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<document id="1183D8EE94E7FA0445D5FF3CFAB7494C" ID-CLB-Dataset="45647" ID-DOI="10.11646/zootaxa.3802.4.6" ID-GBIF-Dataset="9a1086b4-468f-472e-9dc6-17b9e73d1e8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="249896" ID-ZooBank="4A83AAAE-6F0B-4173-A066-DBBCFBC3BDD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460427674668" checkinUser="plazi" docAuthor="Macgown, Joe A., Boudinot, Brendon, Deyrup, Mark &amp; Sorger, D. Magdalena" docDate="2014" docId="C66787A55B22FF9F9AE0FCE4FB63F9CC" docLanguage="en" docName="zt03802p552.pdf" docOrigin="Zootaxa 3802 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Odontomachus brunneus Patton 1894" docType="treatment" docVersion="18" lastPageNumber="529" masterDocId="3A5EFFDD5B29FF919A77FFBAFFEDFFC4" masterDocTitle="A review of the Nearctic Odontomachus (Hymenoptera: Formicidae: Ponerinae) with a treatment of the males" masterLastPageNumber="552" masterPageNumber="515" pageNumber="526" updateTime="1732493224424" updateUser="ExternalLinkService">
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<mods:title id="B6F782B2F67221453AB3E1CFBA98D7D2">A review of the Nearctic Odontomachus (Hymenoptera: Formicidae: Ponerinae) with a treatment of the males</mods:title>
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<mods:namePart id="AD27AD3305BA6048B3E1BBD244008CFF">Macgown, Joe A.</mods:namePart>
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<mods:namePart id="D1F6997A51EAB8C4E7EDA2D0E8992D9C">Sorger, D. Magdalena</mods:namePart>
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<treatment id="C66787A55B22FF9F9AE0FCE4FB63F9CC" ID-DOI="http://doi.org/10.5281/zenodo.6129408" ID-GBIF-Taxon="119429108" ID-Zenodo-Dep="6129408" LSID="urn:lsid:plazi:treatment:C66787A55B22FF9F9AE0FCE4FB63F9CC" httpUri="http://treatment.plazi.org/id/C66787A55B22FF9F9AE0FCE4FB63F9CC" lastPageId="14" lastPageNumber="529" pageId="11" pageNumber="526" scope_family="Formicidae" scope_order="Hymenoptera" scope_subFamily="Ponerinae">
<subSubSection id="06D465385B22FF9A9AE0FCE4FDEDFC5E" pageId="11" pageNumber="526" type="nomenclature">
<paragraph id="4E7136B35B22FF9A9AE0FCE4FD6DFCBC" blockId="11.[151,640,862,922]" box="[151,640,862,888]" pageId="11" pageNumber="526">
<heading id="153981DF5B22FF9A9AE0FCE4FD6DFCBC" bold="true" box="[151,640,862,888]" fontSize="11" level="1" pageId="11" pageNumber="526" reason="1">
<taxonomicName id="89CE4D305B22FF9A9AE0FCE4FD6DFCBC" ID-CoL="6SH5N" authority="Patton, 1894" authorityName="Patton" authorityYear="1894" box="[151,640,862,888]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9AE0FCE4FD6DFCBC" bold="true" box="[151,640,862,888]" pageId="11" pageNumber="526">
<emphasis id="7CBAEAA15B22FF9A9AE0FCE4FE27FCBC" bold="true" box="[151,458,862,888]" italics="true" pageId="11" pageNumber="526">Odontomachus brunneus</emphasis>
(
<bibRefCitation id="2A5F4B425B22FF9A9BAEFCE4FD95FCBC" author="Patton" box="[473,632,862,888]" pageId="11" pageNumber="526" refString="Patton, W. H. (1894) Habits of the leaping-ant of southern Georgia. American Naturalist, 28, 618 - 619." type="journal article" year="1894">Patton, 1894</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4E7136B35B22FF9A9AE0FC3BFDEDFC5E" blockId="11.[151,640,862,922]" box="[151,512,897,922]" pageId="11" pageNumber="526">
(
<figureCitation id="D6F52A365B22FF9A9AE8FC3BFF03FC5D" box="[159,238,897,922]" captionStart="FIGURE 4" captionStartId="7.[151,249,1425,1447]" captionTargetBox="[151,1435,486,1403]" captionTargetId="figure@7.[151,1435,486,1403]" captionTargetPageId="7" captionText="FIGURE 4. First gastral tergites of workers, dorsal view: (A) O. brunneus and (B) O. clarus." httpUri="https://zenodo.org/record/249900/files/figure.png" pageId="11" pageNumber="526">Figs. 4</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9A8DFC38FEE7FC5D" box="[250,266,898,921]" captionStart="FIGURE 7" captionStartId="9.[151,249,1080,1102]" captionTargetBox="[151,1435,410,1059]" captionTargetId="figure@9.[151,1435,410,1059]" captionTargetPageId="9" captionText="FIGURE 7. Male heads, full-face view: (A) O. brunneus and (B) O. haematodus. LOC = lateral ocelli, MOC = median ocellus." httpUri="https://zenodo.org/record/249903/files/figure.png" pageId="11" pageNumber="526">7</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9B61FC3BFEC8FC5D" box="[278,293,897,921]" captionStart="FIGURE 8" captionStartId="9.[151,249,1900,1922]" captionTargetBox="[151,1435,1331,1878]" captionTargetId="figure@9.[151,1435,1331,1879]" captionTargetPageId="9" captionText="FIGURE 8. Male mesoscuta, dorsal view: (A) O. brunneus (inset shows transverse striae) and (B) O. relictus (inset shows longitudinal striae)." httpUri="https://zenodo.org/record/249904/files/figure.png" pageId="11" pageNumber="526">8</figureCitation>
, 12, 13, 24, 25, 26)
</paragraph>
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<subSubSection id="06D465385B22FF9A9AE0FC70FED1FB9E" pageId="11" pageNumber="526" type="reference_group">
<paragraph id="4E7136B35B22FF9A9AE0FC70FD21FC1B" blockId="11.[151,1436,969,1114]" box="[151,716,969,991]" pageId="11" pageNumber="526">
<treatmentCitationGroup id="6EDE119D5B22FF9A9AE0FC70FD21FC1B" box="[151,716,969,991]" pageId="11" pageNumber="526">
<taxonomicName id="89CE4D305B22FF9A9AE0FC70FE22FC1B" ID-CoL="CL7YN" authority="Patton 1894: 618" authorityName="Patton" authorityPageNumber="618" authorityYear="1894" box="[151,463,969,991]" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="7CBAEAA15B22FF9A9AE0FC70FEF6FC1B" box="[151,283,969,991]" italics="true" pageId="11" pageNumber="526">Atta brunnea</emphasis>
<treatmentCitation id="CF6F10A25B22FF9A9B55FC70FE22FC1B" author="Patton" box="[290,463,969,991]" page="618" pageId="11" pageNumber="526" year="1894">
<bibRefCitation id="2A5F4B425B22FF9A9B55FC70FE72FC1B" author="Patton" box="[290,415,969,991]" pageId="11" pageNumber="526" refString="Patton, W. H. (1894) Habits of the leaping-ant of southern Georgia. American Naturalist, 28, 618 - 619." type="journal article" year="1894">Patton 1894</bibRefCitation>
: 618
</treatmentCitation>
</taxonomicName>
(worker)
<collectingCountry id="36D976235B22FF9A9843FC73FD65FC1B" box="[564,648,969,991]" name="Georgia" pageId="11" pageNumber="526">Georgia</collectingCountry>
,
<collectingCountry id="36D976235B22FF9A98E5FC73FD25FC1B" box="[658,712,969,991]" name="United States of America" pageId="11" pageNumber="526">USA</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E7136B35B22FF9A9AE0FC52FED1FB9E" blockId="11.[151,1436,969,1114]" pageId="11" pageNumber="526">
<treatmentCitationGroup id="6EDE119D5B22FF9A9AE0FC52FED1FB9E" pageId="11" pageNumber="526">
<taxonomicName id="89CE4D305B22FF9A9AE0FC52FDDDFC3A" ID-CoL="6SH5N" authority="Patton, 1894" authorityName="Patton" authorityYear="1894" box="[151,560,1000,1022]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9AE0FC52FE78FC3A" box="[151,405,1000,1022]" italics="true" pageId="11" pageNumber="526">Odontomachus brunneus</emphasis>
(
<bibRefCitation id="2A5F4B425B22FF9A9BD4FC52FDC4FC3A" author="Patton" box="[419,553,1000,1022]" pageId="11" pageNumber="526" refString="Patton, W. H. (1894) Habits of the leaping-ant of southern Georgia. American Naturalist, 28, 618 - 619." type="journal article" year="1894">Patton, 1894</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B22FF9A984CFC53FD1CFC3A" author="Emery" box="[571,753,1000,1022]" page="268" pageId="11" pageNumber="526" year="1895">
<bibRefCitation id="2A5F4B425B22FF9A984CFC53FD2CFC3A" author="Emery" box="[571,705,1000,1022]" pageId="11" pageNumber="526" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery, 1895</bibRefCitation>
: 268
</treatmentCitation>
(as junior synonym of
<taxonomicName id="89CE4D305B22FF9A9996FC52FF13FBD8" ID-CoL="48QBW" authority="Guerin-Meneville, 1844" authorityName="Guerin-Meneville" authorityYear="1844" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="7CBAEAA15B22FF9A9996FC52FB37FC3A" box="[993,1242,1000,1022]" italics="true" pageId="11" pageNumber="526">Odontomachus insularis</emphasis>
<bibRefCitation id="2A5F4B425B22FF9A9E97FC52FF13FBD8" author="Guerin-Meneville" pageId="11" pageNumber="526" refString="Guerin-Meneville, F. E. (1844) Iconographie du regne animal de G. Cuvier, ou representation d'apres nature de l'une des especes les plus remarquables, et souvent non encore figurees, de chaque genre d'animaux. Insectes. J. B. Bailliere, Paris, 576 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10331" type="book" year="1844">Guérin-Méneville, 1844</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="CF6F10A25B22FF9A9B78FBBDFE24FBD8" author="Brown" box="[271,457,1030,1052]" page="103" pageId="11" pageNumber="526" year="1976">
<bibRefCitation id="2A5F4B425B22FF9A9B78FBBDFE75FBD8" author="Brown" box="[271,408,1030,1052]" pageId="11" pageNumber="526" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
, 139 (as senior synonym of
<taxonomicName id="89CE4D305B22FF9A9891FBBDFCA9FBD8" box="[742,836,1030,1052]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="7CBAEAA15B22FF9A9891FBBDFCA9FBD8" box="[742,836,1030,1052]" italics="true" pageId="11" pageNumber="526">O. fuscus</emphasis>
</taxonomicName>
);
<treatmentCitation id="CF6F10A25B22FF9A992DFBBDFB2AFBD8" author="Deyrup" box="[858,1223,1030,1052]" page="191" pageId="11" pageNumber="526" year="1985">
<bibRefCitation id="2A5F4B425B22FF9A992DFBBDFB7AFBD8" author="Deyrup" box="[858,1175,1030,1052]" pageId="11" pageNumber="526" refString="Deyrup, M., Trager, J. &amp; Carlin, N. (1985) The genus Odontomachus in the southeastern United States (Hymenoptera: Formicidae). Entomological News, 96, 188 - 195." type="journal article" year="1985">Deyrup, Trager &amp; Carlin, 1985</bibRefCitation>
: 191
</treatmentCitation>
(male).
<taxonomicName id="89CE4D305B22FF9A9AE0FB9FFDCDFBFF" authority="Stitz, 1925:115" authorityName="Stitz" authorityPageNumber="115" authorityYear="1925" box="[151,544,1061,1083]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="7CBAEAA15B22FF9A9AE0FB9FFE95FBFF" box="[151,376,1061,1083]" italics="true" pageId="11" pageNumber="526">Odontomachus fuscus</emphasis>
<treatmentCitation id="CF6F10A25B22FF9A9BF6FB9FFDCDFBFF" author="Stitz" box="[385,544,1061,1083]" page="115" pageId="11" pageNumber="526" year="1925">
<bibRefCitation id="2A5F4B425B22FF9A9BF6FB9FFE18FBFF" author="Stitz" box="[385,501,1061,1083]" pageId="11" pageNumber="526" refString="Stitz, H. (1925) Ameisen von den Philippinen, den malayischen und ozeanischen Inseln. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1923, 110 - 136." type="journal article" year="1925">Stitz, 1925</bibRefCitation>
:115
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="CF6F10A25B22FF9A985AFB9CFD01FBFF" author="Brown" box="[557,748,1061,1083]" page="103" pageId="11" pageNumber="526" year="1976">
<bibRefCitation id="2A5F4B425B22FF9A985AFB9CFD55FBFF" author="Brown" box="[557,696,1061,1083]" pageId="11" pageNumber="526" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown, 1976</bibRefCitation>
: 103
</treatmentCitation>
, 139 (unresolved junior homonym of
<taxonomicName id="89CE4D305B22FF9A9E0AFB9FFEDCFB9E" authority="Stitz, 1916: 372" authorityName="Stitz" authorityPageNumber="372" authorityYear="1916" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="7CBAEAA15B22FF9A9E0AFB9FFAB3FBFF" box="[1149,1374,1061,1083]" italics="true" pageId="11" pageNumber="526">Odontomachus fuscus</emphasis>
<treatmentCitation id="CF6F10A25B22FF9A9F1FFB9FFEDCFB9E" author="Stitz" page="372" pageId="11" pageNumber="526" year="1916">
<bibRefCitation id="2A5F4B425B22FF9A9F1FFB9FFF10FB9E" author="Stitz" pageId="11" pageNumber="526" refString="Stitz, H. (1916) Formiciden. Ergebnisse der Zweiten Deutschen Zentral-Afrika Expedition, 1, 369 - 405." type="journal article" year="1916">Stitz, 1916</bibRefCitation>
: 372
</treatmentCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="06D465385B22FF9A9AE0FB3CFF02F98B" pageId="11" pageNumber="526" type="diagnosis">
<paragraph id="4E7136B35B22FF9A9AE0FB3CFF02F98B" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="526">
<emphasis id="7CBAEAA15B22FF9A9AE0FB3CFEFCFB5B" bold="true" box="[151,273,1158,1183]" pageId="11" pageNumber="526">Diagnosis.</emphasis>
Workers of this species are distinguished from others in our region by the much finer and denser pubescence on the first gastral tergite. Males are yellow-colored and most easily confused with those of
<taxonomicName id="89CE4D305B22FF9A9FF6FB11FEF3FB22" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="haematodus">
<emphasis id="7CBAEAA15B22FF9A9FF6FB11FEF3FB22" italics="true" pageId="11" pageNumber="526">O. haematodus</emphasis>
</taxonomicName>
; those of
<taxonomicName id="89CE4D305B22FF9A9BE1FB75FDC9FB22" box="[406,548,1230,1254]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9BE1FB75FDC9FB22" box="[406,548,1230,1254]" italics="true" pageId="11" pageNumber="526">O. brunneus</emphasis>
</taxonomicName>
are clearly separated by the following characters: 1) ocelli larger, projecting beyond the posterior border of the head; 2) metasternal processes short, not elongate and spine-like; 3) posterior face of propodeum not offset from dorsal and lateral faces by distinct carina, and by numerous genitalic characters (see
<figureCitation id="D6F52A365B22FF9A9AB9FA80FEC0FA97" box="[206,301,1338,1363]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="11" pageNumber="526">Figs. 24</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9B4BFA80FEB6FA97" box="[316,347,1338,1363]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="11" pageNumber="526">25</figureCitation>
). Male
<taxonomicName id="89CE4D305B22FF9A9BC1FA81FDA9FA97" box="[438,580,1339,1363]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9BC1FA81FDA9FA97" box="[438,580,1339,1363]" italics="true" pageId="11" pageNumber="526">O. brunneus</emphasis>
</taxonomicName>
are further separated from other
<collectingCountry id="36D976235B22FF9A99BEFA81FC00FA97" box="[969,1005,1339,1363]" name="United States of America" pageId="11" pageNumber="526">US</collectingCountry>
species by the transverse mesoscutal striae, and by the anteroposteriorly slender mesosoma. Genitalia of
<taxonomicName id="89CE4D305B22FF9A99D5FAE5FBC2FAB2" box="[930,1071,1374,1398]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A99D5FAE5FBC2FAB2" box="[930,1071,1374,1398]" italics="true" pageId="11" pageNumber="526">O. brunneus</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="89CE4D305B22FF9A9FF6FAE5FEEEFA5F" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9FF6FAE5FEEEFA5F" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B22FF9A9B40FA39FE73FA5F" box="[311,414,1411,1435]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9B40FA39FE73FA5F" box="[311,414,1411,1435]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
, but are distinguished from these two species by the following characters (see
<figureCitation id="D6F52A365B22FF9A9F7FFA38FA89FA5F" box="[1288,1380,1410,1435]" captionStart="FIGURE 24" captionStartId="31.[151,250,1961,1983]" captionTargetBox="[151,1435,553,1937]" captionTargetId="figure@31.[151,1435,553,1939]" captionTargetPageId="31" captionText="FIGURE 24. Male Odontomachus abdominal sternum IX, mesal view; all scale bars 0.2 mm. (A) Odontomachus brunneus; (B) O. clarus; (C) O. desertorum; (D) O. haematodus; (E) O. relictus; (F) O. ruginodis." httpUri="https://zenodo.org/record/249918/files/figure.png" pageId="11" pageNumber="526">Figs. 24</figureCitation>
,
<figureCitation id="D6F52A365B22FF9A9F18FA38FA62FA5F" box="[1391,1423,1410,1435]" captionStart="FIGURE 25" captionStartId="32.[151,250,1512,1534]" captionTargetBox="[151,1435,193,1489]" captionTargetId="figure@32.[151,1435,193,1491]" captionTargetPageId="32" captionText="FIGURE 25. Male Odontomachus genitalic sclerites, left-hand side; arrows indicate anterior; all scale bars 0.2 mm. (A F) Paramere and volsella, medial view; (G L) penisvalva, lateral view. (A, G) Odontomachus brunneus; (B, H) O. clarus; (C, I) O. desertorum; (D, J) O. haematodus; (E, K) O. relictus; (F, L) O. ruginodis. Abbreviations: ADL = apicodorsal lobe; AP = apicoventral process; AVP = anteroventral process; BM = basimere; CU = cuspis; DI = digitus; DLC = dorsolateral carina; DS = diagonal sclerotization; SAL = subapical lamina; TM = telomere; VC = valviceps; VO = volsella; VU = valvura." httpUri="https://zenodo.org/record/249919/files/figure.png" pageId="11" pageNumber="526">25</figureCitation>
): abdominal sternum IX narrower and longer than
<taxonomicName id="89CE4D305B22FF9A98B5FA1DFCC1FA7A" box="[706,812,1446,1470]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A98B5FA1DFCC1FA7A" box="[706,812,1446,1470]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
, and broader than
<taxonomicName id="89CE4D305B22FF9A9E76FA1DFB62FA7A" box="[1025,1167,1446,1470]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9E76FA1DFB62FA7A" box="[1025,1167,1446,1470]" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
; telomere apex broadly rounded, posterodorsal margin nearly linear (apex narrower and posterodorsal margin sinuate in
<taxonomicName id="89CE4D305B22FF9A9EA9FA71FAAAFA27" box="[1246,1351,1483,1507]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9EA9FA71FAAAFA27" box="[1246,1351,1483,1507]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B22FF9A9FF6FA71FEEFF9C2" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9FF6FA71FEEFF9C2" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
); valviceps apicodorsal lobe broad (narrow in
<taxonomicName id="89CE4D305B22FF9A9963FA55FC91F9C2" box="[788,892,1518,1542]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9963FA55FC91F9C2" box="[788,892,1518,1542]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
), subapical lamina narrow (broad
<taxonomicName id="89CE4D305B22FF9A9F74FA55FA86F9C2" box="[1283,1387,1518,1542]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="clarus">
<emphasis id="7CBAEAA15B22FF9A9F74FA55FA86F9C2" box="[1283,1387,1518,1542]" italics="true" pageId="11" pageNumber="526">O. clarus</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B22FF9A9AE0F9A9FECAF9EF" box="[151,295,1555,1579]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B22FF9A9AE0F9A9FECAF9EF" box="[151,295,1555,1579]" italics="true" pageId="11" pageNumber="526">O. ruginodis</emphasis>
</taxonomicName>
).
<taxonomicName id="89CE4D305B22FF9A9B34F9A9FDB3F9EF" box="[323,606,1555,1579]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B22FF9A9B34F9A9FDB3F9EF" box="[323,606,1555,1579]" italics="true" pageId="11" pageNumber="526">Odontomachus brunneus</emphasis>
</taxonomicName>
has the proportionally shortest (dorsoventral) valviceps of the Nearctic species.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B22FF9F9AB0F9E0FDD4FD6C" lastPageId="14" lastPageNumber="529" pageId="11" pageNumber="526" type="description">
<paragraph id="4E7136B35B22FF9D9AB0F9E0FB43FED8" blockId="11.[151,1437,1158,2011]" lastBlockId="12.[151,1436,151,285]" lastPageId="12" lastPageNumber="527" pageId="11" pageNumber="526">
<emphasis id="7CBAEAA15B22FF9A9AB0F9E0FEB4F9B7" bold="true" box="[199,345,1626,1651]" pageId="11" pageNumber="526">Description.</emphasis>
Worker: HL 2.102.44, HW 1.581.92, SL 1.842.10, EL 0.340.42, ML 1.161.36, WL 2.443.00, PTH 0.961.20, PTL 0.480.52 (n=10). Entire body generally shiny except where obscured by dense pubescence; head, mesosoma, and petiole dark reddish-brown to dark brown; gaster darker brown; scape and legs brown. Head with fine, longitudinal striae covering much of the head in full-face view, striae beginning from frontal lobes and diverging toward posterior corners of head, fading at corners and sides; sides and underside of head lacking sculpture; dorsally with numerous, fine, appressed pubescence and scattered elongate, erect setae present. Pronotum with sub-circular, concentric striae that become longitudinal near rear margin; pubescence appressed, abundant; 58 elongate, erect setae present. Mesonotum and propodeum with deep transverse striae; propleuron, mesopleuron, and basalar lobe lacking sculpture; pubescence abundant dorsally, appressed. Metasternum lacking paired, elongate, spiniform processes between hind coxae. Petiole widest at base, gradually tapering apically to a short spine directed rearward; mostly lacking striae with only faint striae present near base; subpetiolar process rounded triangular; appressed pubescence present anteriorly and laterally, but mostly absent posteriorly. Gaster mostly shiny beneath pubescence, lacking striae or other strong sculpture, but with fine coriaceous sculpture (seen at high magnification); fine, appressed pubescence dense, spaces between hairs less than 1/3 the length of a hair, often overlapping one another; scattered erect, elongate setae present.
</paragraph>
<paragraph id="4E7136B35B25FF9D9AE0F8A4FE82F8B6" pageId="12" pageNumber="527">
<table id="3CCEC4135B25006E9AE0F8A4FA71F8B6" box="[151,1436,1822,1906]" gridcols="4" gridrows="3" pageId="12" pageNumber="527">
<tr id="F0FE34F15B25006E9AE0F8A4FA71F8F0" box="[151,1436,1822,1844]" gridrow="0" pageId="12" pageNumber="527">
<th id="B32F5D8D5B25006E9AE0F8A4FDD7F8F0" box="[151,570,1822,1844]" gridcol="0" gridrow="0" pageId="12" pageNumber="527">
<emphasis id="7CBAEAA15B25FF9D9AE0F8A4FEC8F8F0" bold="true" box="[151,293,1822,1844]" pageId="12" pageNumber="527">FIGURE 12.</emphasis>
<taxonomicName id="89CE4D305B25FF9D9B46F8A4FDD9F8F0" box="[305,564,1822,1844]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="527" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B25FF9D9B46F8A4FDD9F8F0" box="[305,564,1822,1844]" italics="true" pageId="12" pageNumber="527">Odontomachus brunneus</emphasis>
</taxonomicName>
,
</th>
<th id="B32F5D8D5B25006E9836F8A4FD76F8F0" box="[577,667,1822,1844]" gridcol="1" gridrow="0" pageId="12" pageNumber="527">worker:</th>
<th id="B32F5D8D5B25006E98D5F8A4FBADF8F0" box="[674,1088,1822,1844]" gridcol="2" gridrow="0" pageId="12" pageNumber="527">(A) head in full-face view, (B) lateral</th>
<th id="B32F5D8D5B25006E9E30F8A4FA71F8F0" box="[1095,1436,1822,1844]" gridcol="3" gridrow="0" pageId="12" pageNumber="527">habitus, (C) dorsal habitus, (D)</th>
</tr>
<tr id="F0FE34F15B25006E9AE0F887FA71F897" box="[151,1436,1853,1875]" gridrow="1" pageId="12" pageNumber="527">
<th id="B32F5D8D5B25006E9AE0F887FDD7F897" box="[151,570,1853,1875]" gridcol="0" gridrow="1" pageId="12" pageNumber="527">mesosoma in dorsal view, (E) mesosoma</th>
<td id="B32F5D8D5B25006E9836F887FD76F897" box="[577,667,1853,1875]" gridcol="1" gridrow="1" pageId="12" pageNumber="527">in lateral</td>
<td id="B32F5D8D5B25006E98D5F887FBADF897" box="[674,1088,1853,1875]" gridcol="2" gridrow="1" pageId="12" pageNumber="527">view, (F) petiole in lateral view, and (G)</td>
<td id="B32F5D8D5B25006E9E30F887FA71F897" box="[1095,1436,1853,1875]" gridcol="3" gridrow="1" pageId="12" pageNumber="527">first gastral tergite in dorsal view</td>
</tr>
<tr id="F0FE34F15B25006E9AE0F8E6FA71F8B6" box="[151,1436,1884,1906]" gridrow="2" pageId="12" pageNumber="527" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="B32F5D8D5B25006E9AE0F8E6FDD7F8B6" box="[151,570,1884,1906]" gridcol="0" gridrow="2" pageId="12" pageNumber="527">showing pubescence.</th>
</tr>
</table>
</paragraph>
<caption id="1AB1663B5B24FF9C9AE0F892FB92F899" ID-DOI="http://doi.org/10.5281/zenodo.249908" httpUri="https://zenodo.org/record/249908/files/figure.png" pageId="13" pageNumber="528" targetBox="[151,1436,193,1810]" targetPageId="13">
<paragraph id="4E7136B35B24FF9C9AE0F892FB92F899" blockId="13.[151,1436,1832,1885]" pageId="13" pageNumber="528">
<emphasis id="7CBAEAA15B24FF9C9AE0F892FEF2F8FA" bold="true" box="[151,287,1832,1854]" pageId="13" pageNumber="528">FIGURE 13.</emphasis>
<taxonomicName id="89CE4D305B24FF9C9B51F892FDCEF8FA" box="[294,547,1832,1854]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B24FF9C9B51F892FDCEF8FA" box="[294,547,1832,1854]" italics="true" pageId="13" pageNumber="528">Odontomachus brunneus</emphasis>
</taxonomicName>
, male: (A) head in full-face view, (B) head in lateral view, (C) head in dorsal view, (D) lateral habitus, (E) mesosoma in lateral view, (F) dorsal habitus, and (G) mesosoma in dorsal view.
</paragraph>
</caption>
<paragraph id="4E7136B35B24FF9F9AB0F83EFA82FD9B" blockId="13.[151,1437,1924,2021]" lastBlockId="14.[151,1437,151,1544]" lastPageId="14" lastPageNumber="529" pageId="13" pageNumber="528">Male: HL 1.001.08, HW 1.241.34, SL 0.180.22, EL 0.680.76, EW 0.400.44, OL 0.24, OES 0.16, WL 2.522.76, PTH 0.720.88, PTL 0.460.54, FWL 5.105.65 (n=5). Head, mesosoma, and petiole generally shiny except where obscured by dense pubescence; head, meso- and metasoma, legs, scape and pedicel yellow to yellowish-brown, remainder of funiculus brown. Head and mesosoma with abundant fine, white pubescence except on anepisternum where pubescence is mostly absent. Eyes extremely large, maximum diameter of each eye at least 70% of the length of the head in full-face view. Ocelli large, the length of each ocellus wider than distance between lateral ocellus and eye margin; in full-face view, lateral ocelli protrude beyond posterior border of head. Mesosoma: pronotum lacking sculpture; mesoscutum with fine transversely arcuate striae anteriorly, striae becoming longitudinal posteriorly; mesoscutellum raised and convex, lacking sculpture; propodeum with weak to moderately strong transverse striae laterally, and especially posterodorsally; mesopleuron mostly lacking striae. Petiole bluntly rounded apically, with rounded triangular subpetiolar process anteriorly; densely pubescent anteriorly and laterally, but reduced pubescence posteriorly. Disc of abdominal sternum IX trapezoidal, breadth slightly greater than length; posterior lobe length approximately twice maximum lobe width, basal half narrowed. Telomeral apex broadly rounded; telomere length distinctly greater than height; valviceps ventral apex strongly produced, narrow; valviceps apical margin broadly concave; vertical portion of dorsolateral carina and lateral margin of subapical lamina meeting at an angle; apicodorsal lobe of valviceps broad; subapical lamina narrow.</paragraph>
<paragraph id="4E7136B35B27FF9F9AB0FDD6FDD4FD6C" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">Queen: No specimens measured, but similar to workers in color and general appearance except slightly larger, with mesosoma developed for wings.</paragraph>
</subSubSection>
<subSubSection id="06D465385B27FF9F9AB0FD0EFED8FBD4" pageId="14" pageNumber="529" type="distribution">
<paragraph id="4E7136B35B27FF9F9AB0FD0EFED8FBD4" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">
<emphasis id="7CBAEAA15B27FF9F9AB0FD0EFDF2FD09" bold="true" box="[199,543,692,717]" pageId="14" pageNumber="529">
Distribution (
<figureCitation id="D6F52A365B27FF9F9B18FD0EFE26FD09" box="[367,459,692,717]" captionStart="FIGURE 26" captionStartId="33.[151,250,1854,1876]" captionTargetBox="[151,1435,1060,1832]" captionTargetId="figure@33.[151,1435,1060,1833]" captionTargetPageId="33" captionText="FIGURE 26. Map of the southern United States (by county) and northern Mexico (by state) showing the known occurrences of Odontomachus brunneus (black square), O. clarus (black circle), O. desertorum (magenta square), O. haematodus (yellow square), O. relictus (red triangle), and O. ruginodis (orange and white circle)." httpUri="https://zenodo.org/record/249920/files/figure.png" pageId="14" pageNumber="529">Fig. 26</figureCitation>
).
<collectingCountry id="36D976235B27FF9F9B90FD0EFDF2FD09" box="[487,543,692,717]" name="United States of America" pageId="14" pageNumber="529">USA</collectingCountry>
</emphasis>
. Alabama: Baldwin and Houston Counties. Florida: Alachua,
<collectingCountry id="36D976235B27FF9F9F65FD0EFABAFD09" box="[1298,1367,692,717]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="529">Baker</collectingCountry>
, Bay, Bradford, Broward, Citrus, Clay, Collier,
<collectingCountry id="36D976235B27FF9F9818FD6DFD0DFD34" box="[623,736,727,752]" name="United States of America" pageId="14" pageNumber="529">Columbia</collectingCountry>
, Dade, De Soto, Duval, Franklin, Gadsden, Gilchrist, Glades, Hamilton, Hendry, Hernando, Highlands, Hillsborough, Indian River, Jackson, Jefferson, Lake, Lee, Leon, Levy, Liberty, Madison, Marion, Martin, Monroe, Nassau, Okeechobee, Orange, Osceola, Palm Beach, Pasco, Polk, Putnam, Sarasota, St. Lucie, Sumter, Taylor, Volusia, Wakulla, and Walton Counties.
<collectingCountry id="36D976235B27FF9F9E1DFCFEFB2EFC99" box="[1130,1219,836,861]" name="Georgia" pageId="14" pageNumber="529">Georgia</collectingCountry>
: Chatham, Clinch, Dougherty, Glynn, Lowndes, and Ware Counties. Mississippi: Harrison County. According to
<bibRefCitation id="2A5F4B425B27FF9F9EBEFCD2FF09FC60" author="Deyrup" pageId="14" pageNumber="529" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
, the occurrence of this species in the Caribbean region and Central and South
<collectingCountry id="36D976235B27FF9F9E05FC36FB3FFC61" box="[1138,1234,908,933]" name="United States of America" pageId="14" pageNumber="529">America</collectingCountry>
is unclear due to historical misidentifications and nomenclatural changes. Distributional information from
<bibRefCitation id="2A5F4B425B27FF9F9E0AFC0AFA71FC0C" author="Deyrup" box="[1149,1436,943,968]" pageId="14" pageNumber="529" refString="Deyrup, M. &amp; Cover, S. (2004) A new species of Odontomachus ant (Hymenoptera: Formicidae) from inland ridges of Florida, with a key to Odontomachus of the United States. Florida Entomologist, 87, 136 - 144. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0136: ansooa] 2.0. co; 2" type="journal article" year="2004">Deyrup and Cover (2004)</bibRefCitation>
and examination of specimens from the ABS, MEM, UGA, USNM, and personal collections of Dan Suitor and Doug Booher.
</paragraph>
</subSubSection>
<subSubSection id="06D465385B27FF9F9AB0FBA6FB63F9CC" pageId="14" pageNumber="529" type="discussion">
<paragraph id="4E7136B35B27FF9F9AB0FBA6FD29FB2C" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">
<emphasis id="7CBAEAA15B27FF9F9AB0FBA6FEA1FBF1" bold="true" box="[199,332,1052,1077]" pageId="14" pageNumber="529">Discussion.</emphasis>
<taxonomicName id="89CE4D305B27FF9F9B23FBA6FD82FBF0" box="[340,623,1052,1076]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9B23FBA6FD82FBF0" box="[340,623,1052,1076]" italics="true" pageId="14" pageNumber="529">Odontomachus brunneus</emphasis>
</taxonomicName>
appears to be restricted to the southeastern
<collectingCountry id="36D976235B27FF9F9E10FBA6FB63FBF0" box="[1127,1166,1052,1076]" name="United States of America" pageId="14" pageNumber="529">US</collectingCountry>
. Previous records of
<taxonomicName id="89CE4D305B27FF9F9FF6FBA6FF12FB9C" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9FF6FBA6FF12FB9C" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
from the Caribbean, and Central and South
<collectingCountry id="36D976235B27FF9F9885FB85FCBEFB9C" box="[754,851,1087,1112]" name="United States of America" pageId="14" pageNumber="529">America</collectingCountry>
(
<bibRefCitation id="2A5F4B425B27FF9F9915FBFAFC1DFB9C" author="Brown" box="[866,1008,1087,1112]" pageId="14" pageNumber="529" refString="Brown, W. L. (1976) Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters Genus Odontomachus. Stutia Entomologica, 19, 67 - 171." type="journal article" year="1976">Brown 1976</bibRefCitation>
) all appear to be of
<taxonomicName id="89CE4D305B27FF9F9EA3FBFAFA8CFB9C" box="[1236,1377,1088,1112]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B27FF9F9EA3FBFAFA8CFB9C" box="[1236,1377,1088,1112]" italics="true" pageId="14" pageNumber="529">O. ruginodis</emphasis>
</taxonomicName>
. The situation was clarified by
<bibRefCitation id="2A5F4B425B27FF9F9BBAFBDFFD2FFBB9" box="[461,706,1124,1149]" pageId="14" pageNumber="529" refString="Deyrup, M., Trager, J. &amp; Carlin, N. (1985) The genus Odontomachus in the southeastern United States (Hymenoptera: Formicidae). Entomological News, 96, 188 - 195." type="journal article">
Deyrup
<emphasis id="7CBAEAA15B27FF9F9859FBDFFD86FBB8" box="[558,619,1124,1148]" italics="true" pageId="14" pageNumber="529">et al.</emphasis>
(1985)
</bibRefCitation>
who recognized the distinction between
<taxonomicName id="89CE4D305B27FF9F9EDAFBDEFAADFBB8" box="[1197,1344,1124,1148]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="ruginodis">
<emphasis id="7CBAEAA15B27FF9F9EDAFBDEFAADFBB8" box="[1197,1344,1124,1148]" italics="true" pageId="14" pageNumber="529">O. ruginodis</emphasis>
</taxonomicName>
and
<taxonomicName id="89CE4D305B27FF9F9FF6FBDEFF12FB64" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9FF6FBDEFF12FB64" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
and revived the former from synonymy with the latter. Browns statement (1976) that
<taxonomicName id="89CE4D305B27FF9F9EB5FB32FAA1FB64" box="[1218,1356,1160,1184]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9EB5FB32FAA1FB64" box="[1218,1356,1160,1184]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
is well adapted to “marginal habitats is consistent with the ecology of
<emphasis id="7CBAEAA15B27FF9F992FFB16FC00FB00" box="[856,1005,1196,1220]" italics="true" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F992FFB16FC04FB00" box="[856,1001,1196,1220]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="ruginodis">O. ruginodis</taxonomicName>
,
</emphasis>
whereas
<taxonomicName id="89CE4D305B27FF9F9E20FB16FB0CFB00" box="[1111,1249,1196,1220]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9E20FB16FB0CFB00" box="[1111,1249,1196,1220]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
in the Southeast is generally found in undisturbed natural habitats.
</paragraph>
<paragraph id="4E7136B35B27FF9F9AB0FB4FFB63F9CC" blockId="14.[151,1437,151,1544]" pageId="14" pageNumber="529">
In the
<collectingCountry id="36D976235B27FF9F9B64FB4EFED4FAC8" box="[275,313,1268,1292]" name="United States of America" pageId="14" pageNumber="529">US</collectingCountry>
,
<taxonomicName id="89CE4D305B27FF9F9B30FB4EFE39FAC8" box="[327,468,1268,1292]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9B30FB4EFE39FAC8" box="[327,468,1268,1292]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
occurs in a wide variety of natural habitats including flatwoods, mesic forests, pine savannas, swamp forests, oak-pine scrub, upland scrub, sandhills, bayheads, edges of seasonal ponds, and elevated tussocks. Nests of
<taxonomicName id="89CE4D305B27FF9F9B1AFA86FE17FA90" box="[365,506,1340,1364]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="7CBAEAA15B27FF9F9B1AFA86FE17FA90" box="[365,506,1340,1364]" italics="true" pageId="14" pageNumber="529">O. brunneus</emphasis>
</taxonomicName>
have been found in leaf litter, rotting logs, at tree bases, and in open to partially covered sandy areas. Nest architecture has been explored and discussed by
<bibRefCitation id="2A5F4B425B27FF9F9E50FADAFA75FABC" author="Cerquera" box="[1063,1432,1375,1400]" pageId="14" pageNumber="529" refString="Cerquera, L. M &amp; Tschinkel, W. R. (2010) The nest architecture of the ant Odontomachus brunneus. Journal of Insect Science, 10, 1 - 12. http: // dx. doi. org / 10.1673 / 031.010.6401" type="journal article" year="2010">Cerquera and Tschinkel (2010)</bibRefCitation>
. Workers occasionally forage during the day, but are more active at night. Upon colony disturbance, workers are not aggressive, but instead quickly retreat or vacate the nest. This is in sharp contrast to the aggressive defensive stinging behavior of
<emphasis id="7CBAEAA15B27FF9F9BFFFA76FDD1FA20" box="[392,572,1484,1508]" italics="true" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F9BFFFA76FDD5FA20" box="[392,568,1484,1508]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">O. haematodus</taxonomicName>
.
</emphasis>
Alates have been collected from May through December. This broad time frame for alate activity contrasts sharply with the early summer activity of
<emphasis id="7CBAEAA15B27FF9F99AAFA4AFB63F9CC" box="[989,1166,1520,1544]" italics="true" pageId="14" pageNumber="529">
<taxonomicName id="89CE4D305B27FF9F99AAFA4AFB67F9CC" box="[989,1162,1520,1544]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="haematodus">O. haematodus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>