185 lines
22 KiB
XML
185 lines
22 KiB
XML
<document id="3BD9E7611325D051C0303CCB6FF8DFF3" ID="10.11646/zootaxa.4136.2.12" ID-DOI="10.11646/zootaxa.4136.2.12" ID-GBIF-Dataset="41bc911e-8bbf-4937-9d57-0000acd661bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="256508" ID-ZooBank="C1A1EC33-18E8-47F4-BDE4-2AA72F66113A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1467808610410" checkinUser="plazi" docAuthor="Nunes, Rafael V. & Vaz-De-Mello, Fernando Z." docDate="2016" docId="B56C87AEFFC6CD01FF79898EFEDEFCC7" docLanguage="en" docName="zootaxa.4136.2.12.pdf" docOrigin="Zootaxa 4136 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holocephalus pokornyi Nunes & Vaz-De-Mello, 2016, new species" docType="treatment" docVersion="7" lastPageNumber="399" masterDocId="4955FFD6FFC6CD03FFEE8C09FFB9FF86" masterDocTitle="A new species of Holocephalus Hope from Paraguay (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini s. str.)" masterLastPageNumber="400" masterPageNumber="397" pageNumber="397" updateTime="1698363811211" updateUser="plazi">
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<mods:mods id="D4A897BA1A54AD227EBDC68F28A48FF1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4077F2309F7613AE29DABAC36EE0FAC0">
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<mods:title id="E3CB3E4EE593B0DAC0C85D1C36BAA3C3">A new species of Holocephalus Hope from Paraguay (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini s. str.)</mods:title>
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</mods:titleInfo>
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<mods:name id="56B1F36D14E7EA47246142BA96527108" type="personal">
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<mods:role id="C30DD2DC2C947BDCA7BE9CCD3F25B266">
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<mods:roleTerm id="61E90A182B6C964B0265F538027A3A43">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2F155CDC3AC05287B4D265E4CE6F5A2B">Nunes, Rafael V.</mods:namePart>
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</mods:name>
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<mods:name id="D780DAEE752D0C757174DBF71436F79D" type="personal">
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<mods:role id="3C0E71E35AAD7D51857C4823338057A2">
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<mods:roleTerm id="9FE4BF4B22843CA08E0DB5B45A40B226">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BE5CE3EFE2CDA48AFFBC2220431E1819">Vaz-De-Mello, Fernando Z.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="922DBD1CF4A5F5092764973F6057B9DC">text</mods:typeOfResource>
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<mods:relatedItem id="8C67D52E1359F74F54B27A5C4BED2578" type="host">
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<mods:titleInfo id="575590C574005132D0073ADAC94DA6A1">
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<mods:title id="06759A577A6A2EA116848B1B3FDD8C8B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C1FB521EBF55E562985516D25913BC10">
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<mods:date id="91A2147CD344BD98DEA43A05580C9EAA">2016</mods:date>
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<mods:detail id="E7C27D64887A17D78FC0945ED3B8313C" type="volume">
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<mods:number id="F2E26B6195DDBFC3415C7489B820AFC2">4136</mods:number>
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</mods:detail>
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<mods:detail id="8941693913DD588C69F2FFC9D6581D48" type="issue">
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<mods:number id="B7185160AB9176A70BBAAA01D611B1CC">2</mods:number>
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</mods:detail>
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<mods:extent id="958272A14FAE28B2E4B80AD40F0009F1" unit="page">
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<mods:start id="154B39C4FE0B676008C95C9EAB68CE9F">397</mods:start>
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<mods:end id="D622680657C822C3CBC7FA68788E6CF3">400</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="F7E32AB4D566696ECB1C0218F4F16128">journal article</mods:classification>
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<mods:identifier id="10F6FC40FB47743BB75AD8837F758950" type="DOI">10.11646/zootaxa.4136.2.12</mods:identifier>
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<mods:identifier id="19FE13983A7E37A037B9AD1D05E6EF0A" type="GBIF-Dataset">41bc911e-8bbf-4937-9d57-0000acd661bd</mods:identifier>
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<mods:identifier id="2C4E4AA018D579F8E297436E34867987" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="65DED10BD2765420CF48388BF386039C" type="Zenodo-Dep">256508</mods:identifier>
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<mods:identifier id="22C6B501E6A8703DB6D9B336F2A0639F" type="ZooBank">C1A1EC33-18E8-47F4-BDE4-2AA72F66113A</mods:identifier>
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</mods:mods>
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<treatment id="B56C87AEFFC6CD01FF79898EFEDEFCC7" ID-DOI="http://doi.org/10.5281/zenodo.6085641" ID-GBIF-Taxon="121544413" ID-Zenodo-Dep="6085641" LSID="urn:lsid:plazi:treatment:B56C87AEFFC6CD01FF79898EFEDEFCC7" httpUri="http://treatment.plazi.org/id/B56C87AEFFC6CD01FF79898EFEDEFCC7" lastPageId="2" lastPageNumber="399" pageId="0" pageNumber="397">
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<subSubSection id="75DF6533FFC6CD03FF79898EFDF1FA24" box="[151,584,1415,1442]" pageId="0" pageNumber="397" type="nomenclature">
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<paragraph id="3D7A36B8FFC6CD03FF79898EFDF1FA24" blockId="0.[151,1436,1415,2028]" box="[151,584,1415,1442]" pageId="0" pageNumber="397">
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<heading id="663281D4FFC6CD03FF79898EFDF1FA24" bold="true" box="[151,584,1415,1442]" fontSize="11" level="1" pageId="0" pageNumber="397" reason="1">
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<emphasis id="0FB1EAAAFFC6CD03FF79898EFDF1FA24" bold="true" box="[151,584,1415,1442]" pageId="0" pageNumber="397">
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<taxonomicName id="FAC54D3BFFC6CD03FF79898EFE08FA27" ID-CoL="3M8M8" box="[151,433,1415,1441]" class="Insecta" family="Scarabaeidae" genus="Holocephalus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="397" phylum="Arthropoda" rank="species" species="pokornyi" status="sp. nov.">
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<emphasis id="0FB1EAAAFFC6CD03FF79898EFE08FA27" bold="true" box="[151,433,1415,1441]" italics="true" pageId="0" pageNumber="397">Holocephalus pokornyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="148257D1FFC6CD03FE578981FDF1FA24" box="[441,584,1416,1442]" pageId="0" pageNumber="397" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="75DF6533FFC6CD03FF7989A2FA9CFA44" box="[151,1317,1451,1474]" pageId="0" pageNumber="397" type="materials_examined">
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<paragraph id="3D7A36B8FFC6CD03FF7989A2FA9CFA44" blockId="0.[151,1436,1415,2028]" box="[151,1317,1451,1474]" pageId="0" pageNumber="397">
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<emphasis id="0FB1EAAAFFC6CD03FF7989A2FEC1FA44" bold="true" box="[151,376,1451,1474]" pageId="0" pageNumber="397">Examined material.</emphasis>
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<typeStatus id="E27E881AFFC6CD03FE9089A2FE58FA44" box="[382,481,1451,1474]" pageId="0" pageNumber="397" type="holotype">Holotype</typeStatus>
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Ƌ.
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<collectingCountry id="45D27628FFC6CD03FDE789A2FD20FA44" box="[521,665,1451,1474]" name="Paraguay" pageId="0" pageNumber="397">PARAGUAY</collectingCountry>
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.
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<emphasis id="0FB1EAAAFFC6CD03FD4D89A2FD55FA44" bold="true" box="[675,748,1451,1474]" pageId="0" pageNumber="397">Chaco</emphasis>
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, Cerro Corá NP.
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<date id="497B1078FFC6CD03FC4D89A2FB8DFA44" box="[931,1076,1451,1474]" pageId="0" pageNumber="397" value="2001-01-19" valueMax="2001-01-21" valueMin="2001-01-19">19-21-I-2001</date>
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, Lgt Mráček [
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<collectionCode id="5BD4AE7DFFC6CD03FB2689A2FAA3FA44" box="[1224,1306,1451,1474]" pageId="0" pageNumber="397">CEMT</collectionCode>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="75DF6533FFC6CD03FF2989C4FA25F981" pageId="0" pageNumber="397" type="diagnosis">
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<paragraph id="3D7A36B8FFC6CD03FF2989C4FA25F981" blockId="0.[151,1436,1415,2028]" pageId="0" pageNumber="397">
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<emphasis id="0FB1EAAAFFC6CD03FF2989C4FE85FA62" bold="true" box="[199,316,1485,1508]" pageId="0" pageNumber="397">Diagnosis.</emphasis>
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The tridentate cephalic carina (
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<figureCitation id="A5FE2A3DFFC6CD03FD6789C4FD7DFA62" box="[649,708,1485,1508]" captionStart="FIGURE 1" captionStartId="1.[151,244,1961,1983]" captionTargetBox="[164,1425,212,1920]" captionTargetId="figure@1.[151,1435,191,1939]" captionTargetPageId="1" captionText="FIGURE 1. Holocephalus pokornyi n. sp. (Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera." httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="0" pageNumber="397">Fig. 1</figureCitation>
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C), anterior pronotal angles produced into acute carina (
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<figureCitation id="A5FE2A3DFFC6CD03FAF989C4FAE2FA62" box="[1303,1371,1485,1508]" captionStart="FIGURE 1" captionStartId="1.[151,244,1961,1983]" captionTargetBox="[164,1425,212,1920]" captionTargetId="figure@1.[151,1435,191,1939]" captionTargetPageId="1" captionText="FIGURE 1. Holocephalus pokornyi n. sp. (Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera." httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="0" pageNumber="397">Figs. 1</figureCitation>
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A–C) and ocellate coarse puncture extending beyond outer edge of eyes will separate this species from all other
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<emphasis id="0FB1EAAAFFC6CD03FAEF89F9FA25F981" box="[1281,1436,1520,1543]" italics="true" pageId="0" pageNumber="397">
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<taxonomicName id="FAC54D3BFFC6CD03FAEF89F9FA21F981" box="[1281,1432,1520,1543]" class="Insecta" family="Scarabaeidae" genus="Holocephalus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="397" phylum="Arthropoda" rank="genus">Holocephalus</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="75DF6533FFC6CD01FF298A1AFB38FDC9" lastPageId="2" lastPageNumber="399" pageId="0" pageNumber="397" type="description">
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<paragraph id="3D7A36B8FFC6CD03FF298A1AFD67F86A" blockId="0.[151,1436,1415,2028]" pageId="0" pageNumber="397">
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<emphasis id="0FB1EAAAFFC6CD03FF298A1AFEEBF9AC" bold="true" box="[199,338,1555,1578]" pageId="0" pageNumber="397">Description.</emphasis>
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<typeStatus id="E27E881AFFC6CD03FEB58A1AFE7BF9AC" box="[347,450,1555,1578]" pageId="0" pageNumber="397" type="holotype">Holotype</typeStatus>
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: Male, body length
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<quantity id="FA3D9B5DFFC6CD03FD4F8A1AFD48F9AC" box="[673,753,1555,1578]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="0" pageNumber="397" unit="mm" value="21.0">21 mm</quantity>
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, pronotum width:
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<quantity id="FA3D9B5DFFC6CD03FC508A1AFBB7F9AC" box="[958,1038,1555,1578]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="0" pageNumber="397" unit="mm" value="11.0">11 mm</quantity>
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. Color: black, glossy.
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<emphasis id="0FB1EAAAFFC6CD03FAEB8A1AFAF2F9AC" bold="true" box="[1285,1355,1555,1578]" pageId="0" pageNumber="397">Head:</emphasis>
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Dorsal surface strongly, transversely wrinkled all over its extension except posterior to cephalic horn, at interocular space. Outer margin of clypeous slightly thickened apically. Clypeo-genal suture distinct, straight. Cephalic carina tridentate, teeth of equal size (
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<figureCitation id="A5FE2A3DFFC6CD03FEFC8A72FEE8F914" box="[274,337,1659,1682]" captionStart="FIGURE 1" captionStartId="1.[151,244,1961,1983]" captionTargetBox="[164,1425,212,1920]" captionTargetId="figure@1.[151,1435,191,1939]" captionTargetPageId="1" captionText="FIGURE 1. Holocephalus pokornyi n. sp. (Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera." httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="0" pageNumber="397">Fig. 1</figureCitation>
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C). Eye 1.5 times longer than wide, interocular width equals 7 times the transverse eyes diameter (in dorsal view). Labrum with dense setae; setae long, reddish brown. First segment of labial palpus greatly enlarged, concealing the second segment in ventral view (
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<figureCitation id="A5FE2A3DFFC6CD03FD4A8AC9FD5AF951" box="[676,739,1728,1751]" captionStart="FIGURE 1" captionStartId="1.[151,244,1961,1983]" captionTargetBox="[164,1425,212,1920]" captionTargetId="figure@1.[151,1435,191,1939]" captionTargetPageId="1" captionText="FIGURE 1. Holocephalus pokornyi n. sp. (Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera." httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="0" pageNumber="397">Fig. 1</figureCitation>
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B); first and second segments of labial palpus with dense setae; setae long, reddish-brown. Second and third segments of labial palpus not enlarged; third segment lacking setae. Antenna 9-segmented, first segmented elongated, sub-equal to segments 2–6. Antennal club (segments 7–9) spherical.
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<emphasis id="0FB1EAAAFFC6CD03FF798B21FEAAF8B9" bold="true" box="[151,275,1832,1855]" pageId="0" pageNumber="397">Pronotum:</emphasis>
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disc lacking punctures except for impressed patch of dense, ocellate punctures medioapically; row of punctures along inside of basal/posterior margin. Pronotal anterior declivity with coarse ocellate puncture extending beyond outer edge of eyes. Anterior angles acute. Pronotal margin at anterior angle carinating, producing a spiniform anterior angle (
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<figureCitation id="A5FE2A3DFFC6CD03FEAF8B99FE32F821" box="[321,395,1936,1959]" captionStart="FIGURE 1" captionStartId="1.[151,244,1961,1983]" captionTargetBox="[164,1425,212,1920]" captionTargetId="figure@1.[151,1435,191,1939]" captionTargetPageId="1" captionText="FIGURE 1. Holocephalus pokornyi n. sp. (Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera." httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="0" pageNumber="397">Figs. 1</figureCitation>
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A–C). Disc with transverse swelling on anterior fourth.
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<emphasis id="0FB1EAAAFFC6CD03FBE88B99FB2EF821" bold="true" box="[1030,1175,1936,1959]" pageId="0" pageNumber="397">Hypomeron:</emphasis>
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bearing ocellate setose punctures equally spaced.
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<emphasis id="0FB1EAAAFFC6CD03FE3D8BBAFDDDF84C" bold="true" box="[467,612,1971,1994]" pageId="0" pageNumber="397">Prosternum:</emphasis>
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glabrous, with micro-sculpture (chagrination, visible with
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<date id="497B1078FFC6CD03FA9A8BBAFA22F84C" box="[1396,1435,1971,1994]" pageId="0" pageNumber="397" value="1930-10">30x</date>
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magnification).
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<emphasis id="0FB1EAAAFFC6CD03FEAF8BDCFE2AF86A" bold="true" box="[321,403,2005,2028]" pageId="0" pageNumber="397">Elytra:</emphasis>
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interstriae glabrous, strongly
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</paragraph>
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<caption id="69BA6630FFC7CD02FF798BA0FBE0F85D" httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="1" pageNumber="398" targetBox="[164,1425,212,1920]" targetPageId="1">
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<paragraph id="3D7A36B8FFC7CD02FF798BA0FBE0F85D" blockId="1.[151,1435,1961,2011]" pageId="1" pageNumber="398">
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<emphasis id="0FB1EAAAFFC7CD02FF798BA0FEB7F839" bold="true" box="[151,270,1961,1983]" pageId="1" pageNumber="398">FIGURE 1.</emphasis>
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<taxonomicName id="FAC54D3BFFC7CD02FEFA8BA0FE4FF838" box="[276,502,1961,1982]" class="Insecta" family="Scarabaeidae" genus="Holocephalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="pokornyi" status="sp. nov.">
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<emphasis id="0FB1EAAAFFC7CD02FEFA8BA0FE4FF838" box="[276,502,1961,1982]" italics="true" pageId="1" pageNumber="398">Holocephalus pokornyi</emphasis>
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</taxonomicName>
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<emphasis id="0FB1EAAAFFC7CD02FE138BA0FD8CF839" bold="true" box="[509,565,1961,1983]" pageId="1" pageNumber="398">
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<taxonomicNameLabel id="148257D1FFC7CD02FE138BA0FD8CF839" box="[509,565,1961,1983]" pageId="1" pageNumber="398" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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(Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera.
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</paragraph>
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</caption>
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<paragraph id="3D7A36B8FFC4CD01FF798C91FB38FDC9" blockId="2.[151,1436,152,833]" pageId="2" pageNumber="399">
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convex and lacking punctures. Striae strongly impressed having ocellate punctures spaced by their diameter; striae 1–6 extending to basis, striae 7–8 almost reaching the basis; striae 9 interrupted since the apical sixth.
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<emphasis id="0FB1EAAAFFC4CD01FB198CB3FA25FF57" bold="true" box="[1271,1436,186,209]" pageId="2" pageNumber="399">Mesosternum:</emphasis>
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laterally, with ocellate setose punctures equally spaced; setae scattered, only visible in
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<date id="497B1078FFC4CD01FBC38CD4FBECFF72" box="[1069,1109,221,244]" pageId="2" pageNumber="399" value="1930-10">30x</date>
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magnification. Medial portion of mesosternum glabrous and lacking punctures.
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<emphasis id="0FB1EAAAFFC4CD01FD598D09FCE3FE91" bold="true" box="[695,858,256,279]" pageId="2" pageNumber="399">Metasternum:</emphasis>
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anterior lobe bearing ocellate setose punctures only close the margins. Lateral portions of metasternum with ocellate setose puncutres equally spaced.
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<emphasis id="0FB1EAAAFFC4CD01FB0D8D2BFA25FEBF" bold="true" box="[1251,1436,290,313]" pageId="2" pageNumber="399">Mesepisternum:</emphasis>
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smooth, with few sparse ocellate punctures. Upper margin straight.
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<emphasis id="0FB1EAAAFFC4CD01FC958D4CFC00FEDA" bold="true" box="[891,953,325,348]" pageId="2" pageNumber="399">Legs:</emphasis>
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Protibiae bearing four teeth. Protibiae inner and outer surface with thin, longitudinal line of dense, yelllowish-brown setae. Ventral surface of femura with few yellowish-brown setae on apical side, other portions of structure smooth and glabrous. All calcar spiniform, truncated. Meso and metatibiae with sinuous apex, apical angles producing external teeth.
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<emphasis id="0FB1EAAAFFC4CD01FC098DA4FBE6FE42" bold="true" box="[999,1119,429,452]" pageId="2" pageNumber="399">Abdomen:</emphasis>
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sternites bearing few ocellate setose punctures concentrated on the sides; setae short and scattered. Ocellate punctures present along the anterior margin of each sternite. Sternites medially glabrous and lacking punctures. Sixth sternite strongly constricted medially.
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<emphasis id="0FB1EAAAFFC4CD01FF798E1CFEBDFDAA" bold="true" box="[151,260,533,556]" pageId="2" pageNumber="399">Pygidium</emphasis>
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: width 2 times length medially. Surface glabrous; margin complete, slightly convex surface in lateral view.
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<emphasis id="0FB1EAAAFFC4CD01FF798E31FEA2FDC9" bold="true" box="[151,283,568,591]" pageId="2" pageNumber="399">Aedeagous:</emphasis>
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as in Figs (1E–F). Paramera symetrical. On lateral view apex truncated (
|
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<figureCitation id="A5FE2A3DFFC4CD01FBCA8E31FBD9FDC9" box="[1060,1120,568,591]" captionStart="FIGURE 1" captionStartId="1.[151,244,1961,1983]" captionTargetBox="[164,1425,212,1920]" captionTargetId="figure@1.[151,1435,191,1939]" captionTargetPageId="1" captionText="FIGURE 1. Holocephalus pokornyi n. sp. (Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera." httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75DF6533FFC4CD01FF298E53FB95FDF7" box="[199,1068,602,625]" pageId="2" pageNumber="399" type="distribution">
|
||
<paragraph id="3D7A36B8FFC4CD01FF298E53FB95FDF7" blockId="2.[151,1436,152,833]" box="[199,1068,602,625]" pageId="2" pageNumber="399">
|
||
<emphasis id="0FB1EAAAFFC4CD01FF298E53FEE0FDF7" bold="true" box="[199,345,602,625]" pageId="2" pageNumber="399">Distribution.</emphasis>
|
||
Until now, only known from Cerro Corá, Chaco,
|
||
<collectingCountry id="45D27628FFC4CD01FC828E53FC76FDF7" box="[876,975,602,625]" name="Paraguay" pageId="2" pageNumber="399">Paraguay</collectingCountry>
|
||
(
|
||
<figureCitation id="A5FE2A3DFFC4CD01FC338E53FBA6FDF7" box="[989,1055,602,625]" captionStart="FIGURE 2" captionStartId="2.[151,244,1593,1615]" captionTargetBox="[339,1247,887,1571]" captionTargetId="figure@2.[339,1248,887,1571]" captionTargetPageId="2" captionText="FIGURE 2. Literature (grey) and new (black) geographical records for Holocephalus species. Question marks (?) refer to localities that are doubtful in our opinion." httpUri="https://zenodo.org/record/256510/files/figure.png" pageId="2" pageNumber="399">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75DF6533FFC4CD01FF298E74FDF3FD31" pageId="2" pageNumber="399" type="etymology">
|
||
<paragraph id="3D7A36B8FFC4CD01FF298E74FDF3FD31" blockId="2.[151,1436,152,833]" pageId="2" pageNumber="399">
|
||
<emphasis id="0FB1EAAAFFC4CD01FF298E74FEFEFD12" bold="true" box="[199,327,637,660]" pageId="2" pageNumber="399">Etymology.</emphasis>
|
||
Named after Svatopluk Pokorný, Czech scarabaeoidologist who kindly provided us with that and several other very interesting specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="75DF6533FFC4CD01FF298ECBFEDEFCC7" pageId="2" pageNumber="399" type="discussion">
|
||
<paragraph id="3D7A36B8FFC4CD01FF298ECBFEDEFCC7" blockId="2.[151,1436,152,833]" pageId="2" pageNumber="399">
|
||
<emphasis id="0FB1EAAAFFC4CD01FF298ECBFE8CFD5F" bold="true" box="[199,309,706,729]" pageId="2" pageNumber="399">Remarks.</emphasis>
|
||
<taxonomicName id="FAC54D3BFFC4CD01FED28ECBFE04FD5F" box="[316,445,706,729]" class="Insecta" family="Scarabaeidae" genus="Holocephalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="pokornyi">
|
||
<emphasis id="0FB1EAAAFFC4CD01FED28ECBFE04FD5F" box="[316,445,706,729]" italics="true" pageId="2" pageNumber="399">H. pokornyi</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="E27E881AFFC4CD01FE2A8ECBFD98FD5F" box="[452,545,706,729]" pageId="2" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
came to us with broken protarsi, some broken teeth of protibiae and a longitudinal crack on pronotum (can be seen in
|
||
<figureCitation id="A5FE2A3DFFC4CD01FDF48EECFDEEFD7A" box="[538,599,741,764]" captionStart="FIGURE 1" captionStartId="1.[151,244,1961,1983]" captionTargetBox="[164,1425,212,1920]" captionTargetId="figure@1.[151,1435,191,1939]" captionTargetPageId="1" captionText="FIGURE 1. Holocephalus pokornyi n. sp. (Holotype) (A) dorsal habitus (B) ventral habitus (C) frontal view of head and pronotum (D) lateral habitus (E) lateral habitus of aedeagus (F) frontal view of aedeagus evidencing paremera." httpUri="https://zenodo.org/record/256509/files/figure.png" pageId="2" pageNumber="399">Fig. 1</figureCitation>
|
||
A). Until now this species is only known by the
|
||
<typeStatus id="E27E881AFFC4CD01FB9F8EECFB6BFD7A" box="[1137,1234,741,764]" pageId="2" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
. The species lacks natural history and ecological data. However, at least one
|
||
<taxonomicName id="FAC54D3BFFC4CD01FD108F01FC2BFC99" box="[766,914,776,799]" class="Insecta" family="Scarabaeidae" genus="Holocephalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0FB1EAAAFFC4CD01FD108F01FC2BFC99" box="[766,914,776,799]" italics="true" pageId="2" pageNumber="399">Holocephalus</emphasis>
|
||
</taxonomicName>
|
||
species is myrmecophilous according to Vaz-de- Mello
|
||
<emphasis id="0FB1EAAAFFC4CD01FF338F23FEAAFCC7" box="[221,275,810,833]" italics="true" pageId="2" pageNumber="399">et al.</emphasis>
|
||
(1998).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |