185 lines
21 KiB
XML
185 lines
21 KiB
XML
<document ID-CLB-Dataset="263454" ID-DOI="http://dx.doi.org/10.3897/zookeys.1174.103324" ID-GBIF-Dataset="7797aa47-8d4c-45a9-a2e8-6fc406472d3c" ID-PMC="PMC10848836" ID-Pensoft-Pub="1313-2970-1174-85" ID-Pensoft-UUID="C382EDC7029755EFA1BC59D090835B37" ID-PubMed="38328308" ID-ZooBank="5C6509E838A34F6A878B08C6868E0D2A" ModsDocID="1313-2970-1174-85" checkinTime="1691559543624" checkinUser="pensoft" docAuthor="McKamey, Stuart H." docDate="2023" docId="B5E38D40232C5BBC8AC11C03E9D715F7" docLanguage="en" docName="ZooKeys 1174: 85-95" docOrigin="ZooKeys 1174" docPubDate="2023-08-08" docSource="http://dx.doi.org/10.3897/zookeys.1174.103324" docTitle="Smilidarnis erwini McKamey 2023, sp. nov." docType="treatment" docUuid="DF05FB0E-5FC5-47EE-9F82-3160A941B6A4" docUuidSource="ZooBank" docVersion="4" id="C382EDC7029755EFA1BC59D090835B37" lastPageNumber="85" masterDocId="C382EDC7029755EFA1BC59D090835B37" masterDocTitle="Three new species of the Neotropical genus Smilidarnis Andrade (Hemiptera, Membracidae)" masterLastPageNumber="95" masterPageNumber="85" pageNumber="85" updateTime="1732480995221" updateUser="ExternalLinkService">
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<mods:mods id="D0B470A95612838CC3A9E91A4A052178" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D9C290D1083E4BC2D82652664A7E1D52">
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<mods:title id="1929E4FB3D8668A7F80389D23984366F">Three new species of the Neotropical genus Smilidarnis Andrade (Hemiptera, Membracidae)</mods:title>
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<mods:name id="EAC19A8DA5242637C819B9E9DD5A74CD" type="personal">
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<mods:namePart id="F4FF5DCF60AD69CC58D493BF2F8CC045">McKamey, Stuart H.</mods:namePart>
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<mods:nameIdentifier id="D055BD7029D755571953E04B7FB5B78C" type="ORCID">https://orcid.org/0000-0002-9617-0594</mods:nameIdentifier>
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<mods:affiliation id="2998079F67063900EFF0DC93C6A063D7">Systematic Entomology Laboratory, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum of Natural History, P. O. Box 37012, Washington, D. C. 20013, USA</mods:affiliation>
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<mods:nameIdentifier id="FF3D92056C9552235FCA2B9FF18D3A72" type="email">stuart.mckamey@usda.gov</mods:nameIdentifier>
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<mods:typeOfResource id="ADDF6EFA30839E2984CE3338F83773C6">text</mods:typeOfResource>
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<mods:title id="8093012E7563BED454D4F71E09D2F502">ZooKeys</mods:title>
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<mods:part id="35A31A29C5340616076ABD179E801FDF">
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<mods:date id="0204CCAC41552EDB8EF3F0457B466FB6">2023</mods:date>
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<mods:number id="2BB05303009BEC5F09569F0F75E2DD1D">2023-08-08</mods:number>
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<mods:number id="D2DA73D5B71FF73AC812757B021D2AF6">1174</mods:number>
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<mods:start id="19EC1026E8A4E80E7E7841A8E45B0336">85</mods:start>
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<mods:url id="82FACD34D7F501EE78FC39232B5AD12C">http://dx.doi.org/10.3897/zookeys.1174.103324</mods:url>
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<mods:classification id="E58CBAF2B073E828191F3CB5FFFE01D2">journal article</mods:classification>
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<mods:identifier id="BD118B3357DD05B743C25B2465E55AA2" type="DOI">http://dx.doi.org/10.3897/zookeys.1174.103324</mods:identifier>
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<mods:identifier id="C889BF1EEF00DF9D04DA9CF2D55B39BE" type="ZooBank">5C6509E838A34F6A878B08C6868E0D2A</mods:identifier>
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<treatment id="B5E38D40232C5BBC8AC11C03E9D715F7" ID-GBIF-Taxon="212426714" LSID="urn:lsid:zoobank.org:act:DF05FB0E-5FC5-47EE-9F82-3160A941B6A4" httpUri="http://treatment.plazi.org/id/B5E38D40232C5BBC8AC11C03E9D715F7" lastPageNumber="85" pageId="0" pageNumber="85" scope_family="Membracidae" scope_order="Hemiptera">
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<subSubSection id="42CBC7591E48341627BC9CFB276DB4E6" pageId="0" pageNumber="85" type="nomenclature">
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<paragraph id="3E046A5EF7A4E90DFDB148689AD987EE" pageId="0" pageNumber="85">
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<taxonomicName id="5E56858E350AEA9BD63B39267726F64F" ID-CoL="CMTLG" LSID="https://zoobank.org/DF05FB0E-5FC5-47EE-9F82-3160A941B6A4" authority="McKamey, 2023" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis erwini" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="erwini" status="sp. nov.">Smilidarnis erwini</taxonomicName>
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<taxonomicNameLabel id="5CCF9EDAC348EDBFA931FA551DDC8828" pageId="0" pageNumber="85">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="3CCFFC5B36B985E755CF9486FCCA890C" pageId="0" pageNumber="85" type="description">
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<paragraph id="C46DADABFAB138B3F7D30022881E5C7F" pageId="0" pageNumber="85">
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<figureCitation id="33EBB855E17596B6912B194C7DA5EE9A" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">Figs 8-13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FBE5FDEF2AD4CFF36EAF93C79F96A0D9" pageId="0" pageNumber="85" type="diagnosis">
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<paragraph id="AA2CFF55667CCA57917C6E54D6A54F06" pageId="0" pageNumber="85">Diagnosis.</paragraph>
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<paragraph id="08732B820C23FE8AF1543736CCB78781" pageId="0" pageNumber="85">Pronotum without pair of suprahumeral spines; vertex of head with four dark lines, outer pair adjacent to eyes and inner pair ventrally converging.</paragraph>
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</subSubSection>
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<subSubSection id="FA3DE7D5B09D2656F7C126CDC86D6B63" pageId="0" pageNumber="85" type="description">
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<paragraph id="CDC0BCA61DB071011081E6BB34FDC3CD" pageId="0" pageNumber="85">Description of male.</paragraph>
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<paragraph id="F9D2E6355B6D60085F8DFF644F662F46" pageId="0" pageNumber="85">
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Dimensions (mm). Length of pronotum 8.5; length including wings in repose 10.0; width across humeri 3.7; width across posterolateral spine apices 1.5; height in anterior view 4.2. Head (Fig.
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<figureCitation id="95C645905B04D5A48E35BF926E96D5B9" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">9</figureCitation>
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). Ocelli circular, above imaginary middle line between eyes; distance to eye 0.8
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<normalizedToken id="D0149067A7FCFA9B249756FFDF4664CA" originalValue="×">x</normalizedToken>
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distance between ocelli; vertex with adjacent depression ventrolaterally; frontoclypeus with indistinct sutures and evenly convex in line with ventrolateral margins. Pronotum (Figs
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<figureCitation id="CB60BF55384FDAD488266846B55891DF" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">8-10</figureCitation>
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). Humeral angles rounded, only slightly produced laterally from wing bases. Forewing (as in Fig.
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<figureCitation id="9E4B307FB48DE6D09063E4EE3E27C732" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Smilidarnis robustus sp. nov., forewing and hind wing." figureDoi="10.3897/zookeys.1174.103324.figure1" httpUri="https://binary.pensoft.net/fig/890186" pageId="0" pageNumber="85">1</figureCitation>
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) R and M not confluent for short distance near apex. Leg chaetotaxy. Femora of pro- and metathorax lacking cucullate setae and spines (other femora missing); metathoracic tibiae missing. Abdominal terga without pits, fossae, or digitate processes.
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</paragraph>
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</subSubSection>
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<subSubSection id="A7C54386A207FE2A4D725E7FFA482BB3" pageId="0" pageNumber="85" type="male terminalia">
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<paragraph id="D05A9A89759AD4024B5986488EB1A515" pageId="0" pageNumber="85">Male terminalia</paragraph>
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<paragraph id="A4E7F564186033FBF91A8B6B72AEF924" pageId="0" pageNumber="85">
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(Figs
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<figureCitation id="9F4334C160ED5FF856DB4C4A75507975" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">11-13</figureCitation>
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). Pygofer (Fig.
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<figureCitation id="9E445D9C50E5B8B660DF7B6297FC02B3" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">11</figureCitation>
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) including lateral plate subquadrate, lateral plate unarmed, ovate, pilose throughout, subgenital plate (Fig.
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<figureCitation id="D4000ACD50DAFCD0C2F61D61DDCBA020" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">13</figureCitation>
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) fused basally, its sides subparallel, slightly narrowing distally, length in ventral view about 2.3
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<normalizedToken id="52E0B585D0691CB83E4CB72D04317E95" originalValue="×">x</normalizedToken>
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width; style (Figs
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<figureCitation id="A4770C8729B1A5CB8723FC714874E40D" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">11</figureCitation>
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,
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<figureCitation id="4A164FDC68076F2A41806AF5BCF98E71" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">12</figureCitation>
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) recurved with short acute apex; aedeagus (Figs
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<figureCitation id="98B28AB92B50DFB0B2ADF64AF35C6ADA" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">11</figureCitation>
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,
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<figureCitation id="E2EC92F81FAEC14FF9567840E5FC675E" captionStart="Figures 8–13" captionStartId="F3" captionText="Figures 8 - 13. Smilidarnis erwini sp. nov. 8 - 10 habitus in dorsal, anterior, and lateral views, respectively 11 pygofer, styles, and aedeagus, lateral view 12 aedeagus and styles, anterior view 13 subgenital plate, ventral view. lp lateral plate." figureDoi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85">12</figureCitation>
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) U-shaped in lateral view, shaft subparallel in anterior and lateral views, lacking dentae or other texture on distoanterior portion; gonopore dorsal. Color. Pale brown throughout except 4 dark lines on head vertex (outer pair adjacent to eyes, inner pair overlapping ocelli and ventrally convergent), 1 large median, dark brown, anteriorly divergent, V-shaped mark on pronotal metopidium, all 3 apical spines with bases and apices black but pale at mid length.
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</paragraph>
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<caption id="348DFBCEC0848F61D8434BCFDF49F4C9" doi="10.3897/zookeys.1174.103324.figures8-13" httpUri="https://binary.pensoft.net/fig/890188" pageId="0" pageNumber="85" start="Figures 8–13" startId="F3">
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<paragraph id="D92CB96386B1D06A32964C4E6E7EC64D" pageId="0" pageNumber="85">
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<emphasis id="237F63ACBC37179D43709ABA7200007E" bold="true" pageId="0" pageNumber="85">Figures 8-13.</emphasis>
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<taxonomicName id="120301DA58D0991ACAA2E000F78FBA80" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis erwini" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="erwini">
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<emphasis id="B8E670C3132BF58678A43CA3548CDF21" italics="true" pageId="0" pageNumber="85">Smilidarnis erwini</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="BB6E60E8BFCF05EE19B3BCA938CACE5A" bold="true" pageId="0" pageNumber="85">8-10</emphasis>
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habitus in dorsal, anterior, and lateral views, respectively
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<emphasis id="2949952FEAFDA425455B68C95171693D" bold="true" pageId="0" pageNumber="85">11</emphasis>
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pygofer, styles, and aedeagus, lateral view
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<emphasis id="744DD13F2CD8187E3867994951AE8CD2" bold="true" pageId="0" pageNumber="85">12</emphasis>
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aedeagus and styles, anterior view
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<emphasis id="5BB747DF5858EC9580D98617ABD60ABE" bold="true" pageId="0" pageNumber="85">13</emphasis>
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subgenital plate, ventral view.
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<emphasis id="86E87603A7BBEE2FF9C919CEB7E94E6C" italics="true" pageId="0" pageNumber="85">lp</emphasis>
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lateral plate.
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</paragraph>
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</caption>
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<paragraph id="4D01DD29811EAFD4E4C1A576DA0EE12B" pageId="0" pageNumber="85">
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<emphasis id="92EA19EEF6BCB4983B0819E55C5FAF7B" bold="true" pageId="0" pageNumber="85">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="8C803733987E232EE4050B2DF3643CE9" pageId="0" pageNumber="85" type="materials_examined">
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<paragraph id="58C7087369E870C61D6B10F3B5D68BFC" pageId="0" pageNumber="85">Material examined.</paragraph>
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<paragraph id="C44C4830748C505FA8CFEC9F098DE00C" pageId="0" pageNumber="85">
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<materialsCitation id="6BD85C62B1CA137EBF5D3F939784C6DF" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="S. OnkoneGare Camp Transect Ent, S. H. McKamey" country="ECUADOR" elevation="216" latitude="-0.6571389" location="Reserva Etnica Waorani" longLatPrecision="1" longitude="-76.452995" specimenCount="♂" stateProvince="ORELLANA" typeStatus="Holotype">
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<emphasis id="F82531997C91578941DB3CACA89FA715" bold="true" italics="true" pageId="0" pageNumber="85">
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<typeStatus id="570FF639C4E1E88EDD2DD0C310101FCF">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="52855F5AD9A7709FABC032BEA070718B">♂</specimenCount>
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(EPNC) with labels "
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<collectingCountry id="CB170965AE54CB9299C11229ABB0E7D1" name="Ecuador">ECUADOR</collectingCountry>
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:
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<collectingRegion id="48142750776890E3A17A4D081C5651C2" country="Ecuador" name="Orellana">ORELLANA</collectingRegion>
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:
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<location id="B740CDF5670DE8DDE43957DE32D87ADF" LSID="urn:lsid:plazi:treatment:B5E38D40232C5BBC8AC11C03E9D715F7:B740CDF5670DE8DDE43957DE32D87ADF" country="ECUADOR" latitude="-0.6571389" longLatPrecision="1" longitude="-76.452995" name="Reserva Etnica Waorani" stateProvince="ORELLANA">Reserva Etnica Waorani</location>
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, 1 | km
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<collectorName id="B94EC592DE82C9208D03874DFC37E1F0">S. OnkoneGare Camp Transect Ent</collectorName>
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<quantity id="FA6AFC4EAFE3A0EC61DB59F9EAFB9523" metricMagnitude="2" metricUnit="m" metricValue="2.163" unit="m" value="216.3">
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<elevation id="8653BA63FDFE6F8519FC3A1575884A2A" metricMagnitude="2" metricUnit="m" metricValue="2.163" unit="m" value="216.3">216.3m</elevation>
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</quantity>
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| 15-Jan-94
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<geoCoordinate id="C248FA7E2538BA01DD6094135AB0D47A" degrees="00" direction="south" minutes="39" orientation="latitude" precision="1" seconds="25.7" value="-0.6571389">00°39'25.7"S</geoCoordinate>
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,
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<geoCoordinate id="08C2A85619B8E01845390F8D10BAD946" degrees="076" direction="west" minutes="27" orientation="longitude" precision="1" seconds="10.8" value="-76.452995">076°27'10.8"W</geoCoordinate>
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T.L | Erwin: et al Fogging Terre firme forest Lot 579", and a red "
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<typeStatus id="5CF78C2D05D6E9D17A274D8A93648FF0">HOLOTYPE</typeStatus>
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|
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<taxonomicName id="D6B63F3F16650FC14024EF5CF31858C4" authorityName="Andrade" authorityYear="1989" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="genus">Smilidarnis</taxonomicName>
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||
<taxonomicName id="3487BCBE791667B5AEEB32C7FC015AE3" authorityName="McKamey" authorityYear="2023" class="Insecta" family="Membracidae" genus="Smilidarnis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smilidarnis erwini" order="Hemiptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="erwini">Smilidarnis erwini</taxonomicName>
|
||
|
|
||
<collectorName id="82291A048BFA079FFB01C9A670A1A427">S.H. McKamey</collectorName>
|
||
". All legs are missing or badly damaged but the specimen, excluding its abdominal dissection, is otherwise intact.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5348A416D1C6DD441BD7A83ED08BA2CA" pageId="0" pageNumber="85" type="etymology">
|
||
<paragraph id="51EC52B453DAE1FB3F5A1223D8D4D1DA" pageId="0" pageNumber="85">Etymology.</paragraph>
|
||
<paragraph id="D9B5F6585680FDAEAAA4C239700CCEA4" pageId="0" pageNumber="85">The species is named to honor Terry Erwin for his innovative and revolutionary method of collecting insects through insecticidal fogging of the forest canopy, which has collected many new species, including this one.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC86A38EA05A83FD1DED461062BB423A" pageId="0" pageNumber="85" type="notes">
|
||
<paragraph id="3D9D42321FC5093BB8F79B2D7C680D6F" pageId="0" pageNumber="85">Notes.</paragraph>
|
||
<paragraph id="7334350D41EE64E1BBAA60F9B0817EEE" pageId="0" pageNumber="85">
|
||
The pronotal apex of
|
||
<taxonomicName id="6F4F564ECF65675AA75405EDD7C2CA27" lsidName="S. erwini" pageId="0" pageNumber="85" rank="species" species="erwini">
|
||
<emphasis id="4FD3F623C0DA0075DA84EEF8CDA36EEF" italics="true" pageId="0" pageNumber="85">S. erwini</emphasis>
|
||
</taxonomicName>
|
||
resembles that
|
||
<emphasis id="B76E7A1563D419D44603DBC89879F4E3" italics="true" pageId="0" pageNumber="85">
|
||
of
|
||
<taxonomicName id="CD4FF8A8375DBBB2A7144533801C926F" lsidName="S. fasciatus" pageId="0" pageNumber="85" rank="species" species="fasciatus">S. fasciatus</taxonomicName>
|
||
</emphasis>
|
||
in having the bases and tips of the posterolateral spines black. In
|
||
<taxonomicName id="D7BFAB1E1C24B641BFB4193D2A05C483" lsidName="S. fasciatus" pageId="0" pageNumber="85" rank="species" species="fasciatus">
|
||
<emphasis id="4394D572C62D77DED050BAA8DAF96843" italics="true" pageId="0" pageNumber="85">S. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
, however, in addition to other differences, those lateral spines are nearly as long as the middle spine, in comparison to
|
||
<taxonomicName id="540D1F27350AF4FADA7E42D3CC6DBED6" lsidName="S. erwini" pageId="0" pageNumber="85" rank="species" species="erwini">
|
||
<emphasis id="92859761D70F2897DCB5A36EA3530548" italics="true" pageId="0" pageNumber="85">S. erwini</emphasis>
|
||
</taxonomicName>
|
||
, in which they are about half the length of the middle spine, and
|
||
<taxonomicName id="0E054F0A040EBE39324A4230D7CE5DAC" lsidName="S. concolor" pageId="0" pageNumber="85" rank="species" species="concolor">
|
||
<emphasis id="CC9BF089C00A1D6C321C718CBB14C530" italics="true" pageId="0" pageNumber="85">S. concolor</emphasis>
|
||
</taxonomicName>
|
||
, in which they are even more reduced.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |