="18" lastPageNumber="533" pageId="17" pageNumber="532" type="discussion">
<paragraph id="4E7136B35B38FF839AB0F9E6FA84FE84" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,1077]" lastPageId="18" lastPageNumber="533" pageId="17" pageNumber="532">
<emphasis id="7CBAEAA15B38FF809AB0F9E6FEA1F9B1" bold="true" box="[199,332,1628,1653]" pageId="17" pageNumber="532">Discussion.</emphasis>
<taxonomicName id="89CE4D305B38FF809B23F9E6FDA6F9B0" box="[340,587,1628,1652]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B38FF809B23F9E6FDA6F9B0" box="[340,587,1628,1652]" italics="true" pageId="17" pageNumber="532">Odontomachus clarus</emphasis>
</taxonomicName>
has had four specific synonyms until the present paper. While the Buckley name may be safely ignored as superfluous and
<taxonomicName id="89CE4D305B38FF8098CAF93AFC8CF95C" box="[701,865,1664,1688]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="desertorum">
<emphasis id="7CBAEAA15B38FF8098CAF93AFC8CF95C" box="[701,865,1664,1688]" italics="true" pageId="17" pageNumber="532">O. desertorum</emphasis>
</taxonomicName>
is here recognized as a valid species, the speciesstatus of the other two names stands to be reassessed.
<taxonomicName id="89CE4D305B38FF809979F91EFBEAF978" box="[782,1031,1700,1724]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B38FF809979F91EFBEAF978" box="[782,1031,1700,1724]" italics="true" pageId="17" pageNumber="532">Odontomachus clarus</emphasis>
</taxonomicName>
is variable across its range, with a broad longitudinal gradient from Texas and Louisiana South to Hidalgo,
<collectingCountry id="36D976235B38FF809992F97DFBD0F924" box="[997,1085,1735,1760]" name="Mexico" pageId="17" pageNumber="532">Mexico</collectingCountry>
, and a very narrow shift from Texas into the mountains of Arizona. The US-Mexico cline is manifested in a North-to-South trend of increasing intensity of gastral sculpturation, erection of gastral pubescence, and anteroposterior compression of the petiolar node. The Mexican High Plateau population is otherwise identical to those of the Great and South Central Plains. The montane Arizonan populations, located in the Chiricahua, Huachua, and Santa Rita Mountains, as well as the Colorado Plateau Transition Zone, have previously been provided with the subspecific name
<taxonomicName id="89CE4D305B38FF809ECBF8C6FEA6F87D" authority="Wheeler 1915" authorityName="Wheeler" authorityYear="1915" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="subSpecies" species="clarus" subSpecies="coninodis">
<emphasis id="7CBAEAA15B38FF809ECBF8C6FA71F850" box="[1212,1436,1916,1940]" italics="true" pageId="17" pageNumber="532">O. clarus coninodis</emphasis>
(
<bibRefCitation id="2A5F4B425B38FF809AE9F81AFEAFF87D" author="Wheeler" box="[158,322,1952,1977]" pageId="17" pageNumber="532" refString="Wheeler, W. M. (1915) Some additions to the North American ant-fauna. Bulletin of the American Museum of Natural History, 34, 389 - 421." type="journal article" year="1915">Wheeler 1915</bibRefCitation>
)
</taxonomicName>
due to their nub-like dorsomedian petiolar process (see
<figureCitation id="D6F52A365B38FF8099A8F81AFBC9F87D" box="[991,1060,1952,1977]" captionStart="FIGURE 6" captionStartId="8.[151,250,1946,1968]" captionTargetBox="[229,1361,1133,1929]" captionTargetId="figure@8.[229,1361,1133,1929]" captionTargetPageId="8" captionText="FIGURE 6. Worker propleurae (prp) and pronota (prn), ventrolateral oblique view: (A) O. desertorum, note striations on propleurae, and (B) O. clarus, note the smooth propleurae; and worker petiolar nodes, anterior view: (C) O. desertorum and (D, E) O. clarus." httpUri="https://zenodo.org/record/249902/files/figure.png" pageId="17" pageNumber="532">Fig. 6</figureCitation>
E). Upon careful examination of material across the range of
<taxonomicName id="89CE4D305B38FF809BA1F87EFDD2F818" box="[470,575,1988,2012]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B38FF809BA1F87EFDD2F818" box="[470,575,1988,2012]" italics="true" pageId="17" pageNumber="532">O. clarus</emphasis>
</taxonomicName>
it is clear that this is the only distinguishing character for these populations, and that the conical dorsomedian petiolar process merely represents a foreshortening of the process as the rest of the petiole is otherwise unmodified. Moreover, males attributable to “
<taxonomicName id="89CE4D305B3BFF8399C5FF06FBCCFF11" box="[946,1057,188,213]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="coninodis">coninodis</taxonomicName>
are inseparable from those of
<taxonomicName id="89CE4D305B3BFF839FF6FF06FF30FF3C" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839FF6FF06FF30FF3C" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
except for reduced size. These observations strongly suggest that the name
<taxonomicName id="89CE4D305B3BFF839E16FF5AFAABFF3C" box="[1121,1350,224,248]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="subSpecies" species="clarus" subSpecies="coninodis">
<emphasis id="7CBAEAA15B3BFF839E16FF5AFAABFF3C" box="[1121,1350,224,248]" italics="true" pageId="18" pageNumber="533">O. clarus coninodis</emphasis>
</taxonomicName>
should remain in synonymy with
<taxonomicName id="89CE4D305B3BFF839BB6FEBEFDC6FED8" box="[449,555,260,284]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BB6FEBEFDC6FED8" box="[449,555,260,284]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
. It would still be valuable to conduct a sampling transect from Texas through the southeastern Arizona mountains to detect intergradation of petiolar node form as
<bibRefCitation id="2A5F4B425B3BFF839E39FE92FB01FE84" author="Brown" box="[1102,1260,295,320]" pageId="18" pageNumber="533" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
suggested.
</paragraph>
<paragraph id="4E7136B35B3BFF839AB0FEF6FB4AFD41" blockId="18.[151,1437,151,1077]" pageId="18" pageNumber="533">
Finally, the status of the name
<taxonomicName id="89CE4D305B3BFF839844FEF6FCDEFEA0" box="[563,819,332,356]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="subSpecies" species="clarus" subSpecies="clarionensis">
<emphasis id="7CBAEAA15B3BFF839844FEF6FCDEFEA0" box="[563,819,332,356]" italics="true" pageId="18" pageNumber="533">O. clarus clarionensis</emphasis>
</taxonomicName>
remains to be absolutely determined. The name was provided by
<bibRefCitation id="2A5F4B425B3BFF839B53FED5FE38FE4C" author="Wheeler" box="[292,469,367,392]" pageId="18" pageNumber="533" refString="Wheeler, W. M. (1934) Ants from the islands off the west coast of Lower California and Mexico. Pan-Pacific Entomologist, 10, 132 - 144." type="journal article" year="1934">Wheeler (1934)</bibRefCitation>
for an insular population on Clarión Island (about
<quantity id="89369B565B3BFF839E78FECAFB89FE4C" box="[1039,1124,367,392]" metricMagnitude="5" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="533" unit="km" value="700.0">700 km</quantity>
WSW from the tip of Baja) of what he considered to be
<taxonomicName id="89CE4D305B3BFF839BA5FE2EFD96FE68" box="[466,635,404,428]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B3BFF839BA5FE2EFD96FE68" box="[466,635,404,428]" italics="true" pageId="18" pageNumber="533">O. haematodus</emphasis>
</taxonomicName>
. Wheeler erected the subspecies based on the larger size, broader head, and smoother posterolateral head corners than compared to
<taxonomicName id="89CE4D305B3BFF839944FE02FC3FFE14" box="[819,978,440,464]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="haematoda">
<emphasis id="7CBAEAA15B3BFF839944FE02FC3FFE14" box="[819,978,440,464]" italics="true" pageId="18" pageNumber="533">O. haematoda</emphasis>
</taxonomicName>
(an artificial assembly not recognized as such at the time).
<bibRefCitation id="2A5F4B425B3BFF839B2AFE66FD91FE31" author="Taylor" box="[349,636,476,501]" pageId="18" pageNumber="533" refString="Taylor, R. W. &amp; Wilson, E. O. (1962 [&quot; 1961 &quot;]) Ants from three remote oceanic islands. Psyche 68, 137 - 144. http: // dx. doi. org / 10.1155 / 1961 / 87982" type="journal article" year="1962">Taylor and Wilson (1962)</bibRefCitation>
synonymized the subspecies with
<taxonomicName id="89CE4D305B3BFF839989FE66FB8BFE30" box="[1022,1126,476,500]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839989FE66FB8BFE30" box="[1022,1126,476,500]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
without discussion, and the population has been noted as “typical for
<taxonomicName id="89CE4D305B3BFF83980DFDBAFD08FDDC" box="[634,741,512,536]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF83980DFDBAFD08FDDC" box="[634,741,512,536]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
by
<bibRefCitation id="2A5F4B425B3BFF839963FDBAFC5AFDDC" author="Brown" box="[788,951,511,536]" pageId="18" pageNumber="533" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
. Examination of one of the cotypes at the MCZ does not reveal any salient distinctions of the Clarión Island population from mainland
<taxonomicName id="89CE4D305B3BFF839E87FD9EFAB0FDF8" box="[1264,1373,548,572]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839E87FD9EFAB0FDF8" box="[1264,1373,548,572]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
. The synonymy is thus tentatively confirmed here; however, it would be valuable to sequence genetic material of the Clarión Island population of
<taxonomicName id="89CE4D305B3BFF839BADFDD6FDAFFD40" box="[474,578,620,644]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BADFDD6FDAFFD40" box="[474,578,620,644]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
to compare to material from across the species’ range.
</paragraph>
<paragraph id="4E7136B35B3BFF839AB0FD35FEF0FBF0" blockId="18.[151,1437,151,1077]" pageId="18" pageNumber="533">
The first report of
<taxonomicName id="89CE4D305B3BFF839BD7FD2AFD77FD6C" box="[416,666,656,680]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BD7FD2AFD77FD6C" box="[416,666,656,680]" italics="true" pageId="18" pageNumber="533">Odontomachus clarus</emphasis>
</taxonomicName>
from Louisiana was based on specimens collected in Provencal in Natchitoches Parish (
<bibRefCitation id="2A5F4B425B3BFF839BF3FD0EFDE6FD08" author="Smith" box="[388,523,692,717]" pageId="18" pageNumber="533" refString="Smith, M. R. (1939) A study of the subspecies of Odontomachus haematoda (L.) of the United States (Hymenoptera: Formicid). Journal of the New York Entomological Society, 47, 125 - 130." type="journal article" year="1939">Smith 1939</bibRefCitation>
). Unfortunately, we were unable to locate Smith’s specimens to verify this record. Subsequent authorities implicitly questioned this disjunct record:
<bibRefCitation id="2A5F4B425B3BFF839905FD62FBE2FD34" author="Brown" box="[882,1039,727,752]" pageId="18" pageNumber="533" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown (1976)</bibRefCitation>
omitted it entirely from his lengthy monograph, and this record was later omitted by
<bibRefCitation id="2A5F4B425B3BFF8398B0FD47FC02FCD0" author="Deyrup" box="[711,1007,764,789]" pageId="18" pageNumber="533" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
. Smith's efforts were later vindicated, however, as specimens of
<taxonomicName id="89CE4D305B3BFF839BCCFC9AFDCEFCFC" box="[443,547,800,824]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839BCCFC9AFDCEFCFC" box="[443,547,800,824]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
were collected nesting under large rocks in sandy soil in the Kisatchie National Forest in Natchitoches Parish just
<quantity id="89369B565B3BFF83986DFCFEFD8FFC99" box="[538,610,836,861]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="533" unit="km" value="20.0">20 km</quantity>
SW of where Smith reported this species (
<bibRefCitation id="2A5F4B425B3BFF839E32FCFEFAF6FC98" author="Adams" box="[1093,1307,836,861]" pageId="18" pageNumber="533" refString="Adams, B. J., Chen, X. &amp; Hooper-Bui, L. M. (2010) Odontomachus clarus Roger (Hymenoptera: Formicidae) reported in Kisatchie National Forest, Louisiana. Midsouth Entomologist, 3, 104 - 105." type="journal article" year="2010">
Adams
<emphasis id="7CBAEAA15B3BFF839EE9FCFFFB3DFC98" box="[1182,1232,836,860]" italics="true" pageId="18" pageNumber="533">et al</emphasis>
. 2010
</bibRefCitation>
). Similarly, in Texas, nesting sites are typically in dry sites. In Arizona, its nesting habits appear more flexible, as it has been collected under rocks and grass tussocks in dry to mesic habitats (
<bibRefCitation id="2A5F4B425B3BFF8399FEFC36FB4FFC60" author="Deyrup" box="[905,1186,908,933]" pageId="18" pageNumber="533" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover 2004</bibRefCitation>
). These differences in nest site selection may yet reveal whether the “
<taxonomicName id="89CE4D305B3BFF8398CBFC15FCC6FC0C" box="[700,811,943,968]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="coninodis">coninodis</taxonomicName>
populations represent a distinct species, although this seems unlikely. Regardless of the results of increased sampling and molecular work on the Arizona populations, the Louisiana populations of
<taxonomicName id="89CE4D305B3BFF839B95FC42FDA6FBD4" box="[482,587,1016,1040]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B3BFF839B95FC42FDA6FBD4" box="[482,587,1016,1040]" italics="true" pageId="18" pageNumber="533">O. clarus</emphasis>
</taxonomicName>
would certainly retain that specific epithet as the
<typeStatus id="917588115B3BFF839EF4FC42FB5EFBD4" box="[1155,1203,1016,1040]" pageId="18" pageNumber="533">type</typeStatus>
series was collected from Texas.
</paragraph>
</subSubSection>
</treatment>
</document>
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