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<document id="745B32069D2CE1635763CA6768B5E0C6" ID-CLB-Dataset="22233" ID-DOI="10.11646/zootaxa.4777.1.1" ID-GBIF-Dataset="75de85af-39c5-41ca-9ccc-4ff7b743eefa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3821174" ID-ZooBank="BF4149EF-478F-4FE4-BE7B-B497272F99C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589267332095" checkinUser="plazi" docAuthor="Shen, Jia-Wei &amp; Leschen, Richard A. B." docDate="2020" docId="C0438794D872FF9EFF017412FE16FD89" docLanguage="en" docName="zootaxa.4777.1.1.pdf.imf" docOrigin="Zootaxa 4777 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eupines (Byraxis) mayae Shen &amp; Leschen 2020, sp. nov." docType="treatment" docUuid="4CF5D50D-65F4-477F-B2CE-E32144FC8D9F" docUuidSource="ZooBank" docVersion="6" lastPageNumber="50" masterDocId="3C7AFFECD842FFAFFF967608FFA4FE0A" masterDocTitle="Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)" masterLastPageNumber="84" masterPageNumber="1" pageNumber="49" updateTime="1698839235073" updateUser="ExternalLinkService">
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<mods:title id="8F51CB91D380EEADA5F0800C98AF766B">Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)</mods:title>
</mods:titleInfo>
<mods:name id="21E7D5FC5480841ECC1F568B7F09008C" type="personal">
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<mods:namePart id="AEAC5F7224FE7C0BCFAF6A58552D2BE7">Shen, Jia-Wei</mods:namePart>
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<mods:namePart id="9B57830BDF500EA84444F39D50D83F92">Leschen, Richard A. B.</mods:namePart>
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<mods:title id="88BCAD44D48020EF6D039732FB760771">Zootaxa</mods:title>
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<mods:date id="873D4BAE841685247DC0A5D595D2E085">2020</mods:date>
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<mods:number id="6E36F0DEDB99137532C262548E81253D">2020-05-12</mods:number>
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<mods:number id="B4EBEE8FF25E0FBC2045DDE054B6E47A">4777</mods:number>
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<treatment id="C0438794D872FF9EFF017412FE16FD89" ID-DOI="http://doi.org/10.5281/zenodo.3844500" ID-GBIF-Taxon="164123578" ID-Zenodo-Dep="3844500" LSID="urn:lsid:zoobank.org:act:4CF5D50D-65F4-477F-B2CE-E32144FC8D9F" httpUri="http://treatment.plazi.org/id/C0438794D872FF9EFF017412FE16FD89" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<subSubSection id="00F06509D872FF9FFF017412FD89FC3E" box="[151,557,538,564]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="48553682D872FF9FFF017412FD89FC3E" blockId="48.[151,557,538,600]" box="[151,557,538,564]" pageId="48" pageNumber="49">
<heading id="131D81EED872FF9FFF017412FD89FC3E" bold="true" box="[151,557,538,564]" fontSize="11" level="1" pageId="48" pageNumber="49" reason="1">
<emphasis id="7A9EEA90D872FF9FFF017412FD89FC3E" bold="true" box="[151,557,538,564]" pageId="48" pageNumber="49">
<taxonomicName id="8FEA4D01D872FF9FFF017412FE6CFC3E" ID-CoL="8XMD9" authority="Shen &amp; Leschen, 2020" authorityName="Shen &amp; Leschen" authorityYear="2020" box="[151,456,538,564]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="mayae" status="sp. nov." subGenus="Byraxis">
<emphasis id="7A9EEA90D872FF9FFF017412FF5EFC3E" bold="true" box="[151,250,538,564]" italics="true" pageId="48" pageNumber="49">Eupines</emphasis>
(
<emphasis id="7A9EEA90D872FF9FFE9A7412FECDFC3E" bold="true" box="[268,361,538,564]" italics="true" pageId="48" pageNumber="49">Byraxis</emphasis>
)
<emphasis id="7A9EEA90D872FF9FFEEC7412FE6CFC3E" bold="true" box="[378,456,538,564]" italics="true" pageId="48" pageNumber="49">mayae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61AD57EBD872FF9FFE597412FD89FC3E" box="[463,557,538,564]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="00F06509D872FF9FFF017436FED8FC52" box="[151,380,573,600]" pageId="48" pageNumber="49" type="description">
<paragraph id="48553682D872FF9FFF017436FED8FC52" blockId="48.[151,557,538,600]" box="[151,380,573,600]" pageId="48" pageNumber="49">
<figureCitation id="D0D12A07D872FF9FFF017436FF4EFC52" box="[151,234,574,600]" captionStart="FIGURE 31" captionStartId="48.[151,250,1901,1926]" captionTargetBox="[181,1406,880,1868]" captionTargetId="figure@48.[168,1420,868,1875]" captionTargetPageId="48" captionText="FIGURE 31. Diagnostic characters of E. (B.) mayae sp. n. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, BF = 0.2 mm, GJ = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821240" httpUri="https://zenodo.org/record/3821240/files/figure.png" pageId="48" pageNumber="49">Fig. 31</figureCitation>
; map
<figureCitation id="D0D12A07D872FF9FFEBB7436FED8FC52" box="[301,380,573,600]" captionStart="FIGURE 52" captionStartId="78.[151,250,1945,1970]" captionTargetBox="[202,1383,197,1920]" captionTargetId="figure@78.[194,1394,181,1920]" captionTargetPageId="78" captionText="FIGURE 52. Geographic distributions of E. (B.) impar (Sharp, 1874), E. (B.) impressifrons (Broun, 1880), E. (B.) insolita sp. n., E. (B.) lewisi Broun, 1910, E. (B.) longiceps Raffray, 1904, E. (B.) mayae sp. n., and E. (B.) micans (Sharp, 1874) in New Zealand." figureDoi="http://doi.org/10.5281/zenodo.3821282" httpUri="https://zenodo.org/record/3821282/files/figure.png" pageId="48" pageNumber="49">Fig. 52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="00F06509D872FF9FFF017488FC6BFD44" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="48553682D872FF9FFF017488FC6BFD44" blockId="48.[151,1436,640,846]" pageId="48" pageNumber="49">
<emphasis id="7A9EEA90D872FF9FFF017488FEB0FC91" bold="true" box="[151,276,640,667]" pageId="48" pageNumber="49">Diagnosis:</emphasis>
Head with vertexal foveae punctiform; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7, A8 and A10 unmodified, A9 asymmetrical and enlarged, slightly shorter than A10, with ventromedial excavation and minute protuberance at lateral side, A10 largest and conical. Abdominal ventrite 2 with pair of large discal protuberances close to each other, inserted with slightly curved medium-sized apical setae; ventrite 6 with pair of long and acute median struts, about one-third of ventral length. Protrochanters with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified.
</paragraph>
</subSubSection>
<caption id="1C95660AD872FF9FFF017165FC3BF9C7" ID-DOI="http://doi.org/10.5281/zenodo.3821240" ID-Zenodo-Dep="3821240" httpUri="https://zenodo.org/record/3821240/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1901,1926]" targetBox="[181,1406,880,1868]" targetPageId="48">
<paragraph id="48553682D872FF9FFF017165FC3BF9C7" blockId="48.[151,1436,1901,2034]" pageId="48" pageNumber="49">
<emphasis id="7A9EEA90D872FF9FFF017165FEBBF98C" bold="true" box="[151,287,1901,1926]" pageId="48" pageNumber="49">FIGURE 31.</emphasis>
Diagnostic characters of
<taxonomicName id="8FEA4D01D872FF9FFDB67166FD0EF98F" authority="Shen &amp; Leschen, 2020" authorityName="Shen &amp; Leschen" authorityYear="2020" box="[544,682,1901,1925]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="11" phylum="Arthropoda" rank="species" species="mayae" status="sp. nov." subGenus="Byraxis">
<emphasis id="7A9EEA90D872FF9FFDB67166FD8BF98F" box="[544,559,1902,1925]" italics="true" pageId="48" pageNumber="49">E</emphasis>
. (
<emphasis id="7A9EEA90D872FF9FFDD27166FDF7F98F" box="[580,595,1902,1925]" italics="true" pageId="48" pageNumber="49">B</emphasis>
.)
<emphasis id="7A9EEA90D872FF9FFDFE7166FD0EF98F" box="[616,682,1902,1925]" italics="true" pageId="48" pageNumber="49">mayae</emphasis>
</taxonomicName>
<emphasis id="7A9EEA90D872FF9FFD397165FCACF98C" bold="true" box="[687,776,1901,1926]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="61AD57EBD872FF9FFD397165FD4CF98F" box="[687,744,1901,1925]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
A)
</emphasis>
Habitus.
<emphasis id="7A9EEA90D872FF9FFCFF7165FC27F98C" bold="true" box="[873,899,1901,1926]" pageId="48" pageNumber="49">B)</emphasis>
Antenna, in dorsal view.
<emphasis id="7A9EEA90D872FF9FFB157165FB3AF98C" bold="true" box="[1155,1182,1901,1926]" pageId="48" pageNumber="49">C)</emphasis>
Same, in lateral view.
<emphasis id="7A9EEA90D872FF9FFA177165FA38F98C" bold="true" box="[1409,1436,1901,1926]" pageId="48" pageNumber="49">D)</emphasis>
Same, in ventral view.
<emphasis id="7A9EEA90D872FF9FFE177199FE3EF9A0" bold="true" box="[385,410,1937,1962]" pageId="48" pageNumber="49">E)</emphasis>
Protrochanter.
<emphasis id="7A9EEA90D872FF9FFDA17199FDF4F9A0" bold="true" box="[567,592,1937,1962]" pageId="48" pageNumber="49">F)</emphasis>
Ventrite 2.
<emphasis id="7A9EEA90D872FF9FFD517199FD47F9A0" bold="true" box="[711,739,1937,1962]" pageId="48" pageNumber="49">G)</emphasis>
Ventrite 6.
<emphasis id="7A9EEA90D872FF9FFCCD7199FCD2F9A0" bold="true" box="[859,886,1937,1962]" pageId="48" pageNumber="49">H)</emphasis>
Aedeagus, in dorsal view.
<emphasis id="7A9EEA90D872FF9FFB1D7199FB39F9A0" bold="true" box="[1163,1181,1937,1962]" pageId="48" pageNumber="49">I)</emphasis>
Same, in lateral view.
<emphasis id="7A9EEA90D872FF9FFA117199FA38F9A0" bold="true" box="[1415,1436,1937,1962]" pageId="48" pageNumber="49">J)</emphasis>
Same, in ventral view. Scale bars: A = 1 mm, BF = 0.2 mm, GJ = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="00F06509D872FF9EFF5171DFFAC9FC8D" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
<paragraph id="48553682D872FF9EFF5171DFFAC9FC8D" blockId="48.[151,1436,1901,2034]" lastBlockId="49.[151,1437,153,899]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="7A9EEA90D872FF9FFF5171DFFEFFF9F8" bold="true" box="[199,347,2007,2034]" pageId="48" pageNumber="49">Description:</emphasis>
Body length
<quantity id="8F129B67D872FF9FFE6871D0FD03F9F8" box="[510,679,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" metricValueMax="1.4" metricValueMin="1.34" pageId="48" pageNumber="49" unit="mm" value="1.37" valueMax="1.4" valueMin="1.34">1.341.40 mm</quantity>
; body colour reddish brown to dark brown with lighter coloured elytra and legs.
<emphasis id="7A9EEA90D873FF9EFEDB7691FE29FEB9" bold="true" box="[333,397,153,179]" pageId="49" pageNumber="50">Head</emphasis>
wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed and prominent in dorsal view, with about 34 facets; antennal tubercles moderately developed; postantennal notches broad and distinctive; vertexal foveae punctiform; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7, A8 and A10 unmodified, A9 asymmetrical and enlarged, slightly shorter than A10, with ventromedial excavation and minute protuberance at lateral side, A10 largest and conical.
<emphasis id="7A9EEA90D873FF9EFCDC7745FC67FF6D" bold="true" box="[842,963,333,359]" pageId="49" pageNumber="50">Pronotum</emphasis>
slightly wider than long, as wide as head at eye level; lacking lateral antebasal foveae.
<emphasis id="7A9EEA90D873FF9EFD097779FD4FFF81" bold="true" box="[671,747,369,395]" pageId="49" pageNumber="50">Elytra</emphasis>
slightly longer than wide, widest near middle; humeral calli slightly prominent in dorsal view; sutural stria weak, shallowly extending from base to apex.
<emphasis id="7A9EEA90D873FF9EFB34779CFAB3FFA5" bold="true" box="[1186,1303,404,431]" pageId="49" pageNumber="50">Abdomen</emphasis>
with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 with pair of large discal protuberances close to each other, inserted with slightly curved medium-sized apical setae; ventrite 6 with pair of long and acute median struts, about one-third of ventral length.
<emphasis id="7A9EEA90D873FF9EFEF57409FE3FFC11" bold="true" box="[355,411,513,539]" pageId="49" pageNumber="50">Legs</emphasis>
with protrochanters modified with short ventral spine, about two-fifths of protrochanteral width; protibiae and mesotibiae unmodified.
<emphasis id="7A9EEA90D873FF9EFD1A742CFD5BFC35" bold="true" box="[652,767,548,575]" pageId="49" pageNumber="50">Aedeagus</emphasis>
about
<quantity id="8F129B67D873FF9EFCDD742DFC0AFC35" box="[843,942,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="49" pageNumber="50" unit="mm" value="0.37">0.37 mm</quantity>
long, with long and stout parameres, curved towards each other at apex and bent downwards in lateral view, with three pairs of short subapical setae; median lobe with two slender sclerites, curved towards each other at apex, shorter and much narrower than parameres.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D873FF9EFF517498FBDCFCC5" pageId="49" pageNumber="50" type="discussion">
<paragraph id="48553682D873FF9EFF517498FBDCFCC5" blockId="49.[151,1437,153,899]" pageId="49" pageNumber="50">
<emphasis id="7A9EEA90D873FF9EFF517498FEF6FCA1" bold="true" box="[199,338,656,683]" pageId="49" pageNumber="50">Comments:</emphasis>
This species can be distinguished from all other
<taxonomicName id="8FEA4D01D873FF9EFCE0749AFC74FCA1" authorityName="King. A" authorityYear="1866" box="[886,976,658,683]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D873FF9EFCE0749AFC74FCA1" box="[886,976,658,683]" italics="true" pageId="49" pageNumber="50">Eupines</emphasis>
</taxonomicName>
by its simple antennal modifications and ventrite 2 with pair of large discal protuberances close to each other (
<figureCitation id="D0D12A07D873FF9EFC0874BDFC5CFCC5" box="[926,1016,693,719]" captionStart="FIGURE 31" captionStartId="48.[151,250,1901,1926]" captionTargetBox="[181,1406,880,1868]" captionTargetId="figure@48.[168,1420,868,1875]" captionTargetPageId="48" captionText="FIGURE 31. Diagnostic characters of E. (B.) mayae sp. n. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, BF = 0.2 mm, GJ = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821240" httpUri="https://zenodo.org/record/3821240/files/figure.png" pageId="49" pageNumber="50">Figs. 31</figureCitation>
BD, 31F).
</paragraph>
</subSubSection>
<subSubSection id="00F06509D873FF9EFF5174D0FD90FCF9" box="[199,564,728,755]" pageId="49" pageNumber="50" type="distribution">
<paragraph id="48553682D873FF9EFF5174D0FD90FCF9" blockId="49.[151,1437,153,899]" box="[199,564,728,755]" pageId="49" pageNumber="50">
<emphasis id="7A9EEA90D873FF9EFF5174D0FEC7FCF9" bold="true" box="[199,355,728,755]" pageId="49" pageNumber="50">Distribution:</emphasis>
North Island. ND.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D873FF9EFF5174F5FBEEFD1D" box="[199,1098,765,791]" pageId="49" pageNumber="50" type="etymology">
<paragraph id="48553682D873FF9EFF5174F5FBEEFD1D" blockId="49.[151,1437,153,899]" box="[199,1098,765,791]" pageId="49" pageNumber="50">
<emphasis id="7A9EEA90D873FF9EFF5174F5FEF6FD1D" bold="true" box="[199,338,765,791]" pageId="49" pageNumber="50">Etymology:</emphasis>
This species is named after the collector of the
<typeStatus id="97518820D873FF9EFCFF74F5FC6AFD1D" box="[873,974,765,791]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
, V. A. May
</paragraph>
</subSubSection>
<subSubSection id="00F06509D873FF9EFF517529FE16FD89" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="48553682D873FF9EFF517529FE16FD89" blockId="49.[151,1437,153,899]" pageId="49" pageNumber="50">
<emphasis id="7A9EEA90D873FF9EFF517529FDF9FD31" bold="true" box="[199,605,800,827]" pageId="49" pageNumber="50">
Type material (
<collectionCode id="2EFBAE47D873FF9EFEE87528FE70FD31" box="[382,468,800,827]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="49" pageNumber="50" type="Other institutional type">NZAC</collectionCode>
):
<typeStatus id="97518820D873FF9EFE7F7529FDFCFD31" box="[489,600,801,827]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
.
</emphasis>
<materialsCitation id="F8823CDFD873FF9EFDF37529FCCEFD55" ID-GBIF-Occurrence="2612164359" collectingDate="1976-12-13" collectorName="V. A. May Litter" country="New Zealand" elevation="386" location="NEW ZEALAND" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="5EECFD0BD873FF9EFDF37529FDD8FD31" box="[613,636,801,827]" pageId="49" pageNumber="50" type="male"></specimenCount>
, specimen entire, card-mounted: “
<collectingCountry id="30FD7612D873FF9EFC6A7529FB69FD31" box="[1020,1229,801,827]" name="New Zealand" pageId="49" pageNumber="50">NEW ZEALAND</collectingCountry>
, ND, Mangamuka Smt.,
<quantity id="8F129B67D873FF9EFF4C754DFEB9FD55" box="[218,285,837,863]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="49" pageNumber="50" unit="m" value="386.0">
<elevation id="C3C7D1B1D873FF9EFF4C754DFEB9FD55" box="[218,285,837,863]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="49" pageNumber="50" unit="m" value="386.0">386m</elevation>
</quantity>
,
<date id="3C541042D873FF9EFEBF754DFE1AFD55" box="[297,446,837,863]" pageId="49" pageNumber="50" value="1976-12-13">
<collectingDate id="2C10E9AAD873FF9EFEBF754DFE1AFD55" box="[297,446,837,863]" pageId="49" pageNumber="50" value="1976-12-13">13 Dec 1976</collectingDate>
</date>
,
<collectorName id="E51F5354D873FF9EFE5F754DFDDAFD55" box="[457,638,837,863]" pageId="49" pageNumber="50">V.A. May Litter</collectorName>
76/109”.
<typeStatus id="97518820D873FF9EFD66754DFCCEFD55" box="[752,874,837,863]" pageId="49" pageNumber="50" type="paratype">
<emphasis id="7A9EEA90D873FF9EFD66754DFCCEFD55" bold="true" box="[752,874,837,863]" pageId="49" pageNumber="50">Paratypes</emphasis>
</typeStatus>
</materialsCitation>
<emphasis id="7A9EEA90D873FF9EFCFC754DFCCBFD55" bold="true" box="[874,879,837,863]" pageId="49" pageNumber="50">.</emphasis>
<specimenCount id="5EECFD0BD873FF9EFCE1754DFC16FD55" box="[887,946,837,863]" pageId="49" pageNumber="50" type="generic">1 ex.</specimenCount>
:
<emphasis id="7A9EEA90D873FF9EFC28754DFB3FFD55" bold="true" box="[958,1179,836,863]" pageId="49" pageNumber="50">Unknown locality:</emphasis>
T. Broun Collection // A.E. Brookes Collection.
</paragraph>
</subSubSection>
</treatment>
</document>