treatments-xml/data/B8/09/87/B80987C2FFFC117DFF42FB44FCE1EA2A.xml

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<mods:title id="1B9F9688E8E353ECFF09A0E7BF97DD4D">Long-legged flies (Diptera: Dolichopodidae) in the Czech Republic- 2. Corindia and Thrypticus</mods:title>
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<mods:namePart id="8C2A43DF8D0704A61C9FE48CFCEFE84C">Kejval, Zbyněk</mods:namePart>
<mods:affiliation id="884AA56ECF702F92C41A4D83C7A6828F">Muzeum Chodska, Chodské náměstí 96, CZ- 344 01 Domažlice, Czech Republic</mods:affiliation>
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<mods:namePart id="DD8A0156C649C848488F221E8CBC061D">Pollet, Marc</mods:namePart>
<mods:affiliation id="D8DFD699C5C6AEDFD7FAB9424908EA63">Research Institute for Nature and Forest (INBO), Herman Teirlinckgebouw, Havenlaan 88 bus 73, B- 1000 Brussels, Belgium &amp; Royal Belgian Institute of Natural Sciences (RBINS), Operational Directory Taxonomy and Phylogeny, Entomology, Vautierstraat 29, B- 1000 Brussels, Belgium</mods:affiliation>
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<subSubSection id="78BA655FFFFC117EFF42FB44FD62EC7B" box="[151,734,1266,1293]" pageId="1" pageNumber="412" type="nomenclature">
<paragraph id="301F36D4FFFC117EFF42FB44FD62EC7B" blockId="1.[151,734,1266,1293]" box="[151,734,1266,1293]" pageId="1" pageNumber="412">
<heading id="6B5781B8FFFC117EFF42FB44FD62EC7B" bold="true" box="[151,734,1266,1293]" fontSize="11" level="1" pageId="1" pageNumber="412" reason="1">
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<taxonomicName id="F7A04D57FFFC117EFF42FB44FEA5EC7A" ID-CoL="63WCK" authorityName="Gerstacker" authorityYear="1864" box="[151,281,1266,1293]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="412" phylum="Arthropoda" rank="genus">
<emphasis id="02D4EAC6FFFC117EFF42FB44FEA5EC7A" bold="true" box="[151,281,1266,1293]" italics="true" pageId="1" pageNumber="412">Thrypticus</emphasis>
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—host plants, habitats and localities
</emphasis>
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</paragraph>
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<paragraph id="301F36D4FFFC117EFF42FA8DFA2EEF5A" blockId="1.[151,1437,1339,1977]" pageId="1" pageNumber="412">
The larvae of 
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<emphasis id="02D4EAC6FFFC117EFE9CFA8DFE7DEC22" box="[329,449,1339,1365]" italics="true" pageId="1" pageNumber="412">Thrypticus</emphasis>
species
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are plant-miners and form mines within the stems of wetland or aquatic monocotyledons (
<bibRefCitation id="54314B25FFFC117EFEB4FAE9FE62EC0E" author="Dyte, C. E." box="[353,478,1374,1401]" pageId="1" pageNumber="412" pagination="139 - 143" refId="ref10287" refString="Dyte, C. E. (1959) Some interesting habitats of larval Dolichopodidae (Diptera). Entomologist's monthly Magazine, 95, 139 - 143." type="journal article" year="1959">Dyte 1959</bibRefCitation>
; 
<bibRefCitation id="54314B25FFFC117EFE39FAE9FCA1EC0E" author="Bickel, D. J. &amp; Hernandez, M. C." box="[492,797,1375,1401]" pageId="1" pageNumber="412" pagination="437 - 449" refId="ref9987" refString="Bickel, D. J. &amp; Hernandez, M. C. (2004) Neotropical Thrypticus (Diptera: Dolichopodidae) reared from water hyacinth, Eichhornia crassipes, and other Pontederiaceae. Annals of the Entomological Society of America, 97, 437 - 449. https: // doi. org / 10.1603 / 0013 - 8746 (2004) 097 [0437: NTDDRF] 2.0. CO; 2" type="journal article" year="2004">Bickel &amp; Hernandez 2004</bibRefCitation>
; 
<bibRefCitation id="54314B25FFFC117EFCFEFAE9FBEEEC0E" author="Negrobov, O. P. &amp; Naglis, S." box="[811,1106,1375,1401]" pageId="1" pageNumber="412" pagination="111 - 125" refId="ref11375" refString="Negrobov, O. P. &amp; Naglis, S. (2020) Two new species of Thrypticus Gerstacker from Turkey (Diptera, Dolichopodidae), with checklist and key to Palaearctic species. Zootaxa, 4868 (1), 111 - 125. https: // doi. org / 10.11646 / zootaxa. 4858.1.8" type="journal article" year="2020">Negrobov &amp; Naglis 2020</bibRefCitation>
; 
<bibRefCitation id="54314B25FFFC117EFB8AFAE9FA92EC0E" author="Drake, C. M. &amp; Godfrey, A. &amp; Gibbs, D." box="[1119,1326,1375,1401]" pageId="1" pageNumber="412" pagination="172 - 199" refId="ref10227" refString="Drake, C. M., Godfrey, A. &amp; Gibbs, D. J (2023) Thrypticus and Corindia in Britain, with the description of two new species and the addition of two species to the British list (Diptera, Dolichopodidae, Medeterinae). Dipterists Digest, 30, 172 - 199." type="journal article" year="2023">
Drake 
<emphasis id="02D4EAC6FFFC117EFB78FAE9FB54EC0E" box="[1197,1256,1375,1401]" italics="true" pageId="1" pageNumber="412">et al.</emphasis>
2023
</bibRefCitation>
). For this purpose, females possess a strongly scletorized, blade-like oviscapt (
<figureCitation id="A89B2A51FFFC117EFC4BFA35FBA2ECEA" box="[926,1054,1411,1437]" captionStart="FIGURES 11–14. 11–13" captionStartId="7.[151,264,1143,1167]" captionTargetBox="[151,1436,181,1120]" captionTargetId="figure-17@7.[151,1436,181,1120]" captionTargetPageId="7" captionText="FIGURES 11–14. 11–13, Thrypticus habitus, lateral view. 11, T. atomus Frey (female, Horšovský Týn); 12, T. bolevensis sp. nov. (male, holotype); 13, T. cuneatus (Becker) (male, Plzeň-Bolevec); 14, T. atomus (Horšovský Týn, female ovipositor). All photos by Z. Kejval. Scale (1 mm)—Figs 11–13; (0.1 mm)—Fig. 14." figureDoi="http://doi.org/10.5281/zenodo.10906683" httpUri="https://zenodo.org/record/10906683/files/figure.png" pageId="1" pageNumber="412">Figs 11, 14</figureCitation>
) for piercing and ovipositing into plant tissue, a biological behaviour which is unique in the family (
<emphasis id="02D4EAC6FFFC117EFC41FA11FC00ECB7" box="[916,956,1447,1472]" italics="true" pageId="1" pageNumber="412">e.g.</emphasis>
, 
<bibRefCitation id="54314B25FFFC117EFC19FA11FB42ECB6" author="Johannsen, O. A. &amp; Crosby, C. R." box="[972,1278,1447,1473]" pageId="1" pageNumber="412" pagination="164 - 166" refId="ref10766" refString="Johannsen, O. A. &amp; Crosby, C. R. (1913) The life history of Thrypticus muhlenbergiae sp. nov. (Diptera). Psyche, 20, 164 - 166. https: // doi. org / 10.1155 / 1913 / 21361" type="journal article" year="1913">Johannsen &amp; Crosby 1913</bibRefCitation>
; 
<bibRefCitation id="54314B25FFFC117EFAD9FA11FA2BECB6" author="Parent, O." box="[1292,1431,1447,1473]" pageId="1" pageNumber="412" pagination="46 - 69" refId="ref11899" refString="Parent, O. (1924) Essai sur le genre Thrypticus Gerst. Annales de la Societe scientifique de Bruxelles, 44, 46 - 69." type="journal article" year="1924">Parent 1924</bibRefCitation>
; 
<bibRefCitation id="54314B25FFFC117EFF42FA7DFE04EC92" author="Negrobov, O. P. &amp; Naglis, S." box="[151,440,1483,1509]" pageId="1" pageNumber="412" pagination="111 - 125" refId="ref11375" refString="Negrobov, O. P. &amp; Naglis, S. (2020) Two new species of Thrypticus Gerstacker from Turkey (Diptera, Dolichopodidae), with checklist and key to Palaearctic species. Zootaxa, 4868 (1), 111 - 125. https: // doi. org / 10.11646 / zootaxa. 4858.1.8" type="journal article" year="2020">Negrobov &amp; Naglis 2020</bibRefCitation>
). The known host plants for the world fauna belong to the following families and genera: 
<taxonomicName id="F7A04D57FFFC117EFF42FA59FEA6EF7E" authorityName="de Jussieu" authorityYear="1789" box="[151,282,1519,1545]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
(
<taxonomicName id="F7A04D57FFFC117EFEFCFA59FE6BEF7E" box="[297,471,1519,1545]" class="Liliopsida" family="Cyperaceae" genus="Schoenoplectus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFEFCFA59FE6BEF7E" box="[297,471,1519,1545]" italics="true" pageId="1" pageNumber="412">Schoenoplectus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFC117EFE31FA59FDE1EF7E" box="[484,605,1519,1545]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFE31FA59FDE1EF7E" box="[484,605,1519,1545]" italics="true" pageId="1" pageNumber="412">Eleocharis</emphasis>
</taxonomicName>
), 
<taxonomicName id="F7A04D57FFFC117EFDA1FA59FD6CEF7E" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[628,720,1519,1545]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
(
<taxonomicName id="F7A04D57FFFC117EFD0AFA59FCC1EF7E" box="[735,893,1519,1545]" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFD0AFA59FCC1EF7E" box="[735,893,1519,1545]" italics="true" pageId="1" pageNumber="412">Muhlenbergia</emphasis>
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFC117EFC5FFA59FBB7EF7E" box="[906,1035,1519,1545]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFC5FFA59FBB7EF7E" box="[906,1035,1519,1545]" italics="true" pageId="1" pageNumber="412">Phragmites</emphasis>
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFC117EFBCDFA59FBC4EF7E" box="[1048,1144,1519,1545]" class="Liliopsida" family="Poaceae" genus="Spartina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFBCDFA59FBC4EF7E" box="[1048,1144,1519,1545]" italics="true" pageId="1" pageNumber="412">Spartina</emphasis>
</taxonomicName>
), 
<taxonomicName id="F7A04D57FFFC117EFB5AFA59FABCEF7E" box="[1167,1280,1519,1545]" class="Liliopsida" family="Juncaceae" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="family">Juncaceae</taxonomicName>
(
<taxonomicName id="F7A04D57FFFC117EFAC5FA59FAE2EF7F" baseAuthorityName="Hering" baseAuthorityYear="1957" box="[1296,1374,1519,1544]" class="Liliopsida" family="Juncaceae" genus="Juncus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFAC5FA59FAE2EF7F" box="[1296,1374,1519,1544]" italics="true" pageId="1" pageNumber="412">Juncus</emphasis>
</taxonomicName>
), and 
<taxonomicName id="F7A04D57FFFC117EFF42F9A5FEFCEF5A" authorityName="Kunth" authorityYear="1816" box="[151,320,1555,1581]" class="Liliopsida" family="Pontederiaceae" kingdom="Plantae" order="Commelinales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="family">Pontederiaceae</taxonomicName>
(
<taxonomicName id="F7A04D57FFFC117EFE84F9A5FE73EF5A" box="[337,463,1555,1581]" class="Liliopsida" family="Pontederiaceae" genus="Eichhornia" kingdom="Plantae" order="Commelinales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFE84F9A5FE73EF5A" box="[337,463,1555,1581]" italics="true" pageId="1" pageNumber="412">Eichhornia</emphasis>
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFC117EFE08F9A5FDE5EF5A" box="[477,601,1555,1581]" class="Liliopsida" family="Pontederiaceae" genus="Pontederia" kingdom="Plantae" order="Commelinales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFE08F9A5FDE5EF5A" box="[477,601,1555,1581]" italics="true" pageId="1" pageNumber="412">Pontederia</emphasis>
</taxonomicName>
) (
<bibRefCitation id="54314B25FFFC117EFDA4F9A5FD57EF5A" author="Dyte, C. E." box="[625,747,1555,1581]" pageId="1" pageNumber="412" pagination="81 - 84" refId="ref10344" refString="Dyte, C. E. (1993) The occurrence of Thrypticus smaragdinus Gerst. (Diptera: Dolichopodidae) in Britain, with remarks on plant hosts in the genus. The Entomologist, 112, 81 - 84." type="journal article" year="1993">Dyte 1993</bibRefCitation>
; 
<bibRefCitation id="54314B25FFFC117EFD22F9A5FC0EEF5A" author="Hernandez, A. C." box="[759,946,1555,1581]" pageId="1" pageNumber="412" pagination="1041 - 1049" refId="ref10649" refString="Hernandez, A. C. (2008) Biology of Thrypticus truncatus and Thrypticus sagittatus (Diptera: Dolichopodidae), petiole miners of water hyacinth, in Argentina, with morphological descriptions of larvae and pupae. Annals of the Entomological Society of America, 101, 1041 - 1049. https: // doi. org / 10.1603 / 0013 - 8746 - 101.6.1041" type="journal article" year="2008">Hernández 2008</bibRefCitation>
; 
<taxonomicName id="F7A04D57FFFC117EFC6BF9A5FB78EF5A" box="[958,1220,1555,1581]" pageId="1" pageNumber="412">
<emphasis id="02D4EAC6FFFC117EFC6BF9A5FBD0EF5A" box="[958,1132,1555,1581]" italics="true" pageId="1" pageNumber="412">Schoenoplectus</emphasis>
species
</taxonomicName>
listed as 
<taxonomicName id="F7A04D57FFFC117EFAF8F9A5FA3DEF5A" box="[1325,1409,1555,1581]" class="Liliopsida" family="Cyperaceae" genus="Scirpus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFAF8F9A5FA3DEF5A" box="[1325,1409,1555,1581]" italics="true" pageId="1" pageNumber="412">Scirpus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="301F36D4FFFC117EFF12F981FA82EECE" blockId="1.[151,1437,1339,1977]" pageId="1" pageNumber="412">
The bulk of the faunistic data presented below originates from a survey conducted by the first author during 
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16 wetland localities, mainly in western 
<collectingCountry id="48B77644FFFC117EFD29F9EDFCDDEF02" box="[764,865,1627,1653]" name="Czech Republic" pageId="1" pageNumber="412">Bohemia</collectingCountry>
(westernmost part of the 
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). In most cases, the sites were located on the banks of ponds/pools (or bed, if drained). The two sites that received extra attention are described below. Despite the lack of reliable data (no reared specimens), major or at least important genera of host plants for Palaearctic 
<taxonomicName id="F7A04D57FFFC117EFDE2F971FD13EF96" authorityName="Gerstacker" authorityYear="1864" box="[567,687,1735,1761]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="412" phylum="Arthropoda" rank="genus">
<emphasis id="02D4EAC6FFFC117EFDE2F971FD13EF96" box="[567,687,1735,1761]" italics="true" pageId="1" pageNumber="412">Thrypticus</emphasis>
</taxonomicName>
appear to be 
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<emphasis id="02D4EAC6FFFC117EFC9EF971FC45EF96" box="[843,1017,1735,1761]" italics="true" pageId="1" pageNumber="412">Schoenoplectus</emphasis>
</taxonomicName>
and 
<taxonomicName id="F7A04D57FFFC117EFBE7F971FB17EF96" box="[1074,1195,1735,1761]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFBE7F971FB17EF96" box="[1074,1195,1735,1761]" italics="true" pageId="1" pageNumber="412">Eleocharis</emphasis>
</taxonomicName>
. They are both listed by 
<bibRefCitation id="54314B25FFFC117EFF69F95DFEFAEE72" author="Dyte, C. E." box="[188,326,1771,1797]" pageId="1" pageNumber="412" pagination="81 - 84" refId="ref10344" refString="Dyte, C. E. (1993) The occurrence of Thrypticus smaragdinus Gerst. (Diptera: Dolichopodidae) in Britain, with remarks on plant hosts in the genus. The Entomologist, 112, 81 - 84." type="journal article" year="1993">Dyte (1993)</bibRefCitation>
for the species of the northern, temperate part of the Nearctic Region (Maryland, Michigan) and mentioned (both mostly as 
<taxonomicName id="F7A04D57FFFC117EFE16F8B9FDABEE5E" box="[451,535,1807,1833]" class="Liliopsida" family="Cyperaceae" genus="Scirpus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFE16F8B9FDABEE5E" box="[451,535,1807,1833]" italics="true" pageId="1" pageNumber="412">Scirpus</emphasis>
</taxonomicName>
) in a number of the Palaearctic records, 
<emphasis id="02D4EAC6FFFC117EFC06F8B9FC47EE5F" box="[979,1019,1807,1832]" italics="true" pageId="1" pageNumber="412">e.g.</emphasis>
<bibRefCitation id="54314B25FFFC117EFBD4F8B8FB33EE5E" author="Stora, R." box="[1025,1167,1806,1833]" pageId="1" pageNumber="412" pagination="17 - 22" refId="ref12370" refString="Stora, R. (1956) Fynd av sallsynta dipterer i Osterbotten (Om). [Findings of rare diptera in Ostrobothnia (Om)]. Notulae entomologicae, 36, 17 - 22." type="journal article" year="1956">Storå (1956)</bibRefCitation>
, 
<bibRefCitation id="54314B25FFFC117EFB4CF8B9FAA3EE5E" author="Frey, R." box="[1177,1311,1806,1833]" pageId="1" pageNumber="412" pagination="1 - 11" refId="ref10386" refString="Frey, R. (1957) Die Europaischen Thrypticus - Arten (Dipt., Dolichopodidae). Notulae Entomologicae, 37, 1 - 11." type="journal article" year="1957">Frey (1957)</bibRefCitation>
, and 
<bibRefCitation id="54314B25FFFC117EFA8DF8B9FE9BEE3A" author="Drake, C. M. &amp; Godfrey, A. &amp; Gibbs, D." pageId="1" pageNumber="412" pagination="172 - 199" refId="ref10227" refString="Drake, C. M., Godfrey, A. &amp; Gibbs, D. J (2023) Thrypticus and Corindia in Britain, with the description of two new species and the addition of two species to the British list (Diptera, Dolichopodidae, Medeterinae). Dipterists Digest, 30, 172 - 199." type="journal article" year="2023">
Drake 
<emphasis id="02D4EAC6FFFC117EFF42F885FF6DEE3A" box="[151,209,1843,1869]" italics="true" pageId="1" pageNumber="412">et al.</emphasis>
(2023)
</bibRefCitation>
. Accordingly, the herein presented short term survey brought about the best results (both in terms of the number of species and specimens) precisely in those places where plants of both genera grew in abundance. For more details, see the Ecology section of the respective species records. An exception to this rule seems to be 
<taxonomicName id="F7A04D57FFFC117EFA5DF8CDFE51EECE" authority="Gerstacker, 1864" authorityName="Gerstacker" authorityYear="1864" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="412" phylum="Arthropoda" rank="species" species="smaragdinus">
<emphasis id="02D4EAC6FFFC117EFA5DF8CDFE94EECE" italics="true" pageId="1" pageNumber="412">T. smaragdinus</emphasis>
Gerstäcker, 1864
</taxonomicName>
which has mainly been reported from 
<taxonomicName id="F7A04D57FFFC117EFC4BF829FBA3EECE" box="[926,1055,1951,1977]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="1" pageNumber="412" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFC117EFC4BF829FBA3EECE" box="[926,1055,1951,1977]" italics="true" pageId="1" pageNumber="412">Phragmites</emphasis>
</taxonomicName>
vegetations (see further).
</paragraph>
<paragraph id="301F36D4FFFF117DFF12FF20FAC7E8DA" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="413">
<emphasis id="02D4EAC6FFFF117DFF12FF20FE18E9C6" bold="true" box="[199,420,150,177]" pageId="2" pageNumber="413">Bolevecké rybníky</emphasis>
(
<collectingCountry id="48B77644FFFF117DFE67FF21FDA0E9C6" box="[434,540,151,177]" name="Czech Republic" pageId="2" pageNumber="413">Bohemia</collectingCountry>
, 
<geoCoordinate id="55945013FFFF117DFDF3FF2EFD3FE9C4" box="[550,643,152,179]" degrees="49" direction="north" minutes="47" orientation="latitude" pageId="2" pageNumber="413" precision="925" value="49.783333">49°47′N</geoCoordinate>
<geoCoordinate id="55945013FFFF117DFD5CFF2EFD5AE9C4" box="[649,742,152,179]" degrees="13" direction="east" minutes="23" orientation="longitude" pageId="2" pageNumber="413" precision="925" value="13.383333">13°23′E</geoCoordinate>
, altitude 
<quantity id="F7589B31FFFF117DFC9EFF21FC77E9C6" box="[843,971,151,177]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="2" pageNumber="413" unit="m" value="325.0" valueMax="340.0" valueMin="310.0">310–340 m</quantity>
, 
<figureCitation id="A89B2A51FFFF117DFC03FF21FB89E9C6" box="[982,1077,151,177]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1873,1897]" captionTargetBox="[151,1436,887,1850]" captionTargetId="figure-348@2.[151,1436,885,1850]" captionTargetPageId="2" captionText="FIGURES 1–4. Collecting sites in Bohemia (Czech Republic) for Thrypticus species. 1, 2, Plzeň-Bolevec, NW banks of Velký bolevecký rybník, at extremely low (June 2022) and artificially raised water level (July 2023), with abundant Eleocharis palustris, Juncus spp. and Phragmites australis; 3, 4, Kateřina, Soos NNR, salt marshes with extensive Schoenoplectus tabernaemontani stands. All photos by Z. Kejval." figureDoi="http://doi.org/10.5281/zenodo.10906677" httpUri="https://zenodo.org/record/10906677/files/figure.png" pageId="2" pageNumber="413">Figs 1, 2</figureCitation>
)—a system of nine ponds in the northern suburban part of 
<collectingRegion id="F264F836FFFF117DFE62FF0DFE44E9A2" box="[439,504,187,213]" country="Czech Republic" name="Plzensky" pageId="2" pageNumber="413">Plzeň</collectingRegion>
, with a history dating back to the 15th century. These are mostly smaller ponds, with exception of the Velký bolevecký pond, which has an area of approximately 53 ha. 
<taxonomicName id="F7A04D57FFFF117DFB97FF69FB06E98E" authorityName="Gerstacker" authorityYear="1864" box="[1090,1210,223,249]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="genus">
<emphasis id="02D4EAC6FFFF117DFB97FF69FB06E98E" box="[1090,1210,223,249]" italics="true" pageId="2" pageNumber="413">Thrypticus</emphasis>
</taxonomicName>
flies were collected during a dedicated inventory (see above) on the banks of four ponds (Velký bolevecký, Kamenný, Senecký and Nováček pond), with 
<taxonomicName id="F7A04D57FFFF117DFE5EFE91FD86E836" box="[395,570,295,321]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="02D4EAC6FFFF117DFE5EFE91FDB8E836" box="[395,516,295,321]" italics="true" pageId="2" pageNumber="413">Eleocharis</emphasis>
spp.
</taxonomicName>
(mostly 
<taxonomicName id="F7A04D57FFFF117DFD48FE91FC14E836" authority="(Linnaeus))" baseAuthorityName="Linnaeus" box="[669,936,295,321]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="palustris">
<emphasis id="02D4EAC6FFFF117DFD48FE91FCA3E836" box="[669,799,295,321]" italics="true" pageId="2" pageNumber="413">E. palustris</emphasis>
(Linnaeus))
</taxonomicName>
and/or 
<taxonomicName id="F7A04D57FFFF117DFC2AFE91FA25E836" authority="(Linnaeus)" baseAuthorityName="Linnaeus" box="[1023,1433,295,321]" class="Liliopsida" family="Cyperaceae" genus="Schoenoplectus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="lacustris">
<emphasis id="02D4EAC6FFFF117DFC2AFE91FAA8E836" box="[1023,1300,295,321]" italics="true" pageId="2" pageNumber="413">Schoenoplectus lacustris</emphasis>
(Linnaeus)
</taxonomicName>
. Most records (five species and the majority of specimens) originate from the Velký bolevecký pond, which has the following characteristics (largely shared by other ponds, 
<figureCitation id="A89B2A51FFFF117DFCF7FED9FC36E8FE" box="[802,906,367,393]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1873,1897]" captionTargetBox="[151,1436,887,1850]" captionTargetId="figure-348@2.[151,1436,885,1850]" captionTargetPageId="2" captionText="FIGURES 1–4. Collecting sites in Bohemia (Czech Republic) for Thrypticus species. 1, 2, Plzeň-Bolevec, NW banks of Velký bolevecký rybník, at extremely low (June 2022) and artificially raised water level (July 2023), with abundant Eleocharis palustris, Juncus spp. and Phragmites australis; 3, 4, Kateřina, Soos NNR, salt marshes with extensive Schoenoplectus tabernaemontani stands. All photos by Z. Kejval." figureDoi="http://doi.org/10.5281/zenodo.10906677" httpUri="https://zenodo.org/record/10906677/files/figure.png" pageId="2" pageNumber="413">Figs 1, 2</figureCitation>
): oligotrophic lake-like pond with limited fish stock, and well-developed macrophyte cover, having shallow littoral and largely sandy banks (
<bibRefCitation id="54314B25FFFF117DFB6DFE24FAD3E8DA" author="Juza, T. &amp; Duras, J. &amp; Blabolil, P. &amp; Sajdlova, Z. &amp; Hess, J. &amp; Chocholouskova, Z. &amp; Kubecka, J." box="[1208,1391,402,429]" pageId="2" pageNumber="413" pagination="167 - 176" refId="ref10840" refString="Juza, T., Duras, J., Blabolil, P., Sajdlova, Z., Hess, J., Chocholouskova, Z. &amp; Kubecka, J. (2019) Recovery of the Velky Bolevecky pond (Plzen, Czech Republic) via biomanipulation - Key study for management. Ecological Engineering, 136, 167 - 176. https: // doi. org / 10.1016 / j. ecoleng. 2019.06.025" type="journal article" year="2019">
Jůza 
<emphasis id="02D4EAC6FFFF117DFB26FE25FA97E8DA" box="[1267,1323,403,429]" italics="true" pageId="2" pageNumber="413">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
<paragraph id="301F36D4FFFF117DFF12FE01FBCAE882" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="413">
The species collected: 
<taxonomicName id="F7A04D57FFFF117DFE12FE01FD35E8A6" authorityName="Loew" authorityYear="1869" box="[455,649,439,465]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="02D4EAC6FFFF117DFE12FE01FD35E8A6" box="[455,649,439,465]" italics="true" pageId="2" pageNumber="413">Thrypticus bellus</emphasis>
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFF117DFD4DFE01FC95E8A6" authority="Kejval &amp; Pollet, 2024" authorityName="Kejval &amp; Pollet" authorityYear="2024" box="[664,809,439,465]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="bolevensis" status="sp. nov.">
<emphasis id="02D4EAC6FFFF117DFD4DFE01FC95E8A6" box="[664,809,439,465]" italics="true" pageId="2" pageNumber="413">T. bolevensis</emphasis>
</taxonomicName>
<emphasis id="02D4EAC6FFFF117DFCE5FE01FC36E8A6" bold="true" box="[816,906,439,465]" pageId="2" pageNumber="413">
<taxonomicNameLabel id="19E757BDFFFF117DFCE5FE01FC36E8A6" box="[816,906,439,465]" pageId="2" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="F7A04D57FFFF117DFC4CFE01FB76E8A6" authority="(Becker, 1917)" baseAuthorityName="Becker" baseAuthorityYear="1917" box="[921,1226,439,465]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="cuneatus">
<emphasis id="02D4EAC6FFFF117DFC4CFE01FBA5E8A7" box="[921,1049,439,464]" italics="true" pageId="2" pageNumber="413">T. cuneatus</emphasis>
(Becker, 1917)
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFF117DFB03FE01FF67E882" authority="(Strobl, 1880)" baseAuthorityName="Strobl" baseAuthorityYear="1880" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="divisus">
<emphasis id="02D4EAC6FFFF117DFB03FE01FAFCE8A6" box="[1238,1344,439,465]" italics="true" pageId="2" pageNumber="413">T. divisus</emphasis>
(Strobl, 1880)
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFF117DFF32FE6DFE5DE882" authority="Verrall, 1912" authorityName="Verrall" authorityYear="1912" box="[231,481,475,501]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="02D4EAC6FFFF117DFF32FE6DFEF9E882" box="[231,325,475,501]" italics="true" pageId="2" pageNumber="413">T. laetus</emphasis>
<bibRefCitation id="54314B25FFFF117DFE9EFE6DFE5DE882" author="Verrall, G. H." box="[331,481,475,501]" pageId="2" pageNumber="413" pagination="20 - 27" refId="ref12439" refString="Verrall, G. H. (1912) Another hundred new British species of Diptera. Entomologist's monthly Magazine, 48, 20 - 27 + 56 - 59 + 144 - 147 + 190 - 198." type="journal article" year="1912">Verrall, 1912</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFF117DFE39FE6DFCADE882" authority="Wood, 1913" authorityName="Wood" authorityYear="1913" box="[492,785,475,501]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="02D4EAC6FFFF117DFE39FE6DFD3EE882" box="[492,642,475,501]" italics="true" pageId="2" pageNumber="413">T. nigricauda</emphasis>
Wood, 1913
</taxonomicName>
and 
<taxonomicName id="F7A04D57FFFF117DFC92FE6DFBCDE882" authority="Verrall, 1912" authorityName="Verrall" authorityYear="1912" box="[839,1137,475,501]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="pollinosus">
<emphasis id="02D4EAC6FFFF117DFC92FE6DFC6AE882" box="[839,982,475,501]" italics="true" pageId="2" pageNumber="413">T. pollinosus</emphasis>
<bibRefCitation id="54314B25FFFF117DFC09FE6DFBCDE882" author="Verrall, G. H." box="[988,1137,475,501]" pageId="2" pageNumber="413" pagination="20 - 27" refId="ref12439" refString="Verrall, G. H. (1912) Another hundred new British species of Diptera. Entomologist's monthly Magazine, 48, 20 - 27 + 56 - 59 + 144 - 147 + 190 - 198." type="journal article" year="1912">Verrall, 1912</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="301F36D4FFFF117DFF12FE48FAADEA62" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="413">
<emphasis id="02D4EAC6FFFF117DFF12FE48FF41EB6F" bold="true" box="[199,253,510,536]" pageId="2" pageNumber="413">Soos</emphasis>
(
<collectingCountry id="48B77644FFFF117DFEC4FE49FEC7EB6E" box="[273,379,511,537]" name="Czech Republic" pageId="2" pageNumber="413">Bohemia</collectingCountry>
, National Nature Reserve, 
<geoCoordinate id="55945013FFFF117DFD6BFDB6FCFEEB6C" box="[702,834,512,539]" degrees="50" direction="north" minutes="08" orientation="latitude" pageId="2" pageNumber="413" precision="15" seconds="53" value="50.148056">50°08′53″N</geoCoordinate>
<geoCoordinate id="55945013FFFF117DFC9AFDB6FC6FEB6C" box="[847,979,512,539]" degrees="12" direction="east" minutes="24" orientation="longitude" pageId="2" pageNumber="413" precision="15" seconds="24" value="12.406667">12°24′24″E</geoCoordinate>
, altitude 
<quantity id="F7589B31FFFF117DFB91FE49FB2EEB6E" box="[1092,1170,511,537]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="413" unit="m" value="440.0">440 m</quantity>
, 
<figureCitation id="A89B2A51FFFF117DFB76FE49FAB2EB6E" box="[1187,1294,511,537]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1873,1897]" captionTargetBox="[151,1436,887,1850]" captionTargetId="figure-348@2.[151,1436,885,1850]" captionTargetPageId="2" captionText="FIGURES 1–4. Collecting sites in Bohemia (Czech Republic) for Thrypticus species. 1, 2, Plzeň-Bolevec, NW banks of Velký bolevecký rybník, at extremely low (June 2022) and artificially raised water level (July 2023), with abundant Eleocharis palustris, Juncus spp. and Phragmites australis; 3, 4, Kateřina, Soos NNR, salt marshes with extensive Schoenoplectus tabernaemontani stands. All photos by Z. Kejval." figureDoi="http://doi.org/10.5281/zenodo.10906677" httpUri="https://zenodo.org/record/10906677/files/figure.png" pageId="2" pageNumber="413">Figs 3, 4</figureCitation>
)—protected wetland area with the remains of former volcanic activity, located approximately 
<quantity id="F7589B31FFFF117DFB94FD95FBC0EB4A" box="[1089,1148,547,573]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="413" unit="km" value="6.0">6 km</quantity>
northeast of Františkovy Lázně. A spectacular peatbog site with fens on numerous mineral spring effluents, emanations of pure carbon dioxide in mofettes, saltwater wetlands, and a diatomite shield in its central part (
<bibRefCitation id="54314B25FFFF117DFBC6FDDDFB6EEBF2" author="Horackova, J." box="[1043,1234,619,645]" pageId="2" pageNumber="413" pagination="54 - 57" refId="ref10716" refString="Horackova, J. (2012) Mekkysi Narodni prirodni rezervace Soos. (Molluscs (Mollusca) of the Soos National Nature Reserve). Malacologica Bohemoslovaca, 11, 54 - 57. https: // doi. org / 10.5817 / MaB 2012 - 11 - 54" type="journal article" year="2012">Horáčková 2012</bibRefCitation>
). The major saline habitats are halophytic common reed stands and salt marsh grasslands, with rare plant species, 
<emphasis id="02D4EAC6FFFF117DFB6EFD39FB5FEBDF" box="[1211,1251,655,680]" italics="true" pageId="2" pageNumber="413">e.g.</emphasis>
<taxonomicName id="F7A04D57FFFF117DFB3FFD39FF42EBBA" authority="Linnaeus" authorityName="Linnaeus" class="Liliopsida" family="Pontederiaceae" genus="Glaux" kingdom="Plantae" order="Commelinales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="maritima">
<emphasis id="02D4EAC6FFFF117DFB3FFD39FA21EBDF" box="[1258,1437,655,681]" italics="true" pageId="2" pageNumber="413">Glaux maritima</emphasis>
Linnaeus
</taxonomicName>
and 
<taxonomicName id="F7A04D57FFFF117DFEE6FD05FC1EEBBA" authority="Jacquin (Brabec &amp; Velebil 2019)" authorityName="Jacquin (Brabec &amp; Velebil" authorityYear="2019" box="[307,930,691,717]" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="parviflora">
<emphasis id="02D4EAC6FFFF117DFEE6FD05FD97EBBA" box="[307,555,691,717]" italics="true" pageId="2" pageNumber="413">Scorzonera parviflora</emphasis>
Jacquin (
<bibRefCitation id="54314B25FFFF117DFD41FD05FC26EBBA" author="Brabec, J. &amp; Velebil, J." box="[660,922,691,717]" pageId="2" pageNumber="413" pagination="113 - 116" refId="ref10097" refString="Brabec, J. &amp; Velebil, J. (2019) Za kvety Soosu. I. (Flowers of Soos. I. Halophytic habitats). Ziva, 3, 113 - 116. [https: // ziva. avcr. cz / 2019 - 3 / za-kvety-soosu-i-slaniska. html]" type="journal article" year="2019">Brabec &amp; Velebil 2019</bibRefCitation>
)
</taxonomicName>
. Of the two major genera of host plants, there are extensive, largely monodominant stands of 
<taxonomicName id="F7A04D57FFFF117DFD73FD61FBA6EB86" box="[678,1050,727,753]" class="Liliopsida" family="Cyperaceae" genus="Schoenoplectus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="tabernaemontani">
<emphasis id="02D4EAC6FFFF117DFD73FD61FBA6EB86" box="[678,1050,727,753]" italics="true" pageId="2" pageNumber="413">Schoenoplectus tabernaemontani</emphasis>
</taxonomicName>
whereas 
<taxonomicName id="F7A04D57FFFF117DFB56FD61FB40EB86" box="[1155,1276,727,753]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="genus">
<emphasis id="02D4EAC6FFFF117DFB56FD61FB40EB86" box="[1155,1276,727,753]" italics="true" pageId="2" pageNumber="413">Eleocharis</emphasis>
</taxonomicName>
is also locally abundant, with at least two species, 
<taxonomicName id="F7A04D57FFFF117DFDFFFD4DFD17EA62" box="[554,683,763,789]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="palustris">
<emphasis id="02D4EAC6FFFF117DFDFFFD4DFD17EA62" box="[554,683,763,789]" italics="true" pageId="2" pageNumber="413">E. palustris</emphasis>
</taxonomicName>
and 
<taxonomicName id="F7A04D57FFFF117DFD37FD4DFC7CEA62" authority="(Link)" baseAuthorityName="Link" box="[738,960,763,789]" class="Liliopsida" family="Cyperaceae" genus="Eleocharis" kingdom="Plantae" order="Poales" pageId="2" pageNumber="413" phylum="Tracheophyta" rank="species" species="uniglumis">
<emphasis id="02D4EAC6FFFF117DFD37FD4DFCD3EA62" box="[738,879,763,789]" italics="true" pageId="2" pageNumber="413">E. uniglumis</emphasis>
(Link)
</taxonomicName>
, see 
<bibRefCitation id="54314B25FFFF117DFC23FD4DFAB1EA62" author="Brabec, J. &amp; Velebil, J." box="[1014,1293,763,789]" pageId="2" pageNumber="413" refId="ref10062" refString="Brabec, J. &amp; Velebil, J., Eds. (2018) Kvetena Soosu a okoli [Flora of Soos and environs]. HB print, Cheb, 421 pp." type="book" year="2018">Brabec &amp; Velebil (2018)</bibRefCitation>
.
</paragraph>
<paragraph id="301F36D4FFFF117DFF12FCA9FCE1EA2A" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="413">
The species collected: 
<taxonomicName id="F7A04D57FFFF117DFE04FCA9FC90EA4E" authority="Loew, 1869" authorityName="Loew" authorityYear="1869" box="[465,812,799,825]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="02D4EAC6FFFF117DFE04FCA9FD2AEA4E" box="[465,662,799,825]" italics="true" pageId="2" pageNumber="413">Thrypticus bellus</emphasis>
Loew, 1869
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFF117DFCEEFCA9FB1CEA4E" authority="Negrobov, 1967" authorityName="Negrobov" authorityYear="1967" box="[827,1184,799,825]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="intercedens">
<emphasis id="02D4EAC6FFFF117DFCEEFCA9FC66EA4E" box="[827,986,799,825]" italics="true" pageId="2" pageNumber="413">T. intercedens</emphasis>
Negrobov, 1967
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFF117DFB7AFCA9FF6EEA2A" authority="Wood, 1913" authorityName="Wood" authorityYear="1913" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="nigricauda">
<emphasis id="02D4EAC6FFFF117DFB7AFCA9FAF5EA4E" box="[1199,1353,799,825]" italics="true" pageId="2" pageNumber="413">T. nigricauda</emphasis>
Wood, 1913
</taxonomicName>
, 
<taxonomicName id="F7A04D57FFFF117DFF08FCF5FE5DEA2A" authority="Parent, 1932" authorityName="Parent" authorityYear="1932" box="[221,481,835,861]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="02D4EAC6FFFF117DFF08FCF5FEF1EA2A" box="[221,333,835,861]" italics="true" pageId="2" pageNumber="413">T. tarsalis</emphasis>
Parent, 1932
</taxonomicName>
and 
<taxonomicName id="F7A04D57FFFF117DFDCDFCF5FCE4EA2A" authority="Negrobov, 1967" authorityName="Negrobov" authorityYear="1967" box="[536,856,835,861]" class="Insecta" family="Dolichopodidae" genus="Thrypticus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="02D4EAC6FFFF117DFDCDFCF5FD26EA2B" box="[536,666,835,860]" italics="true" pageId="2" pageNumber="413">T. virescens</emphasis>
Negrobov, 1967
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>