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<document id="11D84FE2F9DE9B1372657EF64375058C" ID-DOI="10.11646/zootaxa.3619.2.5" ID-GBIF-Dataset="549f99ee-2153-4d61-a953-8def30c8159d" ID-ISSN="1175-5326" ID-Zenodo-Dep="216580" ID-ZooBank="DBB8DB78-09DF-4F68-AFB1-AAC8EBC95DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460577749427" checkinUser="plazi" docAuthor="Ren, Li, Alonso-Zarazaga, Miguel A. &amp; Zhang, Runzhi" docDate="2013" docId="C62C2621100AC045FF10FA86FB6C6E1F" docLanguage="en" docName="zt03619p182.pdf" docOrigin="Zootaxa 3619 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geotragus brevidens Ren, Alonso-Zarazaga &amp; Zhang, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="170" masterDocId="3A155E59100CC04CFF87FFA8FF95686B" masterDocTitle="Revision of the Chinese Geotragus Schoenherr with description of three new species (Coleoptera: Curculionidae: Entiminae)" masterLastPageNumber="182" masterPageNumber="161" pageNumber="167" updateTime="1698323549919" updateUser="plazi">
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<mods:title id="62ADDF87C7B9942C096E4FE63A6DA1FE">Revision of the Chinese Geotragus Schoenherr with description of three new species (Coleoptera: Curculionidae: Entiminae)</mods:title>
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<mods:namePart id="C3A2D7F5BD88D293E65B846045DFA980">Ren, Li</mods:namePart>
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<mods:namePart id="F4158213828148D159A869039DEFED67">Alonso-Zarazaga, Miguel A.</mods:namePart>
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<mods:namePart id="19D96F8433A728981808DA48291E3D76">Zhang, Runzhi</mods:namePart>
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<mods:title id="98D700DFB76973D13802C2ABA478E37B">Zootaxa</mods:title>
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<mods:date id="DEB23AA1ECBC03B8B6726C11BBF6236C">2013</mods:date>
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<treatment id="C62C2621100AC045FF10FA86FB6C6E1F" ID-DOI="http://doi.org/10.5281/zenodo.6158917" ID-GBIF-Taxon="119560658" ID-Zenodo-Dep="6158917" LSID="urn:lsid:plazi:treatment:C62C2621100AC045FF10FA86FB6C6E1F" httpUri="http://treatment.plazi.org/id/C62C2621100AC045FF10FA86FB6C6E1F" lastPageId="9" lastPageNumber="170" pageId="6" pageNumber="167">
<subSubSection id="069FC4BC100AC04AFF10FA86FE936D01" pageId="6" pageNumber="167" type="nomenclature">
<paragraph id="4E3A9737100AC04AFF10FA86FE706D2D" blockId="6.[151,485,1326,1386]" box="[151,485,1326,1351]" pageId="6" pageNumber="167">
<heading id="1572205B100AC04AFF10FA86FE706D2D" bold="true" box="[151,485,1326,1351]" fontSize="10" level="2" pageId="6" pageNumber="167" reason="2">
<emphasis id="7CF14B25100AC04AFF10FA86FE706D2D" bold="true" box="[151,485,1326,1351]" pageId="6" pageNumber="167">
<taxonomicName id="8985ECB4100AC04AFF10FA86FE116D2C" ID-CoL="3FT5R" box="[151,388,1326,1351]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="brevidens" status="sp. nov.">
<emphasis id="7CF14B25100AC04AFF10FA86FE116D2C" bold="true" box="[151,388,1326,1351]" italics="true" pageId="6" pageNumber="167">Geotragus brevidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67C2F65E100AC04AFE0CFA86FE706D2D" box="[395,485,1326,1350]" pageId="6" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4E3A9737100AC04AFF10FAFAFE936D01" blockId="6.[151,485,1326,1386]" box="[151,262,1362,1386]" pageId="6" pageNumber="167">
(
<figureCitation id="D6BE8BB2100AC04AFF18FAFAFF766D01" box="[159,227,1362,1386]" captionStart="FIGURE 3" captionStartId="7.[151,250,927,950]" captionTargetBox="[152,1436,301,906]" captionTargetId="figure@7.[151,1436,301,906]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Geotragus brevidens sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216583/files/figure.png" pageId="6" pageNumber="167">Fig. 3</figureCitation>
,
<figureCitation id="D6BE8BB2100AC04AFF68FAFAFF6B6D01" box="[239,254,1362,1386]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="6" pageNumber="167">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC100AC04AFF10FA32FBDE6D91" pageId="6" pageNumber="167" type="diagnosis">
<paragraph id="4E3A9737100AC04AFF10FA32FBDE6D91" blockId="6.[151,1436,1434,2034]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF10FA32FE846DD8" bold="true" box="[151,273,1434,1459]" pageId="6" pageNumber="167">Diagnosis.</emphasis>
Similar to
<taxonomicName id="8985ECB4100AC04AFE15FA33FD176DD9" authority="Chen" authorityName="Chen" box="[402,642,1434,1458]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="7CF14B25100AC04AFE15FA33FE366DD9" box="[402,419,1435,1458]" italics="true" pageId="6" pageNumber="167">G</emphasis>
.
<emphasis id="7CF14B25100AC04AFE35FA32FDA86DD9" box="[434,573,1434,1458]" italics="true" pageId="6" pageNumber="167">tuberculatus</emphasis>
Chen
</taxonomicName>
, but differs by the following characters: scales sparsely distributed on dorsal surface, eyes in frontal view moderately flat (vs. strongly convex in
<taxonomicName id="8985ECB4100AC04AFC5EFA17FB176DBD" box="[985,1154,1470,1494]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="7CF14B25100AC04AFC5EFA17FC7F6DBD" box="[985,1002,1471,1494]" italics="true" pageId="6" pageNumber="167">G</emphasis>
.
<emphasis id="7CF14B25100AC04AFC70FA16FB176DBD" box="[1015,1154,1470,1494]" italics="true" pageId="6" pageNumber="167">tuberculatus</emphasis>
</taxonomicName>
), epistome broad and the elytral interstria 3 only slightly elevated at base, without three consecutive tubercles.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC100AC045FF4EF9ADFA136C3C" lastPageId="9" lastPageNumber="170" pageId="6" pageNumber="167" type="description">
<paragraph id="4E3A9737100AC04AFF4EF9ADFB0B6E29" blockId="6.[151,1436,1434,2034]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF4EF9ADFD8F6E75" bold="true" box="[201,538,1541,1566]" pageId="6" pageNumber="167">Description. Holotype-male.</emphasis>
<emphasis id="7CF14B25100AC04AFDA4F9AFFD506E75" box="[547,709,1543,1566]" italics="true" pageId="6" pageNumber="167">Measurements</emphasis>
(in mm): Standard length: 5.90; pronotal length: 1.65; pronotal width: 1.91; elytral length: 4.30; elytral width: 2.55; rostral length: 0.96; rostral width: 0.84.
</paragraph>
<paragraph id="4E3A9737100AC04AFF4EF9E6FB6C6F09" blockId="6.[151,1436,1434,2034]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF4EF9E6FE066E0D" box="[201,403,1614,1638]" italics="true" pageId="6" pageNumber="167">Habitus and color</emphasis>
(
<figureCitation id="D6BE8BB2100AC04AFE25F9E6FE766E0D" box="[418,483,1614,1638]" captionStart="FIGURE 3" captionStartId="7.[151,250,927,950]" captionTargetBox="[152,1436,301,906]" captionTargetId="figure@7.[151,1436,301,906]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Geotragus brevidens sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216583/files/figure.png" pageId="6" pageNumber="167">Fig. 3</figureCitation>
A, 3B): Integument reddish brown to dark red, antennae and legs reddish brown; with coppery metallic green to greyish brown scales, scales irregularly distributed, moderately sparse, tessellate, but not contiguous; scales on dorsal surface of rostrum sparse, round to oval, on lateral part of rostrum lanceolate; posterior half of antennal scrobes glabrous; pronotum with round scales; scales on ventrites dense, gradually narrowing from ventrite 1, so ventrite 5 with lanceolate scales; legs densely covered by scales; body with recumbent to subrecumbent sparse fine setae; rostrum covered with recumbent fine setae; setae of antennal scape and desmomeres 17 sparse, moderately long and fine; setae of interstriae fine, sparse and subrecumbent; setae on ventral surface fine and dense; setae of tibia long and straight, femora with subrecumbent fine setae.
</paragraph>
<paragraph id="4E3A9737100AC04AFF4EF8C6FADE6FC1" blockId="6.[151,1436,1434,2034]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF4EF8C6FE936FED" box="[201,262,1902,1926]" italics="true" pageId="6" pageNumber="167">Head</emphasis>
(
<figureCitation id="D6BE8BB2100AC04AFE92F8C6FEC26FED" box="[277,343,1902,1926]" captionStart="FIGURE 3" captionStartId="7.[151,250,927,950]" captionTargetBox="[152,1436,301,906]" captionTargetId="figure@7.[151,1436,301,906]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Geotragus brevidens sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216583/files/figure.png" pageId="6" pageNumber="167">Fig. 3</figureCitation>
C): Moderately flat, dorsal surface smooth, punctures small and sparse; eyes moderately flat, with a shallow circumocular sulcus; frons rather convex, much higher than dorsal part of rostrum in lateral view.
</paragraph>
<paragraph id="4E3A9737100AC04BFF4EF81FFAC6689C" blockId="6.[151,1436,1434,2034]" lastBlockId="7.[151,1436,151,248]" lastPageId="7" lastPageNumber="168" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF4EF81FFEBD6FA5" box="[201,296,1975,1998]" italics="true" pageId="6" pageNumber="167">Rostrum</emphasis>
(
<figureCitation id="D6BE8BB2100AC04AFEBFF81EFEE96FA5" box="[312,380,1974,1998]" captionStart="FIGURE 3" captionStartId="7.[151,250,927,950]" captionTargetBox="[152,1436,301,906]" captionTargetId="figure@7.[151,1436,301,906]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Geotragus brevidens sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216583/files/figure.png" pageId="6" pageNumber="167">Fig. 3</figureCitation>
C): In dorsal view, 1.14× length of wide, apex narrower than base; base narrower than frons; dorsal surface depressed in median area of rostrum; with a narrow and deep median sulcus, reaching vertex; epistome broad, posterior angle of epistome 90°, not forming a distinct carina; mandible scars elongate-oval; in dorsal view ventral margin of scrobes visible from apex to antennal insertion; in lateral view, lateral surface between eyes and antennal scrobes flat; lateral surface with a shallow sulcus near ventral margin of rostrum.
</paragraph>
<caption id="1AFAC7BF100BC04BFF10FC37FEB36BBF" httpUri="https://zenodo.org/record/216583/files/figure.png" pageId="7" pageNumber="168" targetBox="[152,1436,301,906]" targetPageId="7">
<paragraph id="4E3A9737100BC04BFF10FC37FEB36BBF" blockId="7.[151,1437,927,980]" pageId="7" pageNumber="168">
<emphasis id="7CF14B25100BC04BFF10FC37FE836BDE" bold="true" box="[151,278,927,950]" pageId="7" pageNumber="168">FIGURE 3.</emphasis>
Habitus of
<taxonomicName id="8985ECB4100BC04BFE10FC09FDF96BDD" box="[407,620,928,950]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="168" phylum="Arthropoda" rank="species" species="brevidens" status="sp. nov.">
<emphasis id="7CF14B25100BC04BFE10FC09FDF96BDD" box="[407,620,928,950]" italics="true" pageId="7" pageNumber="168">Geotragus brevidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67C2F65E100BC04BFDF0FC08FD5F6BDE" box="[631,714,928,949]" pageId="7" pageNumber="168" rank="species">
<emphasis id="7CF14B25100BC04BFDF0FC08FD5F6BDE" bold="true" box="[631,714,928,949]" pageId="7" pageNumber="168">sp. nov.</emphasis>
</taxonomicNameLabel>
, male, holotype:
<emphasis id="7CF14B25100BC04BFC02FC08FC026BDE" bold="true" box="[901,919,928,949]" pageId="7" pageNumber="168">A</emphasis>
dorsal view
<emphasis id="7CF14B25100BC04BFBA3FC08FBA06BDE" bold="true" box="[1060,1077,928,949]" pageId="7" pageNumber="168">B</emphasis>
lateral view
<emphasis id="7CF14B25100BC04BFB44FC37FB406BDD" bold="true" box="[1219,1237,927,950]" pageId="7" pageNumber="168">C</emphasis>
head, dorsal view. Scales: 1 mm.
</paragraph>
</caption>
<paragraph id="4E3A9737100BC04BFF4EFBA9FD956CA6" blockId="7.[151,1437,1025,2021]" pageId="7" pageNumber="168">
<emphasis id="7CF14B25100BC04BFF4EFBA9FEA66C72" box="[201,307,1025,1049]" italics="true" pageId="7" pageNumber="168">Antennae</emphasis>
: Scape short, stout, not reaching middle of eyes at rest, 0.80× length of funicle; desmomere 1 elongate clavate, strongly widened towards apex, apex rather stout, 1.21× length of desmomere 2; desmomere 3 1.17× length of desmomere 4, elongate oval, 0.73× length of 2; desmomeres 46 equal in length, moniliform, shorter than desmomere 3; desmomere 7 1.33× length of and much wider than 6, moniliform; pubescence of funicle similar throughout; club segment 1 1.18× length of 2, segment 2 0.89× length of 3; segment 3 with a marked
<emphasis id="7CF14B25100BC04BFF75FB1DFEDE6CA6" box="[242,331,1205,1229]" italics="true" pageId="7" pageNumber="168">annulus</emphasis>
beyond middle.
</paragraph>
<paragraph id="4E3A9737100BC04BFF4EFB72FE296D36" blockId="7.[151,1437,1025,2021]" pageId="7" pageNumber="168">
<emphasis id="7CF14B25100BC04BFF4EFB72FEAD6C9A" box="[201,312,1242,1265]" italics="true" pageId="7" pageNumber="168">Pronotum</emphasis>
: 0.86× as long as wide, transverse; anterior and posterior margin truncate; disc with a fine, shallow median longitudinal groove; posterior margin distinct carinate, linear; dorsal surface of pronotum with slightly sparse, large, round granules, space between granules convex, each granule with one recumbent fine seta on top; postocular vibrissae short.
</paragraph>
<paragraph id="4E3A9737100BC04BFF4EFAC1FCE06DEA" blockId="7.[151,1437,1025,2021]" box="[201,885,1385,1409]" pageId="7" pageNumber="168">
<taxonomicName id="8985ECB4100BC04BFF4EFAC1FE126DEA" authority="Small" authorityName="Small" box="[201,391,1385,1409]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="7" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="7CF14B25100BC04BFF4EFAC1FEA26DEA" box="[201,311,1385,1409]" italics="true" pageId="7" pageNumber="168">Scutellum</emphasis>
: Small
</taxonomicName>
but distinct, reddish brown, apex V-shaped.
</paragraph>
<paragraph id="4E3A9737100BC04BFF4EFA25FBB26E16" blockId="7.[151,1437,1025,2021]" pageId="7" pageNumber="168">
<emphasis id="7CF14B25100BC04BFF4EFA25FE9A6DCE" box="[201,271,1421,1445]" italics="true" pageId="7" pageNumber="168">Elytra</emphasis>
: Base truncate; in lateral view moderately convex; in dorsal view, lateral margins roundly diverging from base to basal 1/4, then subparallel to 2/3 and strongly narrowed towards apex, apex nearly acute; odd interstriae slightly raised than even ones; interstria 1 moderately raised along basal 1/8, forming a longitudinal crest, strongly raised at declivity, forming a prominent longitudinal crest, top of crest sub-rectangular in lateral view; interstria 3 distinctly raised and expanded at base, much wider than remaining interstriae; interstria 5 gradually raised from base to declivity, most strongly raised at declivity, forming longitudinal crest; interstriae 3 and 9 coalescing, raised at declivity forming a V-shaped tubercle on each elytron.
</paragraph>
<paragraph id="4E3A9737100BC04BFF4EF921FB266E82" blockId="7.[151,1437,1025,2021]" pageId="7" pageNumber="168">
<emphasis id="7CF14B25100BC04BFF4EF921FEA66ECA" box="[201,307,1673,1697]" italics="true" pageId="7" pageNumber="168">Abdomen</emphasis>
(
<figureCitation id="D6BE8BB2100BC04BFEC5F921FE126ECA" box="[322,391,1673,1697]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="7" pageNumber="168">Fig. 4</figureCitation>
A): Surface of ventrite 1 depressed at middle; suture I slightly bisinuate; surface of ventrite 2 moderately flat medially, without apical transverse impression; mesal length of ventrite 1 1.42× length of 2, ventrite 2 as long as ventrite 3 and 4 combined; ventrite 5 broadly parabolic, with flat surface.
</paragraph>
<paragraph id="4E3A9737100BC04BFF4EF95DFA086F12" blockId="7.[151,1437,1025,2021]" pageId="7" pageNumber="168">
<emphasis id="7CF14B25100BC04BFF4EF95DFEEA6F66" box="[201,383,1781,1805]" italics="true" pageId="7" pageNumber="168">Metendosternite</emphasis>
(
<figureCitation id="D6BE8BB2100BC04BFE16F95DFE4D6F66" box="[401,472,1781,1805]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="7" pageNumber="168">Fig. 4</figureCitation>
B): Stalk 0.33× length of furcal arms, trapezoidal and transverse, ventral margin 2× length of dorsal width of stalk, anterior part of longitudinal flange as long as posterior; hemiductus robust, wide and triangular, extending to sheath by a tunnel; anterior tendons positioned near apex of furcal arms, opposite to hemiductus; furcal arms robust, apically slightly bifurcate, diverging at nearly 45° in relation to longitudinal flange.
</paragraph>
<paragraph id="4E3A9737100BC045FF4EF82EFD3168BF" blockId="7.[151,1437,1025,2021]" lastBlockId="9.[151,1437,151,1652]" lastPageId="9" lastPageNumber="170" pageId="7" pageNumber="168">
<emphasis id="7CF14B25100BC04BFF4EF82EFF6B6FF6" box="[201,254,1926,1949]" italics="true" pageId="7" pageNumber="168">Legs</emphasis>
: Protibiae bent inwards at apical 1/3; inner margin of protibiae moderately bisinuate, with ten moderately sharp and small teeth, each tooth with a spiniform seta just behind it, length of the longest denticle less than 1/10 width of tibia at the same point, outer margin slightly arcuate inwards; teeth of meso- and metatibiae fewer and larger than those of protibiae; protibial mucro small, distinct, covered by setae; tarsi short, broad; tarsomere 1 wider and 2× length of 2; tarsomere 5 long and stout.
</paragraph>
<caption id="1AFAC7BF1004C044FF10F8C6FE2E6F8A" httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="8" pageNumber="169" targetBox="[278,1321,222,1868]" targetPageId="8">
<paragraph id="4E3A97371004C044FF10F8C6FE2E6F8A" blockId="8.[151,1436,1902,2017]" pageId="8" pageNumber="169">
<emphasis id="7CF14B251004C044FF10F8C6FE876FEF" bold="true" box="[151,274,1902,1925]" pageId="8" pageNumber="169">FIGURE 4.</emphasis>
<taxonomicName id="8985ECB41004C044FE9EF8D8FE7E6FEE" box="[281,491,1903,1925]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="169" phylum="Arthropoda" rank="species" species="brevidens" status="sp. nov.">
<emphasis id="7CF14B251004C044FE9EF8D8FE7E6FEE" box="[281,491,1903,1925]" italics="true" pageId="8" pageNumber="169">Geotragus brevidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67C2F65E1004C044FE75F8C7FDD76FEF" box="[498,578,1903,1924]" pageId="8" pageNumber="169" rank="species">
<emphasis id="7CF14B251004C044FE75F8C7FDD76FEF" bold="true" box="[498,578,1903,1924]" pageId="8" pageNumber="169">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="7CF14B251004C044FDD7F8C7FDF76FEF" bold="true" box="[592,610,1903,1924]" pageId="8" pageNumber="169">A</emphasis>
male ventrites, ventral view
<emphasis id="7CF14B251004C044FC0FF8C7FC0C6FEF" bold="true" box="[904,921,1903,1924]" pageId="8" pageNumber="169">B</emphasis>
male metendosternite, dorsal view
<emphasis id="7CF14B251004C044FA86F8C6FA866FEE" bold="true" box="[1281,1299,1902,1925]" pageId="8" pageNumber="169">C</emphasis>
male sternite VIII and IX, dorsal view
<emphasis id="7CF14B251004C044FE1DF826FE396FC8" bold="true" box="[410,428,1934,1955]" pageId="8" pageNumber="169">D</emphasis>
penis, dorsal view
<emphasis id="7CF14B251004C044FDF4F826FD116FC8" bold="true" box="[627,644,1934,1955]" pageId="8" pageNumber="169">E</emphasis>
penis, lateral view
<emphasis id="7CF14B251004C044FCCBF826FCCE6FC8" bold="true" box="[844,859,1934,1955]" pageId="8" pageNumber="169">F</emphasis>
tegmen, dorsal view
<emphasis id="7CF14B251004C044FBBFF825FBD96FCF" bold="true" box="[1080,1100,1933,1956]" pageId="8" pageNumber="169">G</emphasis>
female ventrites, ventral view
<emphasis id="7CF14B251004C044FA0FF826FA096FC8" bold="true" box="[1416,1436,1934,1955]" pageId="8" pageNumber="169">H</emphasis>
sternite VIII and female genitalia, lateral view
<emphasis id="7CF14B251004C044FDFCF805FD106FA9" bold="true" box="[635,645,1965,1986]" pageId="8" pageNumber="169">I</emphasis>
coxites and styli, dorsal view
<emphasis id="7CF14B251004C044FC47F805FC586FA9" bold="true" box="[960,973,1965,1986]" pageId="8" pageNumber="169">J</emphasis>
bursal sclerites, lateral view
<emphasis id="7CF14B251004C044FB7BF805FA856FA9" bold="true" box="[1276,1296,1965,1986]" pageId="8" pageNumber="169">K</emphasis>
spermatheca, lateral view. Scales: 0.5 mm.
</paragraph>
</caption>
<paragraph id="4E3A97371005C045FF4EFF48FD9F6A57" blockId="9.[151,1437,151,1652]" pageId="9" pageNumber="170">
<emphasis id="7CF14B251005C045FF4EFF48FDB56893" box="[201,544,224,248]" italics="true" pageId="9" pageNumber="170">Male genitalia and terminalia</emphasis>
: Sternite VIII (
<figureCitation id="D6BE8BB21005C045FD53FF77FC8C689C" box="[724,793,223,247]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
C) divided into two hemisternites, transversely oriented, crescent shape, with discal part much wider than lateral; each hemisternite laterally acuminate, with anterior margin strongly sinuate, and posterior margin arcuate; membrane between sternite VIII and IX with two very small, elongate triangular sclerotizations. Sternite IX (
<figureCitation id="D6BE8BB21005C045FD78FEE4FCD1690F" box="[767,836,332,356]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
C) with basal plate bilobed, each lobe sickle-shaped; spiculum gastrale almost as long as penis, slender, almost straight, anterior 1/3 slightly curved. Penis (
<figureCitation id="D6BE8BB21005C045FA89FEC7FAC769EC" box="[1294,1362,367,391]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
D, 4E) in dorsal view with tube
<quantity id="897D3AD21005C045FE46FE3CFDBC69C7" box="[449,553,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.58" pageId="9" pageNumber="170" unit="mm" value="2.58">2.58 mm</quantity>
in length, ca. 6.0× length of wide, temones
<quantity id="897D3AD21005C045FBBDFE3CFB3769C7" box="[1082,1186,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="9" pageNumber="170" unit="mm" value="1.26">1.26 mm</quantity>
long; lateral margins subparallel, straightly converging towards apex, apex ogival; in lateral view moderately curved, more strongly so at apex; greatest width at base. Tegmen (
<figureCitation id="D6BE8BB21005C045FDC7FE74FD17699F" box="[576,642,476,500]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
F) 0.52× length of penis, with ring narrow, parameroid lobes developed with basal half more sclerotized area triangular; tegminal apodeme slightly stout, forming a Y-shape with basal piece, apex of apodeme rounded.
</paragraph>
<paragraph id="4E3A97371005C045FF4EFDEFFEC86ACC" blockId="9.[151,1437,151,1652]" pageId="9" pageNumber="170">
<emphasis id="7CF14B251005C045FF4EFDEFFD9F6A0B" bold="true" box="[201,522,583,608]" pageId="9" pageNumber="170">
Variation. Male
<typeStatus id="913E29951005C045FE0AFDE0FD906A0B" box="[397,517,584,608]" pageId="9" pageNumber="170" type="paratype">paratypes</typeStatus>
.
</emphasis>
Measurements (in mm): Standard length: 5.906.10; pronotal length: 1.651.72; pronotal width: 1.911.95; elytral length: 4.304.50; elytral width: 2.552.70; rostral length: 0.960.99; rostral width: 0.800.84.
</paragraph>
<paragraph id="4E3A97371005C045FF4EFD1CFA136C3C" blockId="9.[151,1437,151,1652]" pageId="9" pageNumber="170">
<emphasis id="7CF14B251005C045FF4EFD1CFE0E6AA7" bold="true" box="[201,411,692,716]" pageId="9" pageNumber="170">
Female
<typeStatus id="913E29951005C045FEAFFD1CFE036AA7" box="[296,406,692,716]" pageId="9" pageNumber="170" type="paratype">paratype</typeStatus>
.
</emphasis>
Measurements (in mm): Standard length: 4.906.50; pronotal length: 1.451.85; pronotal width: 1.601.98; elytral length: 3.454.60; elytral width: 2.352.80; rostral length: 0.961.02; rostral width: 0.720.82. Elytra slightly wider than male, pronotum less transverse than male, ventrite 5 (
<figureCitation id="D6BE8BB21005C045FB38FD54FA906B7F" box="[1215,1285,764,788]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
G) parabolic, similar in length to ventrite 2. Sternite VIII (
<figureCitation id="D6BE8BB21005C045FD18FCB7FD716B5C" box="[671,740,799,823]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
H) with spiculum ventrale thin, rod-like and almost straight, anteriorly slightly curved; lamina tightly folded, in lateral view triangular, ventral margin straight, strongly sclerotized. Ovipositor (
<figureCitation id="D6BE8BB21005C045FE2EFCCFFE7A6B14" box="[425,495,871,895]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
H,
<date id="3A3BB1F71005C045FD95FCCFFDBE6B14" box="[530,555,871,895]" pageId="9" pageNumber="170">4I</date>
) with coxites shorter than sternite VIII, folded along middle, lateral margins strongly sclerotized; styli relatively small, cylindrical, with 23 long setae inserted apically, strongly sclerotized in comparison to coxites. Bursal sclerites (
<figureCitation id="D6BE8BB21005C045FDECFC07FD266BAC" box="[619,691,943,967]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
J) near the point of union of spermathecal duct and of oviduct, parallel, each sclerite in lateral view chevron-shaped, with mesal area strongly sclerotized, dorsal surface of mesal area with many small spines. Spermatheca (
<figureCitation id="D6BE8BB21005C045FD19FC5FFD736C64" box="[670,742,1015,1039]" captionStart="FIGURE 4" captionStartId="8.[151,250,1902,1925]" captionTargetBox="[278,1321,222,1868]" captionTargetId="figure@8.[257,1335,193,1878]" captionTargetPageId="8" captionText="FIGURE 4. Geotragus brevidens sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216584/files/figure.png" pageId="9" pageNumber="170">Fig. 4</figureCitation>
K) with corpus elongate, slightly shorter than cornu; cornu strongly curved, V-shaped, apically abruptly narrowed, apex narrowly rounded; nodulus prominent, oblong and curved toward ramus at apex, subcontiguous with and angled more than 90° in relation to ramus; ramus reduced.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC1005C045FF4EFBCBFCC76E47" pageId="9" pageNumber="170" type="materials_examined">
<paragraph id="4E3A97371005C045FF4EFBCBFE666E63" blockId="9.[151,1437,151,1652]" pageId="9" pageNumber="170">
<emphasis id="7CF14B251005C045FF4EFBCBFDA86C17" bold="true" box="[201,573,1123,1148]" pageId="9" pageNumber="170">
Material examined.
<typeStatus id="913E29951005C045FE40FBCCFDAD6C17" box="[455,568,1124,1148]" pageId="9" pageNumber="170" type="holotype">Holotype</typeStatus>
:
</emphasis>
3: (white, printed):
<collectingCountry id="3692D7A71005C045FCC6FBCCFC086C17" box="[833,925,1124,1148]" name="China" pageId="9" pageNumber="170">CHINA</collectingCountry>
, Yunnan Prov. / Gongshan County, / Qiqi Reserve,
<quantity id="897D3AD21005C045FF78FB2FFEC56CF4" box="[255,336,1159,1183]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="170" unit="m" value="2000.0">2000m</quantity>
; Sino-America Exped. / 2000.7.12 Liang H. B. [7.12 was deleted by collector] /
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,
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/ 7.9 [handwritten by collector]; (red, printed):
<typeStatus id="913E29951005C045FD3CFB04FCC66CAF" box="[699,851,1196,1220]" pageId="9" pageNumber="170" type="holotype">HOLOTYPE</typeStatus>
; (white, printed): IOZ(E) 905049.
<typeStatus id="913E29951005C045FB61FB04FACB6CAF" box="[1254,1374,1196,1220]" pageId="9" pageNumber="170" type="paratype">
<emphasis id="7CF14B251005C045FB61FB04FACB6CAF" bold="true" box="[1254,1374,1196,1220]" pageId="9" pageNumber="170">Paratypes</emphasis>
</typeStatus>
(23, 2Ƥ): 13: (white, printed):
<collectingCountry id="3692D7A71005C045FE47FB67FD896C8C" box="[448,540,1231,1255]" name="China" pageId="9" pageNumber="170">CHINA</collectingCountry>
, Yunnan Prov. / Gongshan County, / No 12 Bridge-Yakou, /
<quantity id="897D3AD21005C045FB3AFB67FAC16C8C" box="[1213,1364,1231,1255]" metricMagnitude="3" metricUnit="m" metricValue="3.215" metricValueMax="3.6799999999999997" metricValueMin="2.75" pageId="9" pageNumber="170" unit="m" value="3215.0" valueMax="3680.0" valueMin="2750.0">27503680m</quantity>
; Sino-
<collectingCountry id="3692D7A71005C045FF10FB5CFF626D67" box="[151,247,1268,1292]" name="United States of America" pageId="9" pageNumber="170">America</collectingCountry>
Exped. / 2000.7.16 Liang H. B. /
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,
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; (yellow, printed):
<typeStatus id="913E29951005C045FBC2FB5CFB416D67" box="[1093,1236,1268,1292]" pageId="9" pageNumber="170" type="paratype">PARATYPE</typeStatus>
; (white, printed): IOZ(E) 905048. 2Ƥ: (white, printed):
<collectingCountry id="3692D7A71005C045FDC1FABFFD376D44" box="[582,674,1303,1327]" name="China" pageId="9" pageNumber="170">CHINA</collectingCountry>
, Yunnan Prov. / Fugong, Lishadi Town, / Shibali Yakou &amp;
<collectingCountry id="3692D7A71005C045FAC7FAB0FA196D44" box="[1344,1420,1304,1327]" name="Myanmar" pageId="9" pageNumber="170">Burma</collectingCountry>
/
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,
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;
<quantity id="897D3AD21005C045FE47FA94FD846D3F" box="[448,529,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="3.716" pageId="9" pageNumber="170" unit="m" value="3716.0">3716m</quantity>
, 2005.8.17 day / Liang H.B., Zhang J.F. / California Academy &amp; / IOZ., Chinese. Acad. Sci.; (yellow, printed):
<typeStatus id="913E29951005C045FE68FAC8FDEB6D13" box="[495,638,1376,1400]" pageId="9" pageNumber="170" type="paratype">PARATYPE</typeStatus>
; (white, printed): IOZ(E) 1505821 and IOZ(E) 1505822. 13: (white, printed): CASENT / 3005827;
<collectingCountry id="3692D7A71005C045FE73FA2CFDC56DF7" box="[500,592,1412,1436]" name="China" pageId="9" pageNumber="170">CHINA</collectingCountry>
, Yunnan Province, / Gaoligong Shan, Nujiang / Prefecture, Nujiang State / Nature Reserve, Dong Shao / Fang Area, 20 airkm W of / Gongshan,
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/;
<geoCoordinate id="2BB1F1F01005C045FBD3FA0FFB496DD4" box="[1108,1244,1447,1471]" direction="east" orientation="longitude" pageId="9" pageNumber="170" precision="1" value="98.48433">E98.48433°</geoCoordinate>
,
<quantity id="897D3AD21005C045FB6EFA0FFAAF6DD4" box="[1257,1338,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="170" unit="m" value="3300.0">3300m</quantity>
, /
<date id="3A3BB1F71005C045FAD1FA0FFE9F6D8F" pageId="9" pageNumber="170" value="2000-07-16" valueMax="2000-07-17" valueMin="2000-07-16">1617 July 2000</date>
, / Stop#0024, D.H. Kavanaugh, / C.E. Griswold, Liang H.-B., D. / Ubick, &amp; Dong D.-Z. collectors; (yellow, printed):
<typeStatus id="913E29951005C045FEE7FA58FE7A6E63" box="[352,495,1520,1544]" pageId="9" pageNumber="170" type="paratype">PARATYPE</typeStatus>
.
</paragraph>
<paragraph id="4E3A97371005C045FF4EF9BCFCC76E47" blockId="9.[151,1437,151,1652]" box="[201,850,1556,1581]" pageId="9" pageNumber="170">
<typeStatus id="913E29951005C045FF4EF9BCFEA46E47" box="[201,305,1556,1580]" pageId="9" pageNumber="170" type="holotype">Holotype</typeStatus>
and all
<typeStatus id="913E29951005C045FE0DF9BDFE606E46" box="[394,501,1557,1581]" pageId="9" pageNumber="170" type="paratype">paratypes</typeStatus>
are to be conserved in
<collectionCode id="28940FF21005C045FD7EF9BCFCDB6E47" box="[761,846,1556,1580]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="9" pageNumber="170">IZCAS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC1005C045FF4EF99FFAE56E24" box="[201,1392,1591,1616]" pageId="9" pageNumber="170" type="distribution">
<paragraph id="4E3A97371005C045FF4EF99FFAE56E24" blockId="9.[151,1437,151,1652]" box="[201,1392,1591,1616]" pageId="9" pageNumber="170">
<emphasis id="7CF14B251005C045FF4EF99FFEF76E3B" bold="true" box="[201,354,1591,1616]" pageId="9" pageNumber="170">Distribution.</emphasis>
<collectingCountry id="3692D7A71005C045FEEEF99FFE256E24" box="[361,432,1591,1615]" name="China" pageId="9" pageNumber="170">China</collectingCountry>
: Yunnan (Fugong, Gongshan). Altitudinal range of this species is from
<quantity id="897D3AD21005C045FB51F99FFAF96E24" box="[1238,1388,1591,1615]" metricMagnitude="3" metricUnit="m" metricValue="2.858" metricValueMax="3.716" metricValueMin="2.0" pageId="9" pageNumber="170" unit="m" value="2858.0" valueMax="3716.0" valueMin="2000.0">20003716m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC1005C045FF4EF9F4FB6C6E1F" box="[201,1273,1628,1652]" pageId="9" pageNumber="170" type="etymology">
<paragraph id="4E3A97371005C045FF4EF9F4FB6C6E1F" blockId="9.[151,1437,151,1652]" box="[201,1273,1628,1652]" pageId="9" pageNumber="170">
<emphasis id="7CF14B251005C045FF4EF9F4FEC56E1F" bold="true" box="[201,336,1628,1652]" pageId="9" pageNumber="170">Etymology.</emphasis>
The specific epithet is referring to the small size teeth on inner margin of protibiae.
</paragraph>
</subSubSection>
</treatment>
</document>