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<document id="8F86F128647F8A65B4004AE6E8520399" ID-DOI="10.59893/abud.24(2).018" ID-ISSN="1407-8953" ID-Zenodo-Dep="14659830" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736974627243" checkinUser="felipe" docAuthor="Rukmane-Bārbale, Anita" docDate="2024" docId="C004D307FFCCFFFCFD6FCC6BFBE6FF3C" docLanguage="en" docName="ActaBiolUnivDaugavp.24.2.349-358.pdf" docOrigin="Acta Biologica Universitatis Daugavpiliensis 24 (2)" docSource="https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/four-new-species-of-pachyrhynchini-from-southern-luzon-philippines-coleoptera-curculionidae-entiminae/" docStyle="DocumentStyle:DD7D35B4ACFEF3C20AEB77B8656EEADF.5:ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleId="DD7D35B4ACFEF3C20AEB77B8656EEADF" docStyleName="ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleVersion="5" docTitle="Pachyrhynchus masatoshiensis Rukmane-Bārbale, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="353" masterDocId="3C3DAB7FFFCDFFF8FFEFCF06FF97FFAD" masterDocTitle="Four New Species Of Pachyrhynchini From Southern Luzon, Philippines (Coleoptera, Curculionidae, Entiminae)" masterLastPageNumber="358" masterPageNumber="349" pageNumber="350" updateTime="1736975198544" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="AD5E72D472BBA8FABD24B7C5F388D3E5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4A3534CFBADCDEA3495297E787F7CA40">
<mods:title id="5D2070C47050B233448452AAB6D4A08F">Four New Species Of Pachyrhynchini From Southern Luzon, Philippines (Coleoptera, Curculionidae, Entiminae)</mods:title>
</mods:titleInfo>
<mods:name id="0EAFE332F1A663079F0CDA03040AD5DE" type="personal">
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</mods:role>
<mods:namePart id="BC0FBCA259022E62C0F42085CEFF0CE9">Rukmane-Bārbale, Anita</mods:namePart>
<mods:affiliation id="016D7C5890B71C12CECC711EBC834C5D">Corresponding author: Anita Rukmane-Bārbale. Daugavpils University, Institute of Life Sciences and Technology, Coleopterological Research Center, Vienības Str. 13, LV- 5401, Daugavpils, Latvia. E-mail: anita. rukmane @ du. lv</mods:affiliation>
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<mods:typeOfResource id="44AAC22E329094C964322F24F83C8C64">text</mods:typeOfResource>
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<mods:title id="E25A3B7FE5ED62E641EF4B4E6B984CDD">Acta Biologica Universitatis Daugavpiliensis</mods:title>
</mods:titleInfo>
<mods:part id="1167BD722B055B060D5B1F7BE891F7A4">
<mods:date id="A72A842D8FC2893F1EC221A329F72403">2024</mods:date>
<mods:detail id="99ACCDA4ADF41557F0A852F77E4791C7" type="pubDate">
<mods:number id="BEF96FF473D803E1F8DEBC42DF66265D">2024-12-30</mods:number>
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<mods:detail id="75B088D5BE5AB622F6448CFAD80D7702" type="volume">
<mods:number id="0D9D552EDDABE92440B09750639622B0">24</mods:number>
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<mods:url id="BF5AF3132C89124A8B9F554187C6FB6D">https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/four-new-species-of-pachyrhynchini-from-southern-luzon-philippines-coleoptera-curculionidae-entiminae/</mods:url>
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<mods:classification id="501A12290F0CC975626B798FB05348D2">journal article</mods:classification>
<mods:identifier id="BAEF848EC0722890C3AFD91296F841AF" type="DOI">10.59893/abud.24(2).018</mods:identifier>
<mods:identifier id="4DA5183276DE4D5CA5E5F43534E9F68F" type="ISSN">1407-8953</mods:identifier>
<mods:identifier id="38E5923F5AC20AB81161EB43E0B41E3C" type="Zenodo-Dep">14659830</mods:identifier>
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<treatment id="C004D307FFCCFFFCFD6FCC6BFBE6FF3C" LSID="urn:lsid:plazi:treatment:C004D307FFCCFFFCFD6FCC6BFBE6FF3C" httpUri="http://treatment.plazi.org/id/C004D307FFCCFFFCFD6FCC6BFBE6FF3C" lastPageId="4" lastPageNumber="353" pageId="1" pageNumber="350">
<subSubSection id="00B7319AFFCCFFF9FD6FCC6BFBA3FC2B" box="[640,1076,877,902]" pageId="1" pageNumber="350" type="nomenclature">
<paragraph id="48126211FFCCFFF9FD6FCC6BFBA3FC2B" blockId="1.[640,1076,877,934]" box="[640,1076,877,902]" pageId="1" pageNumber="350">
<heading id="135AD57DFFCCFFF9FD6FCC6BFBA3FC2B" bold="true" box="[640,1076,877,902]" fontSize="10" level="2" pageId="1" pageNumber="350" reason="5">
<emphasis id="7AD9BE03FFCCFFF9FD6FCC6BFBA3FC2B" bold="true" box="[640,1076,877,902]" pageId="1" pageNumber="350">
<taxonomicName id="8FAD1992FFCCFFF9FD6FCC6BFC4FFC28" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[640,984,877,901]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="masatoshiensis" status="sp. nov.">
<emphasis id="7AD9BE03FFCCFFF9FD6FCC6BFC4FFC28" bold="true" box="[640,984,877,901]" italics="true" pageId="1" pageNumber="350">Pachyrhynchus masatoshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCCFFF9FC30CC68FBA3FC2B" box="[991,1076,878,902]" pageId="1" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="00B7319AFFCCFFF9FD6FCC88FC1FFC0B" box="[640,904,910,934]" pageId="1" pageNumber="350" type="description">
<paragraph id="48126211FFCCFFF9FD6FCC88FC1FFC0B" blockId="1.[640,1076,877,934]" box="[640,904,910,934]" pageId="1" pageNumber="350">
(
<figureCitation id="D0967E94FFCCFFF9FD67CC88FD55FC0B" box="[648,706,910,934]" captionStart="Figure 1" captionStartId="2.[677,753,1550,1574]" captionTargetBox="[728,1106,750,1529]" captionTargetId="figure-465@2.[727,1123,746,1535]" captionTargetPageId="2" captionText="Figure 1. Pachyrhynchus masatoshiensis sp. nov. habitus of holotype, male (A), female (B); P. chrysocyaneus sp. nov. habitus of holotype, male (C), female (D). Image cour- tesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659832" httpUri="https://zenodo.org/record/14659832/files/figure.png" pageId="1" pageNumber="350">Fig. 1</figureCitation>
A-B; 3F, L; 4D, I)
</paragraph>
</subSubSection>
<subSubSection id="00B7319AFFCCFFF9FD6FCCC8FD74FACB" pageId="1" pageNumber="350" type="materials_examined">
<paragraph id="48126211FFCCFFF9FD6FCCC8FC78FB2B" blockId="1.[640,1136,974,1382]" pageId="1" pageNumber="350">
<emphasis id="7AD9BE03FFCCFFF9FD6FCCC8FCA5FC4B" bold="true" box="[640,818,974,998]" pageId="1" pageNumber="350">Type material.</emphasis>
<materialsCitation id="F8C5684CFFCCFFF9FCD1CCC8FC4BFBEB" collectingDate="2022-09" country="Philippines" county="Luzon" location="Tabaco" municipality="Bicol" pageId="1" pageNumber="350" specimenCount="1" stateProvince="Albay" typeStatus="holotype">
<typeStatus id="9716DCB3FFCCFFF9FCD1CCC8FC3EFC4B" box="[830,937,974,998]" pageId="1" pageNumber="350" type="holotype">Holotype</typeStatus>
, male (
<figureCitation id="D0967E94FFCCFFF9FBECCCC8FBF7FC4B" box="[1027,1120,974,998]" captionStart="Figure 1" captionStartId="2.[677,753,1550,1574]" captionTargetBox="[728,1106,750,1529]" captionTargetId="figure-465@2.[727,1123,746,1535]" captionTargetPageId="2" captionText="Figure 1. Pachyrhynchus masatoshiensis sp. nov. habitus of holotype, male (A), female (B); P. chrysocyaneus sp. nov. habitus of holotype, male (C), female (D). Image cour- tesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659832" httpUri="https://zenodo.org/record/14659832/files/figure.png" pageId="1" pageNumber="350">Fig. 1A</figureCitation>
):
<collectingCountry id="30BA2281FFCCFFF9FD6FCCE8FC89FBAB" box="[640,798,1006,1030]" name="Philippines" pageId="1" pageNumber="350">PHILIPPINES</collectingCountry>
/
<collectingCounty id="A1731A9DFFCCFFF9FCDBCCE8FCECFBA8" box="[820,891,1006,1029]" pageId="1" pageNumber="350">Luzon</collectingCounty>
,
<collectingMunicipality id="A876F86BFFCCFFF9FC68CCE8FC53FBAB" box="[903,964,1006,1030]" pageId="1" pageNumber="350">Bicol</collectingMunicipality>
,
<collectingRegion id="8A69ACF3FFCCFFF9FC22CCE8FB98FBAB" box="[973,1039,1006,1030]" country="Philippines" name="Albay" pageId="1" pageNumber="350">Albay</collectingRegion>
,
<location id="4D7234CAFFCCFFF9FBF4CCE8FBFCFBAB" LSID="urn:lsid:plazi:treatment:C004D307FFCCFFFCFD6FCC6BFBE6FF3C:4D7234CAFFCCFFF9FBF4CCE8FBFCFBAB" box="[1051,1131,1006,1030]" country="Philippines" county="Luzon" municipality="Bicol" name="Tabaco" pageId="1" pageNumber="350" stateProvince="Albay">Tabaco</location>
, Mt. /
<location id="4D7234CAFFCCFFF9FD2FCB08FCF9FB8B" LSID="urn:lsid:plazi:treatment:C004D307FFCCFFFCFD6FCC6BFBE6FF3C:4D7234CAFFCCFFF9FD2FCB08FCF9FB8B" box="[704,878,1038,1062]" country="Philippines" county="Luzon" municipality="Bicol" name="Mayon Volcano" pageId="1" pageNumber="350" stateProvince="Albay">Mayon Volcano</location>
/
<date id="3C1344D1FFCCFFF9FC67CB08FC7CFB8B" box="[904,1003,1038,1062]" pageId="1" pageNumber="350" value="2022-09">
<collectingDate id="2C57BD39FFCCFFF9FC67CB08FC7CFB8B" box="[904,1003,1038,1062]" pageId="1" pageNumber="350" value="2022-09">IX. 2022</collectingDate>
</date>
/ local collector leg. (white printed label) //
</materialsCitation>
<typeStatus id="9716DCB3FFCCFFF9FC0DCB28FBF9FBEB" box="[994,1134,1070,1094]" pageId="1" pageNumber="350" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="8FAD1992FFCCFFF9FD7FCB48FC72FBCB" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[656,997,1102,1126]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="masatoshiensis">
<emphasis id="7AD9BE03FFCCFFF9FD7FCB48FC72FBCB" box="[656,997,1102,1126]" italics="true" pageId="1" pageNumber="350">Pachyrhynchus masatoshiensis</emphasis>
</taxonomicName>
/ Rukmane-Barbale, 2024 (typed on red card).
</paragraph>
<paragraph id="48126211FFCCFFF9FD6FCB88FD74FACB" blockId="1.[640,1136,974,1382]" pageId="1" pageNumber="350">
<typeStatus id="9716DCB3FFCCFFF9FD6FCB88FD7BFB0B" box="[640,748,1166,1190]" pageId="1" pageNumber="350" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="5EABA998FFCCFFF9FCEFCB88FCB4FB0B" box="[768,803,1166,1190]" count="4" pageId="1" pageNumber="350" type="female">4♀</specimenCount>
):
<materialsCitation id="F8C5684CFFCCFFF9FCD2CB88FBFEFB0B" box="[829,1129,1166,1190]" collectingDate="2022-09" country="Philippines" county="Luzon" location="Tabaco" municipality="Bicol" pageId="1" pageNumber="350" specimenCount="3" specimenCount-female="3" stateProvince="Albay" typeStatus="paratype">
<specimenCount id="5EABA998FFCCFFF9FCD2CB88FCC8FB0B" box="[829,863,1166,1190]" count="3" pageId="1" pageNumber="350" type="female">3♀</specimenCount>
same data as holotype
</materialsCitation>
;
<materialsCitation id="F8C5684CFFCCFFF9FD6FCBA8FD48FACB" collectingDate="2024-03" collectionCode="DUBC" collectorName="The" country="Philippines" location="Gen Nakar" municipality="Luzon" pageId="1" pageNumber="350" specimenCount="1" specimenCount-female="1" stateProvince="Quezon" typeStatus="paratype">
<specimenCount id="5EABA998FFCCFFF9FD6FCBA8FD34FB6B" box="[640,675,1198,1222]" count="1" pageId="1" pageNumber="350" type="female">1♀</specimenCount>
<collectingCountry id="30BA2281FFCCFFF9FD5FCBA8FCC1FB6B" box="[688,854,1198,1222]" name="Philippines" pageId="1" pageNumber="350">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="A876F86BFFCCFFF9FC98CBA8FC55FB68" box="[887,962,1198,1221]" pageId="1" pageNumber="350">Luzon</collectingMunicipality>
,
<collectingRegion id="8A69ACF3FFCCFFF9FC3BCBA8FBB8FB6B" box="[980,1071,1198,1222]" country="Philippines" name="Quezon" pageId="1" pageNumber="350">Quezon</collectingRegion>
,
<location id="4D7234CAFFCCFFF9FBAFCBA8FD55FB4B" LSID="urn:lsid:plazi:treatment:C004D307FFCCFFFCFD6FCC6BFBE6FF3C:4D7234CAFFCCFFF9FBAFCBA8FD55FB4B" country="Philippines" municipality="Luzon" name="Gen Nakar" pageId="1" pageNumber="350" stateProvince="Quezon">Gen Nakar</location>
/
<date id="3C1344D1FFCCFFF9FD37CBC8FCA1FB4B" box="[728,822,1230,1254]" pageId="1" pageNumber="350" value="2024-03">
<collectingDate id="2C57BD39FFCCFFF9FD37CBC8FCA1FB4B" box="[728,822,1230,1254]" pageId="1" pageNumber="350" value="2024-03">III. 2024</collectingDate>
</date>
/ local collector leg.
<collectorName id="E55807C7FFCCFFF9FBF3CBC8FBD1FB4B" box="[1052,1094,1230,1254]" pageId="1" pageNumber="350">The</collectorName>
additional label: “
<typeStatus id="9716DCB3FFCCFFF9FCC8CBE8FC27FAAB" box="[807,944,1262,1286]" pageId="1" pageNumber="350" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="8FAD1992FFCCFFF9FC27CBE8FCBCFA8B" authorityName="Rukmane-Bārbale" authorityYear="2024" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="masatoshiensis">
<emphasis id="7AD9BE03FFCCFFF9FC27CBE8FCBCFA8B" italics="true" pageId="1" pageNumber="350">Pachyrhynchus masatoshiensis</emphasis>
</taxonomicName>
/ Rukmane-Barbale, 2024” (typed on red card) was added to all
<typeStatus id="9716DCB3FFCCFFF9FBE7CA28FBF9FAEB" box="[1032,1134,1326,1350]" pageId="1" pageNumber="350" type="paratype">paratypes</typeStatus>
(
<collectionCode id="2EBCFAD4FFCCFFF9FD68CA48FD40FACB" box="[647,727,1358,1382]" pageId="1" pageNumber="350">DUBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="00B7319AFFCCFFFAFD6FCA88FE54FD8B" lastPageId="2" lastPageNumber="351" pageId="1" pageNumber="350" type="diagnosis">
<paragraph id="48126211FFCCFFF9FD6FCA88FBF9F92B" blockId="1.[640,1135,1422,1670]" pageId="1" pageNumber="350">
<emphasis id="7AD9BE03FFCCFFF9FD6FCA88FD6BFA0B" bold="true" box="[640,764,1422,1446]" pageId="1" pageNumber="350">Diagnosis.</emphasis>
<taxonomicName id="8FAD1992FFCCFFF9FCE6CA88FBF9FA0B" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[777,1134,1422,1446]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="masatoshiensis" status="sp. nov.">
<emphasis id="7AD9BE03FFCCFFF9FCE6CA88FBF9FA0B" box="[777,1134,1422,1446]" italics="true" pageId="1" pageNumber="350">Pachyrhynchus masatoshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCCFFF9FD6FCAA8FD42FA68" box="[640,725,1454,1477]" pageId="1" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="8FAD1992FFCCFFF9FC86CAA9FC86FA4B" authority="Yoshitake &amp; Yap, 2017" authorityName="Yoshitake &amp; Yap" authorityYear="2017" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCCFFF9FC86CAA9FC6CFA6B" box="[873,1019,1454,1478]" italics="true" pageId="1" pageNumber="350">P. masatoshii</emphasis>
<bibRefCitation id="2C3C1FE0FFCCFFF9FBEBCAA8FC86FA4B" author="Yoshitake H. &amp; Yap S. A." pageId="1" pageNumber="350" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake &amp; Yap, 2017</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8FAD1992FFCCFFF9FCA2CAC9FD2DF9AB" authority="Barsevskis, 2016" authorityName="Barsevskis" authorityYear="2016" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="rukmaneae">
<emphasis id="7AD9BE03FFCCFFF9FCA2CAC9FC72FA4B" box="[845,997,1486,1510]" italics="true" pageId="1" pageNumber="350">P. rukmaneae</emphasis>
<bibRefCitation id="2C3C1FE0FFCCFFF9FC00CAC8FD2DF9AB" author="Barsevskis A." pageId="1" pageNumber="350" pagination="1 - 6" refId="ref5388" refString="Barsevskis A. 2016. New species of Pachyrhynchus Germar, 1824 and Macrocyrtus Heller, 1912 (Coleoptera: Curculionidae) from the Marinduque Island (Philippines) as a new example of mimetic species pair. Acta Biologica Universitatis Daugavpiliensis 16 (1): 1 - 6." type="journal article" year="2016">Barševskis, 2016</bibRefCitation>
</taxonomicName>
. The new species can be distinguished by 1) thick band markings on prothorax, that lacks in both similar species: prothorax of
<taxonomicName id="8FAD1992FFCCFFF9FD6FC949FB82F9CB" authority="Yoshitake &amp; Yap, 2017" authorityName="Yoshitake &amp; Yap" authorityYear="2017" box="[640,1045,1614,1638]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCCFFF9FD6FC949FC87F9CB" box="[640,784,1614,1638]" italics="true" pageId="1" pageNumber="350">P. masatoshii</emphasis>
<bibRefCitation id="2C3C1FE0FFCCFFF9FCF7C948FB82F9CB" author="Yoshitake H. &amp; Yap S. A." box="[792,1045,1614,1638]" pageId="1" pageNumber="350" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake &amp; Yap, 2017</bibRefCitation>
</taxonomicName>
is without markings (
<figureCitation id="D0967E94FFCCFFF9FCCFC968FCF6F92B" box="[800,865,1646,1670]" captionStart="Figure 2" captionStartId="3.[640,719,526,550]" captionTargetBox="[642,1121,205,495]" captionTargetId="figure-134@3.[640,1131,201,515]" captionTargetPageId="3" captionText="Figure 2. Dorsal habitus of selected spe- cies: Pachyrhynchus caeruleus Yoshitake, 2019, male (A), female (B); P. masatoshii Yoshitake &amp; Yap, 2017, male (C), female (D); P.yuukaeYoshitake, 2019, male (E), female (F); P. rukmaneae Barševskis, 2016, male (G), female (H), P. rukmaneae ssp. pauscignitus Yoshitake, 2017, male. Images courtesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659834" httpUri="https://zenodo.org/record/14659834/files/figure.png" pageId="1" pageNumber="350">Fig. 2</figureCitation>
C-D) while prothorax of
</paragraph>
<paragraph id="48126211FFCFFFFAFF55CF79FBE6FF3C" blockId="2.[186,1137,126,145]" box="[186,1137,126,145]" pageId="2" pageNumber="351">
<emphasis id="7AD9BE03FFCFFFFAFF55CF79FBE6FF3C" box="[186,1137,126,145]" italics="true" pageId="2" pageNumber="351">
Four new species of
<taxonomicName id="8FAD1992FFCFFFFAFE84CF78FD89FF3C" authority="from" authorityName="FROM" box="[363,542,126,145]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini from</taxonomicName>
Southern Luzon,
<collectingCountry id="30BA2281FFCFFFFAFD5ACF78FC81FF3C" box="[693,790,126,145]" name="Philippines" pageId="2" pageNumber="351">Philippines</collectingCountry>
(
<taxonomicName id="8FAD1992FFCFFFFAFCCDCF78FC12FF3C" authorityName="Linnaeus" authorityYear="1758" box="[802,901,126,145]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="2" pageNumber="351" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="8FAD1992FFCFFFFAFC62CF78FB9DFF3C" box="[909,1034,126,145]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
,
<taxonomicName id="8FAD1992FFCFFFFAFBFDCF78FBFCFF3C" box="[1042,1131,126,145]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="subFamily" subFamily="Entiminae">Entiminae</taxonomicName>
)
</emphasis>
</paragraph>
<paragraph id="48126211FFCFFFFAFF78CFC9FE54FD8B" blockId="2.[151,649,206,550]" pageId="2" pageNumber="351">
<taxonomicName id="8FAD1992FFCFFFFAFF78CFC9FE73FF4B" authority="Barsevskis, 2016" authorityName="Barsevskis" authorityYear="2016" box="[151,484,206,230]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="rukmaneae">
<emphasis id="7AD9BE03FFCFFFFAFF78CFC9FEB0FF4B" box="[151,295,206,230]" italics="true" pageId="2" pageNumber="351">P. rukmaneae</emphasis>
<bibRefCitation id="2C3C1FE0FFCFFFFAFEC0CFC8FE73FF4B" author="Barsevskis A." box="[303,484,206,230]" pageId="2" pageNumber="351" pagination="1 - 6" refId="ref5388" refString="Barsevskis A. 2016. New species of Pachyrhynchus Germar, 1824 and Macrocyrtus Heller, 1912 (Coleoptera: Curculionidae) from the Marinduque Island (Philippines) as a new example of mimetic species pair. Acta Biologica Universitatis Daugavpiliensis 16 (1): 1 - 6." type="journal article" year="2016">Barševskis, 2016</bibRefCitation>
</taxonomicName>
is with narrow transverse band of scales along anterior margin and triangular shape band along posterior margin medially (
<figureCitation id="D0967E94FFCFFFFAFEB9CE28FE74FEEB" box="[342,483,302,326]" captionStart="Figure 2" captionStartId="3.[640,719,526,550]" captionTargetBox="[642,1121,205,495]" captionTargetId="figure-134@3.[640,1131,201,515]" captionTargetPageId="3" captionText="Figure 2. Dorsal habitus of selected spe- cies: Pachyrhynchus caeruleus Yoshitake, 2019, male (A), female (B); P. masatoshii Yoshitake &amp; Yap, 2017, male (C), female (D); P.yuukaeYoshitake, 2019, male (E), female (F); P. rukmaneae Barševskis, 2016, male (G), female (H), P. rukmaneae ssp. pauscignitus Yoshitake, 2017, male. Images courtesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659834" httpUri="https://zenodo.org/record/14659834/files/figure.png" pageId="2" pageNumber="351">Fig. 2G, H, I</figureCitation>
); 2) rostrum of
<taxonomicName id="8FAD1992FFCFFFFAFF78CE49FEC5FECB" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[151,338,334,358]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="masatoshiensis" status="sp. nov.">
<emphasis id="7AD9BE03FFCFFFFAFF78CE49FEC5FECB" box="[151,338,334,358]" italics="true" pageId="2" pageNumber="351">P. masatoshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCFFFFAFEB5CE48FE3BFEC8" box="[346,428,334,357]" pageId="2" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
longer, with moderate bulge at apical half while without bulge in
<taxonomicName id="8FAD1992FFCFFFFAFD9DCE69FE51FE0B" authority="Barsevskis, 2016" authorityName="Barsevskis" authorityYear="2016" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="rukmaneae">
<emphasis id="7AD9BE03FFCFFFFAFD9DCE69FE9BFE0B" italics="true" pageId="2" pageNumber="351">P. rukmaneae</emphasis>
<bibRefCitation id="2C3C1FE0FFCFFFFAFEFDCE88FE51FE0B" author="Barsevskis A." box="[274,454,398,422]" pageId="2" pageNumber="351" pagination="1 - 6" refId="ref5388" refString="Barsevskis A. 2016. New species of Pachyrhynchus Germar, 1824 and Macrocyrtus Heller, 1912 (Coleoptera: Curculionidae) from the Marinduque Island (Philippines) as a new example of mimetic species pair. Acta Biologica Universitatis Daugavpiliensis 16 (1): 1 - 6." type="journal article" year="2016">Barševskis, 2016</bibRefCitation>
</taxonomicName>
and smaller bulge in
<taxonomicName id="8FAD1992FFCFFFFAFF5BCEA9FDDCFE6B" authority="Yoshitake &amp; Yap, 2017" authorityName="Yoshitake &amp; Yap" authorityYear="2017" box="[180,587,430,454]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCFFFFAFF5BCEA9FED2FE6B" box="[180,325,430,454]" italics="true" pageId="2" pageNumber="351">P. masatoshii</emphasis>
<bibRefCitation id="2C3C1FE0FFCFFFFAFEA1CEA8FDDCFE6B" author="Yoshitake H. &amp; Yap S. A." box="[334,587,430,454]" pageId="2" pageNumber="351" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake &amp; Yap, 2017</bibRefCitation>
</taxonomicName>
(
<figureCitation id="D0967E94FFCFFFFAFDB4CEA8FF66FE4B" captionStart="Figure 3" captionStartId="3.[640,715,1422,1446]" captionTargetBox="[641,1114,858,1388]" captionTargetId="figure-189@3.[640,1131,832,1408]" captionTargetPageId="3" captionText="Figure 3. Head and rostrum in lateral and dor- sal view. Pachyrhynchus caeruleusYoshitake, 2019 (A, G); P. masatoshii Yoshitake &amp; Yap. 2017 (B, H); P. rukmaneae Barševskis, 2016 (C, I); P. yuukae Yoshitake, 2019 (D, J); P. chrysocyaneus sp. nov. (E, K); P. pseudorukmaneaesp. nov. (F, L). Images courtesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659838" httpUri="https://zenodo.org/record/14659838/files/figure.png" pageId="2" pageNumber="351">Fig. 3B, C, F</figureCitation>
); 3) eyes ovate in
<taxonomicName id="8FAD1992FFCFFFFAFE44CEC9FDF5FE4B" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[427,610,462,486]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="masatoshiensis" status="sp. nov.">
<emphasis id="7AD9BE03FFCFFFFAFE44CEC9FDF5FE4B" box="[427,610,462,486]" italics="true" pageId="2" pageNumber="351">P. masatoshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCFFFFAFD88CEC8FF52FDA8" pageId="2" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
while globular in
<taxonomicName id="8FAD1992FFCFFFFAFE67CEE9FE89FD8B" authority="Yoshitake &amp; Yap, 2017" authorityName="Yoshitake &amp; Yap" authorityYear="2017" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCFFFFAFE67CEE9FD81FDAB" box="[392,534,494,518]" italics="true" pageId="2" pageNumber="351">P. masatoshii</emphasis>
<bibRefCitation id="2C3C1FE0FFCFFFFAFDF3CEE8FE89FD8B" author="Yoshitake H. &amp; Yap S. A." pageId="2" pageNumber="351" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake &amp; Yap, 2017</bibRefCitation>
</taxonomicName>
(
<figureCitation id="D0967E94FFCFFFFAFEC2CD08FE20FD8B" box="[301,439,526,550]" captionStart="Figure 3" captionStartId="3.[640,715,1422,1446]" captionTargetBox="[641,1114,858,1388]" captionTargetId="figure-189@3.[640,1131,832,1408]" captionTargetPageId="3" captionText="Figure 3. Head and rostrum in lateral and dor- sal view. Pachyrhynchus caeruleusYoshitake, 2019 (A, G); P. masatoshii Yoshitake &amp; Yap. 2017 (B, H); P. rukmaneae Barševskis, 2016 (C, I); P. yuukae Yoshitake, 2019 (D, J); P. chrysocyaneus sp. nov. (E, K); P. pseudorukmaneaesp. nov. (F, L). Images courtesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659838" httpUri="https://zenodo.org/record/14659838/files/figure.png" pageId="2" pageNumber="351">Fig. 3H, I, L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="00B7319AFFCFFFFBFF78CD48FDB9FECB" lastPageId="3" lastPageNumber="352" pageId="2" pageNumber="351" type="description">
<paragraph id="48126211FFCFFFFAFF78CD48FEE5FD0B" blockId="2.[151,646,590,1670]" pageId="2" pageNumber="351">
<emphasis id="7AD9BE03FFCFFFFAFF78CD48FE89FDCB" bold="true" box="[151,286,590,614]" pageId="2" pageNumber="351">Description.</emphasis>
<typeStatus id="9716DCB3FFCFFFFAFECBCD48FE1FFDCB" box="[292,392,590,614]" pageId="2" pageNumber="351" type="holotype">Holotype</typeStatus>
. Male. Dimensions: LB: 13.0; LE: 8.7; LP: 4.0; WE: 5,8; WP: 3.9; LR: 2.2; WR: 1.9. N = 1.
</paragraph>
<paragraph id="48126211FFCFFFFAFF78CDA8FF69FCCB" blockId="2.[151,646,590,1670]" pageId="2" pageNumber="351">Integument black, glossy, underside with weak- er luster. Body subovate, with glossy orange, green, yellow, pink and purple scales. Rostrum without scally patch dorsally, moderate patch of green, orange and yellow elongated scales on genae.</paragraph>
<paragraph id="48126211FFCFFFFAFF78CC68FEC2FAE8" blockId="2.[151,646,590,1670]" pageId="2" pageNumber="351">Prothorax with the following yellow to orange scales: 1) broad band of scales along anterior margin in all length; 2) large triangular medial patch along posterior margin, peak connected with anterior band; 3) large lateral patch at each lateroventral part. Each elytron with the following scale markings: 1) longitudinal band of pink scales along suture from middle to apex; 2) longitudinal band along interval II – III in all length, pink scales interrupted by purple scales in middle of band; 3) two additional longitudinal bands, each interrupted by gap from one another, redirected to lateral margin. Each femur with wide band of orange to yellow scales on subapical part.</paragraph>
<paragraph id="48126211FFCFFFFAFF78CA48FF4DF92B" blockId="2.[151,646,590,1670]" pageId="2" pageNumber="351">Head smooth; forehead flattish, without medial furrow; more than two times as wide as eye width; eyes relatively small, slightly convex from outline of the head. Rostrum longer than wide (LR/WR 1.16); deep sub ovate impression at basal half; dorsum smooth, apical half moderately bulging; dorsal contour of rostrum nearly straight, incurved before middle, raised to apex; lateral contour with expressed apical bulge.</paragraph>
<paragraph id="48126211FFCFFFFAFD4ACFC8FBABFE2B" blockId="2.[677,1172,206,742]" pageId="2" pageNumber="351">Antennal scape shorter than funicle; funicular segment 1.5 times as long as wide, longer than II; segment II 1.5 times as long as wide, moderately longer than III; segments III– VII subequal in length, as long as wide; club sub ellipsoidal, two times as long as wide.</paragraph>
<paragraph id="48126211FFCFFFFAFD4ACE88FCCBFD8B" blockId="2.[677,1172,206,742]" pageId="2" pageNumber="351">Pronotum sub spherical, as wide as long (WP/ LP 1.02); dorsal contour widest just before the middle; lateral contour widest at middle; without sub basal groove, subapical groove interrupted at middle.</paragraph>
<paragraph id="48126211FFCFFFFAFD4ACD28FBBAFD6B" blockId="2.[677,1172,206,742]" pageId="2" pageNumber="351">Elytra ovate (LE/WE 1.5), wider than prothorax (WE/WP 1.48), more than twice as long as prothorax, (LE/LP 2.18), smooth, without puncture rows; dorsal contour highest at middle; lateral contour widest at middle.</paragraph>
<paragraph id="48126211FFCFFFFAFD4ACDC8FBC3FD4B" blockId="2.[677,1172,206,742]" box="[677,1108,718,742]" pageId="2" pageNumber="351">
Male genitalia as illustrated (
<figureCitation id="D0967E94FFCFFFFAFC33CDC8FBD1FD4B" box="[988,1094,718,742]" captionStart="Figure 4" captionStartId="5.[113,194,1134,1158]" captionTargetBox="[127,585,528,1043]" captionTargetId="figure-51@5.[113,604,511,1057]" captionTargetPageId="5" captionText="Figure 4. Aedeagus of selected species. Pachyrhynchus caeruleus Yoshitake, 2019 (A, F); P. masatoshii Yoshitake &amp; Yap, 2017 (B, G); P. chrysocyaneus sp. nov. (C, H); P. masatoshiensis sp. nov. (D, I); P. rukmaneae Barševskis, 2016 (E, J). Images courtesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659840" httpUri="https://zenodo.org/record/14659840/files/figure.png" pageId="2" pageNumber="351">Fig. 4D, I</figureCitation>
).
</paragraph>
<caption id="1CD23299FFCFFFFAFD4AC908FC71F90B" ID-DOI="http://doi.org/10.5281/zenodo.14659832" ID-Zenodo-Dep="14659832" httpUri="https://zenodo.org/record/14659832/files/figure.png" pageId="2" pageNumber="351" startId="2.[677,753,1550,1574]" targetBox="[728,1106,750,1529]" targetPageId="2" targetType="figure">
<paragraph id="48126211FFCFFFFAFD4AC908FC71F90B" blockId="2.[677,1173,1550,1702]" pageId="2" pageNumber="351">
<emphasis id="7AD9BE03FFCFFFFAFD4AC908FC99F98B" bold="true" box="[677,782,1550,1574]" pageId="2" pageNumber="351">Figure 1.</emphasis>
<taxonomicName id="8FAD1992FFCFFFFAFCF8C908FBFBF98B" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[791,1132,1550,1574]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="masatoshiensis" status="sp. nov.">
<emphasis id="7AD9BE03FFCFFFFAFCF8C908FBFBF98B" box="[791,1132,1550,1574]" italics="true" pageId="2" pageNumber="351">Pachyrhynchus masatoshiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCFFFFAFB99C908FD6EF9E8" pageId="2" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
habitus of holotype, male (A), female (B);
<taxonomicName id="8FAD1992FFCFFFFAFCEFC949FC29F9CB" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[768,958,1614,1638]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="chrysocyaneus" status="sp. nov.">
<emphasis id="7AD9BE03FFCFFFFAFCEFC949FC29F9CB" box="[768,958,1614,1638]" italics="true" pageId="2" pageNumber="351">P. chrysocyaneus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCFFFFAFC27C948FB88F9C8" box="[968,1055,1614,1637]" pageId="2" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
habitus of holotype, male (C), female (D). Image courtesy A. Rukmane-Bārbale.
</paragraph>
</caption>
<paragraph id="48126211FFCFFFFAFB83C9C2FB03F971" blockId="2.[1132,1172,1732,1756]" box="[1132,1172,1732,1756]" pageId="2" pageNumber="351">351</paragraph>
<paragraph id="48126211FFCEFFFBFDE7CF78FD40FF3C" blockId="3.[520,727,126,145]" box="[520,727,126,145]" pageId="3" pageNumber="352">
<taxonomicName id="8FAD1992FFCEFFFBFDE7CF78FD40FF3C" authority="Rukmane-Barbale" authorityName="Rukmane-Barbale" box="[520,727,126,145]" class="Insecta" family="Notodontidae" genus="Anita" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="7AD9BE03FFCEFFFBFDE7CF78FD40FF3C" box="[520,727,126,145]" italics="true" pageId="3" pageNumber="352">Anita Rukmane-Bārbale</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="48126211FFCEFFFBFF9ECFC8FDB9FECB" blockId="3.[113,608,206,358]" pageId="3" pageNumber="352">
<emphasis id="7AD9BE03FFCEFFFBFF9ECFC8FF5DFF4B" bold="true" box="[113,202,206,230]" pageId="3" pageNumber="352">Female.</emphasis>
(
<figureCitation id="D0967E94FFCEFFFBFF38CFC8FEB1FF4B" box="[215,294,206,230]" captionStart="Figure 1" captionStartId="2.[677,753,1550,1574]" captionTargetBox="[728,1106,750,1529]" captionTargetId="figure-465@2.[727,1123,746,1535]" captionTargetPageId="2" captionText="Figure 1. Pachyrhynchus masatoshiensis sp. nov. habitus of holotype, male (A), female (B); P. chrysocyaneus sp. nov. habitus of holotype, male (C), female (D). Image cour- tesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659832" httpUri="https://zenodo.org/record/14659832/files/figure.png" pageId="3" pageNumber="352">Fig. 1B</figureCitation>
) Dimensions: LB: 13.3 - 15.6 (mean 14.37); LE: 9.2 - 11.0 (mean 9.95); LP: 3.8 - 4.6 (mean 4.1); WE: 7.4 - 7.8 (mean 7.62); WP: 3.9 - 4.3 (mean 4.12); LR: 2.0 - 2.1 (mean 2.075); WR: 1.9 - 2.0 (mean 1.95). N = 4.
</paragraph>
</subSubSection>
<subSubSection id="00B7319AFFCEFFFBFF9ECE88FE12FE4B" pageId="3" pageNumber="352" type="distribution">
<paragraph id="48126211FFCEFFFBFF9ECE88FE12FE4B" blockId="3.[113,608,398,486]" pageId="3" pageNumber="352">
<emphasis id="7AD9BE03FFCEFFFBFF9ECE88FE93FE0B" bold="true" box="[113,260,398,422]" pageId="3" pageNumber="352">Distribution.</emphasis>
The new species is known from two provinces of Luzon Island: General Nakar,
<collectingRegion id="8A69ACF3FFCEFFFBFF9ECEC8FF53FE4B" box="[113,196,462,486]" country="Philippines" name="Quezon" pageId="3" pageNumber="352">Quezon</collectingRegion>
and Bicol,
<collectingRegion id="8A69ACF3FFCEFFFBFED1CEC8FE16FE4B" box="[318,385,462,486]" country="Philippines" name="Albay" pageId="3" pageNumber="352">Albay</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="00B7319AFFCEFFFBFF9ECD08FD81FDCB" pageId="3" pageNumber="352" type="etymology">
<paragraph id="48126211FFCEFFFBFF9ECD08FD81FDCB" blockId="3.[113,608,526,614]" pageId="3" pageNumber="352">
<emphasis id="7AD9BE03FFCEFFFBFF9ECD08FF65FD8B" bold="true" box="[113,242,526,550]" pageId="3" pageNumber="352">Etymology.</emphasis>
The species name
<taxonomicName id="8FAD1992FFCEFFFBFE51CD08FDC8FD8B" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[446,607,526,550]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="masatoshiensis">
<emphasis id="7AD9BE03FFCEFFFBFE51CD08FDC8FD8B" box="[446,607,526,550]" italics="true" pageId="3" pageNumber="352">masatoshiensis</emphasis>
</taxonomicName>
means false
<taxonomicName id="8FAD1992FFCEFFFBFF19CD28FEFBFDEB" authorityName="Barsevskis" authorityYear="2016" box="[246,364,558,582]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="rukmaneae">
<emphasis id="7AD9BE03FFCEFFFBFF19CD28FEFBFDEB" box="[246,364,558,582]" italics="true" pageId="3" pageNumber="352">rukmaneae</emphasis>
</taxonomicName>
, due toits similarity to its
<taxonomicName id="8FAD1992FFCEFFFBFF7ECD49FD81FDCB" authority="Yoshitake &amp; Yap, 2017" authorityName="Yoshitake &amp; Yap" authorityYear="2017" box="[145,534,590,614]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCEFFFBFF7ECD49FD81FDCB" box="[145,534,590,614]" italics="true" pageId="3" pageNumber="352">
P. masatoshii
<bibRefCitation id="2C3C1FE0FFCEFFFBFEC9CD48FD81FDCB" author="Yoshitake H. &amp; Yap S. A." box="[294,534,590,614]" pageId="3" pageNumber="352" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake &amp; Yap, 2017</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<caption id="1CD23299FFCEFFFBFD6FCD08FCDDFC8B" ID-DOI="http://doi.org/10.5281/zenodo.14659834" ID-Zenodo-Dep="14659834" httpUri="https://zenodo.org/record/14659834/files/figure.png" pageId="3" pageNumber="352" startId="3.[640,719,526,550]" targetBox="[642,1121,205,495]" targetPageId="3" targetType="figure">
<paragraph id="48126211FFCEFFFBFD6FCD08FCDDFC8B" blockId="3.[640,1134,526,806]" pageId="3" pageNumber="352">
<emphasis id="7AD9BE03FFCEFFFBFD6FCD08FD66FD8B" bold="true" box="[640,753,526,550]" pageId="3" pageNumber="352">Figure 2.</emphasis>
Dorsal habitus of selected species:
<taxonomicName id="8FAD1992FFCEFFFBFD2ECD28FD2CFDCB" authority="Yoshitake, 2019" authorityName="Yoshitake" authorityYear="2019" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="caeruleus">
<emphasis id="7AD9BE03FFCEFFFBFD2ECD28FC7EFDEB" box="[705,1001,558,582]" italics="true" pageId="3" pageNumber="352">Pachyrhynchus caeruleus</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFC19CD28FD2CFDCB" author="Yoshitake H." pageId="3" pageNumber="352" pagination="22 - 27" refId="ref5764" refString="Yoshitake H. 2019. Two new species of the jewel weevil genus Pachyrhynchus Germar (Coleoptera: Curculionidae: Entiminae) from Luzon, the Philippines. Gekkan-Mushi 576: 22 - 27." type="journal article" year="2019">Yoshitake, 2019</bibRefCitation>
</taxonomicName>
, male (A), female (B);
<taxonomicName id="8FAD1992FFCEFFFBFC3ACD49FCECFD2B" authority="Yoshitake &amp; Yap, 2017" authorityName="Yoshitake &amp; Yap" authorityYear="2017" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCEFFFBFC3ACD49FBF9FDCB" box="[981,1134,590,614]" italics="true" pageId="3" pageNumber="352">P. masatoshii</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFD6FCD68FCECFD2B" author="Yoshitake H. &amp; Yap S. A." box="[640,891,622,646]" pageId="3" pageNumber="352" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake &amp; Yap, 2017</bibRefCitation>
</taxonomicName>
, male (C), female (D);
<taxonomicName id="8FAD1992FFCEFFFBFD6FCD89FC30FD0B" authority="Yoshitake, 2019" authorityName="Yoshitake" authorityYear="2019" box="[640,935,654,678]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="yuukae">
<emphasis id="7AD9BE03FFCEFFFBFD6FCD89FD67FD0B" box="[640,752,654,678]" italics="true" pageId="3" pageNumber="352">P.yuukae</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFD1DCD88FC30FD0B" author="Yoshitake H." box="[754,935,654,678]" pageId="3" pageNumber="352" pagination="22 - 27" refId="ref5764" refString="Yoshitake H. 2019. Two new species of the jewel weevil genus Pachyrhynchus Germar (Coleoptera: Curculionidae: Entiminae) from Luzon, the Philippines. Gekkan-Mushi 576: 22 - 27." type="journal article" year="2019">Yoshitake, 2019</bibRefCitation>
</taxonomicName>
, male (E), female (F);
<taxonomicName id="8FAD1992FFCEFFFBFD41CDA9FC6AFD6B" authority="Barsevskis, 2016" authorityName="Barsevskis" authorityYear="2016" box="[686,1021,686,710]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="rukmaneae">
<emphasis id="7AD9BE03FFCEFFFBFD41CDA9FCA9FD6B" box="[686,830,686,710]" italics="true" pageId="3" pageNumber="352">P. rukmaneae</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFCAACDA8FC6AFD6B" author="Barsevskis A." box="[837,1021,686,710]" pageId="3" pageNumber="352" pagination="1 - 6" refId="ref5388" refString="Barsevskis A. 2016. New species of Pachyrhynchus Germar, 1824 and Macrocyrtus Heller, 1912 (Coleoptera: Curculionidae) from the Marinduque Island (Philippines) as a new example of mimetic species pair. Acta Biologica Universitatis Daugavpiliensis 16 (1): 1 - 6." type="journal article" year="2016">Barševskis, 2016</bibRefCitation>
</taxonomicName>
, male (G), female (H),
<taxonomicName id="8FAD1992FFCEFFFBFCE3CDC9FCADFCAB" authority="Yoshitake, 2017" authorityName="Yoshitake" authorityYear="2017" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="subSpecies" species="rukmaneae" subSpecies="pauscignitus">
<emphasis id="7AD9BE03FFCEFFFBFCE3CDC9FC33FD4B" box="[780,932,718,742]" italics="true" pageId="3" pageNumber="352">P. rukmaneae</emphasis>
ssp.
<emphasis id="7AD9BE03FFCEFFFBFC0DCDC9FBF9FD4B" box="[994,1134,719,742]" italics="true" pageId="3" pageNumber="352">pauscignitus</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFD6FCDE8FCADFCAB" author="Yoshitake H. &amp; Yap S. A." box="[640,826,750,774]" pageId="3" pageNumber="352" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake, 2017</bibRefCitation>
</taxonomicName>
, male. Images courtesy A. Rukmane-Bārbale.
</paragraph>
</caption>
<subSubSection id="00B7319AFFCEFFFCFF9ECD88FBE6FF3C" lastPageId="4" lastPageNumber="353" pageId="3" pageNumber="352" type="discussion">
<paragraph id="48126211FFCEFFFBFF9ECD88FF08FCE8" blockId="3.[113,610,654,837]" pageId="3" pageNumber="352">
<emphasis id="7AD9BE03FFCEFFFBFF9ECD88FF64FD0B" bold="true" box="[113,243,654,678]" pageId="3" pageNumber="352">Comments.</emphasis>
The new species is relatively vari- able in colour and is proposed to be involved in mimicry complex with
<taxonomicName id="8FAD1992FFCEFFFBFE89CDC9FE87FCAB" authority="Yoshitake &amp; Yap, 2017" authorityName="Yoshitake &amp; Yap" authorityYear="2017" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCEFFFBFE89CDC9FE66FD4B" box="[358,497,718,742]" italics="true" pageId="3" pageNumber="352">P. masatoshii</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFE18CDC8FE87FCAB" author="Yoshitake H. &amp; Yap S. A." pageId="3" pageNumber="352" pagination="331 - 341" refId="ref5715" refString="Yoshitake H., Yap S. A. 2017. Four Pachyrhynchini weevils exhibiting allopatric convergence in color and markings, with descriptions of three new taxa from Luzon, the Philippines. Elytra, Tokyo, New Series 7 (2): 331 - 341." type="journal article" year="2017">Yoshitake &amp; Yap, 2017</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="8FAD1992FFCEFFFBFECDCDE9FDC9FCAB" authority="Yoshitake, 2019" authorityName="Yoshitake" authorityYear="2019" box="[290,606,750,774]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="yuukae">
<emphasis id="7AD9BE03FFCEFFFBFECDCDE9FE03FCAB" box="[290,404,750,774]" italics="true" pageId="3" pageNumber="352">P. yuukae</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFE4ECDE8FDC9FCAB" author="Yoshitake H." box="[417,606,750,774]" pageId="3" pageNumber="352" pagination="22 - 27" refId="ref5764" refString="Yoshitake H. 2019. Two new species of the jewel weevil genus Pachyrhynchus Germar (Coleoptera: Curculionidae: Entiminae) from Luzon, the Philippines. Gekkan-Mushi 576: 22 - 27." type="journal article" year="2019">Yoshitake, 2019</bibRefCitation>
</taxonomicName>
(
<figureCitation id="D0967E94FFCEFFFBFF95CC08FEA4FC8B" box="[122,307,782,806]" captionStart="Figure 2" captionStartId="3.[640,719,526,550]" captionTargetBox="[642,1121,205,495]" captionTargetId="figure-134@3.[640,1131,201,515]" captionTargetPageId="3" captionText="Figure 2. Dorsal habitus of selected spe- cies: Pachyrhynchus caeruleus Yoshitake, 2019, male (A), female (B); P. masatoshii Yoshitake &amp; Yap, 2017, male (C), female (D); P.yuukaeYoshitake, 2019, male (E), female (F); P. rukmaneae Barševskis, 2016, male (G), female (H), P. rukmaneae ssp. pauscignitus Yoshitake, 2017, male. Images courtesy A. Rukmane-Bārbale." figureDoi="http://doi.org/10.5281/zenodo.14659834" httpUri="https://zenodo.org/record/14659834/files/figure.png" pageId="3" pageNumber="352">Fig. 2C, D, E, F</figureCitation>
) and
<taxonomicName id="8FAD1992FFCEFFFBFE9ACC09FDAFFC8B" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[373,568,782,806]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="pseudojaysoni" status="sp. nov.">
<emphasis id="7AD9BE03FFCEFFFBFE9ACC09FDAFFC8B" box="[373,568,782,806]" italics="true" pageId="3" pageNumber="352">M. pseudojaysoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCEFFFBFDACCC08FF08FCE8" pageId="3" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<caption id="1CD23299FFCEFFFBFD6FCA88FC5DF92B" ID-DOI="http://doi.org/10.5281/zenodo.14659838" ID-Zenodo-Dep="14659838" httpUri="https://zenodo.org/record/14659838/files/figure.png" pageId="3" pageNumber="352" startId="3.[640,715,1422,1446]" targetBox="[641,1114,858,1388]" targetPageId="3" targetType="figure">
<paragraph id="48126211FFCEFFFBFD6FCA88FC5DF92B" blockId="3.[640,1135,1422,1670]" pageId="3" pageNumber="352">
<emphasis id="7AD9BE03FFCEFFFBFD6FCA88FD72FA0B" bold="true" box="[640,741,1422,1446]" pageId="3" pageNumber="352">Figure 3.</emphasis>
Head and rostrum in lateral and dorsal view.
<taxonomicName id="8FAD1992FFCEFFFBFD0BCAA8FD2FFA4B" authority="Yoshitake, 2019" authorityName="Yoshitake" authorityYear="2019" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="caeruleus">
<emphasis id="7AD9BE03FFCEFFFBFD0BCAA8FC6AFA6B" box="[740,1021,1454,1478]" italics="true" pageId="3" pageNumber="352">Pachyrhynchus caeruleus</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFC11CAA8FD2FFA4B" author="Yoshitake H." pageId="3" pageNumber="352" pagination="22 - 27" refId="ref5764" refString="Yoshitake H. 2019. Two new species of the jewel weevil genus Pachyrhynchus Germar (Coleoptera: Curculionidae: Entiminae) from Luzon, the Philippines. Gekkan-Mushi 576: 22 - 27." type="journal article" year="2019">Yoshitake, 2019</bibRefCitation>
</taxonomicName>
(A, G);
<taxonomicName id="8FAD1992FFCEFFFBFCC2CAC9FD66F9AB" authority="Yoshitake &amp; Yap. 2017" authorityName="Yoshitake &amp; Yap." authorityYear="2017" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="masatoshii">
<emphasis id="7AD9BE03FFCEFFFBFCC2CAC9FC5AFA4B" box="[813,973,1486,1510]" italics="true" pageId="3" pageNumber="352">P. masatoshii</emphasis>
Yoshitake &amp; Yap. 2017
</taxonomicName>
(B, H);
<taxonomicName id="8FAD1992FFCEFFFBFCBDCAE9FD2EF98B" authority="Barsevskis, 2016" authorityName="Barsevskis" authorityYear="2016" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="rukmaneae">
<emphasis id="7AD9BE03FFCEFFFBFCBDCAE9FC7FF9AB" box="[850,1000,1518,1542]" italics="true" pageId="3" pageNumber="352">P. rukmaneae</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFC1DCAE8FD2EF98B" author="Barsevskis A." pageId="3" pageNumber="352" pagination="1 - 6" refId="ref5388" refString="Barsevskis A. 2016. New species of Pachyrhynchus Germar, 1824 and Macrocyrtus Heller, 1912 (Coleoptera: Curculionidae) from the Marinduque Island (Philippines) as a new example of mimetic species pair. Acta Biologica Universitatis Daugavpiliensis 16 (1): 1 - 6." type="journal article" year="2016">Barševskis, 2016</bibRefCitation>
</taxonomicName>
(C, I);
<taxonomicName id="8FAD1992FFCEFFFBFCC8C909FBFAF98B" authority="Yoshitake, 2019" authorityName="Yoshitake" authorityYear="2019" box="[807,1133,1550,1574]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="yuukae">
<emphasis id="7AD9BE03FFCEFFFBFCC8C909FC0AF98B" box="[807,925,1550,1574]" italics="true" pageId="3" pageNumber="352">P. yuukae</emphasis>
<bibRefCitation id="2C3C1FE0FFCEFFFBFC44C908FBFAF98B" author="Yoshitake H." box="[939,1133,1550,1574]" pageId="3" pageNumber="352" pagination="22 - 27" refId="ref5764" refString="Yoshitake H. 2019. Two new species of the jewel weevil genus Pachyrhynchus Germar (Coleoptera: Curculionidae: Entiminae) from Luzon, the Philippines. Gekkan-Mushi 576: 22 - 27." type="journal article" year="2019">Yoshitake, 2019</bibRefCitation>
</taxonomicName>
(D, J);
<taxonomicName id="8FAD1992FFCEFFFBFD34C929FC32F9EB" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[731,933,1582,1606]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="chrysocyaneus" status="sp. nov.">
<emphasis id="7AD9BE03FFCEFFFBFD34C929FC32F9EB" box="[731,933,1582,1606]" italics="true" pageId="3" pageNumber="352">P. chrysocyaneus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCEFFFBFC5FC928FB99F9E8" box="[944,1038,1582,1605]" pageId="3" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
(E, K);
<taxonomicName id="8FAD1992FFCEFFFBFD6FC949FCF2F9CB" authorityName="Rukmane-Bārbale" authorityYear="2024" box="[640,869,1614,1638]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="pseudorukmaneae" status="sp. nov.">
<emphasis id="7AD9BE03FFCEFFFBFD6FC949FCF2F9CB" box="[640,869,1614,1638]" italics="true" pageId="3" pageNumber="352">P. pseudorukmaneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61EA0378FFCEFFFBFC88C948FC28F9C8" box="[871,959,1614,1637]" pageId="3" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
(F, L). Images courtesy A. Rukmane-Bārbale.
</paragraph>
</caption>
<paragraph id="48126211FFCEFFFBFF9EC9C2FF0EF971" blockId="3.[113,153,1732,1756]" box="[113,153,1732,1756]" pageId="3" pageNumber="352">352</paragraph>
<paragraph id="48126211FFC9FFFCFF55CF79FBE6FF3C" blockId="4.[186,1137,126,145]" box="[186,1137,126,145]" pageId="4" pageNumber="353">
<emphasis id="7AD9BE03FFC9FFFCFF55CF79FBE6FF3C" box="[186,1137,126,145]" italics="true" pageId="4" pageNumber="353">
Four new species of
<taxonomicName id="8FAD1992FFC9FFFCFE84CF78FD89FF3C" authority="from" authorityName="FROM" box="[363,542,126,145]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="349" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini from</taxonomicName>
Southern Luzon,
<collectingCountry id="30BA2281FFC9FFFCFD5ACF78FC81FF3C" box="[693,790,126,145]" name="Philippines" pageId="4" pageNumber="353">Philippines</collectingCountry>
(
<taxonomicName id="8FAD1992FFC9FFFCFCCDCF78FC12FF3C" authorityName="Linnaeus" authorityYear="1758" box="[802,901,126,145]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="353" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="8FAD1992FFC9FFFCFC62CF78FB9DFF3C" box="[909,1034,126,145]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
,
<taxonomicName id="8FAD1992FFC9FFFCFBFDCF78FBFCFF3C" box="[1042,1131,126,145]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="subFamily" subFamily="Entiminae">Entiminae</taxonomicName>
)
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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