treatments-xml/data/B5/10/B8/B510B805C1405B1081A1B04ACA258316.xml

<document ID-CLB-Dataset="6375" ID-DOI="http://dx.doi.org/10.3897/phytokeys.178.64564" ID-GBIF-Dataset="dc4dfb29-9f52-4d81-b923-358afd0bb8b9" ID-PMC="PMC8184734" ID-Pensoft-Pub="1314-2003-178-147" ID-Pensoft-UUID="1E320638632250E9B9D5FFC20CC2695B" ID-PubMed="34140828" ModsDocID="1314-2003-178-147" checkinTime="1622492441769" checkinUser="pensoft" docAuthor="Santamaria-Aguilar, Daniel &amp; Lagomarsino, Laura P." docDate="2021" docId="B510B805C1405B1081A1B04ACA258316" docLanguage="en" docName="PhytoKeys 178: 147-170" docOrigin="PhytoKeys 178" docPubDate="2021-05-31" docSource="http://dx.doi.org/10.3897/phytokeys.178.64564" docTitle="Otoba scottmorii D. Santam. 2021, sp. nov." docType="treatment" docVersion="6" id="1E320638632250E9B9D5FFC20CC2695B" lastPageNumber="147" masterDocId="1E320638632250E9B9D5FFC20CC2695B" masterDocTitle="Two new species of Otoba (Myristicaceae) from Colombia" masterLastPageNumber="170" masterPageNumber="147" pageNumber="147" updateTime="1732840509785" updateUser="ExternalLinkService">
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<mods:title id="D85843BE6DDBD7B2884113F21B9434B9">Two new species of Otoba (Myristicaceae) from Colombia</mods:title>
</mods:titleInfo>
<mods:name id="CDD5E4E1C55643957FF76264531E7C1A" type="personal">
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<mods:namePart id="CF9689DCCB68D662792E773B80ADA901">Santamaria-Aguilar, Daniel</mods:namePart>
<mods:nameIdentifier id="37BC2151295DADD596EE747E635D7D02" type="ORCID">https://orcid.org/0000-0002-6339-8250</mods:nameIdentifier>
<mods:affiliation id="F7760B2B248D43AC745859D14164F1D6">Shirley C. Tucker Herbarium, Department of Biological Sciences, Louisiana State, University, Baton Rouge, LA 70803 - 1705, USA &amp; Missouri Botanical Garden St. Louis, MO 63166, USA</mods:affiliation>
<mods:nameIdentifier id="0F4BA5D9D55F211664673A0D4D5517CE" type="email">daniel.santamaria366@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CC9E3C3E971450DA765E38EC46DD8F3C">Lagomarsino, Laura P.</mods:namePart>
<mods:affiliation id="C78AFFE713CBB10B9047A322F94BE526">Shirley C. Tucker Herbarium, Department of Biological Sciences, Louisiana State, University, Baton Rouge, LA 70803 - 1705, USA</mods:affiliation>
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<mods:date id="C32B4B5682CF8153E21FC351BFDE14C6">2021</mods:date>
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<mods:number id="723807B7AD3E4B09E3853A6FF487DEB0">178</mods:number>
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<mods:identifier id="8B3B715FD234FC9CF0ECCC47BA1A14D3" type="DOI">http://dx.doi.org/10.3897/phytokeys.178.64564</mods:identifier>
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<treatment id="B510B805C1405B1081A1B04ACA258316" ID-GBIF-Taxon="182473848" LSID="urn:lsid:plazi:treatment:B510B805C1405B1081A1B04ACA258316" httpUri="http://treatment.plazi.org/id/B510B805C1405B1081A1B04ACA258316" lastPageNumber="147" pageId="0" pageNumber="147" scope_family="Myristicaceae">
<subSubSection id="44D8910D3393B7B54312C9B38AE6AD37" pageId="0" pageNumber="147" type="nomenclature">
<paragraph id="A7D2BCFAF9812347CFF5AEA1372D2B4D" pageId="0" pageNumber="147">
<taxonomicName id="045302EA7906E9B075E194CA3A5358C4" ID-CoL="8MNRF" LSID="B510B805-C140-5B10-81A1-B04ACA258316" authority="D. Santam." authorityName="D. Santam." authorityYear="2021" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii" status="sp. nov.">Otoba scottmorii D. Santam.</taxonomicName>
<taxonomicNameLabel id="1FF18DC204526514EB96B7B6B2397E44" pageId="0" pageNumber="147">sp. nov.</taxonomicNameLabel>
<figureCitation id="1412A5D87D1C871C83E1026391B74445" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Otoba scottmorii A staminate flowering branch, with detail of leaf surface (detail below left) B leaf base, petiole and basal portion of inflorescence C detail of staminate inflorescence D flower bud E staminate flowers, entire (above right) and with one perianth lobe removed, showing the androecium (below center) and perianth lobe (above left) F androecium G stem with infructescence, including an immature, opening fruit H view of dehisced fruit (left, below G) showing the seed and aril (right) Otoba cyclobasis I staminate flowering branch showing leaves with brochidodromous venation, including detail of marginal vein J staminate flower with one perianth lobe removed, showing the androecium and perianth lobe (above left) K androecium. Illustration by Bobbi Angell, W. W. Thomas &amp; C. J. Castano 5501 (NY) (A-F), J. Brand &amp; M. Narvaez 682 (MO) (G, H), E. Narvaez et al. 1072 (MO- 2468293) (I-K)." figureDoi="10.3897/phytokeys.178.64564.figure3" httpUri="https://binary.pensoft.net/fig/550342" pageId="0" pageNumber="147">Figs 3</figureCitation>
<figureCitation id="EBDCAAC1F11CEE6FEC38D69246A57770" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">, 5B</figureCitation>
<figureCitation id="47B7F8323664750872B7021AA3CA699E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparations of herbarium specimens of Otoba acuminata (A L. Poveda et al. 3793, MO) O. cyclobasis (B P. Mendez et al. 156, MO) O. scottmorii (C W. W. Thomas &amp; C. J. Castano 5501, NY), and O. vespertilio (D R. Aguilar et al. 5024, MO)." figureDoi="10.3897/phytokeys.178.64564.figure6" httpUri="https://binary.pensoft.net/fig/550345" pageId="0" pageNumber="147">, 6C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F0FE04B3D4ABF3193CACF631C5A2BEE7" pageId="0" pageNumber="147" type="type">
<paragraph id="1C626A140B3E8449C1535E5E026A299A" pageId="0" pageNumber="147">Type.</paragraph>
<paragraph id="050D797C2BA64A60157ADBAE6B22B153" pageId="0" pageNumber="147">
Colombia. Antioquia: [Municipio] Mpio. Segovia, 24.5 km N of Remedios (17 km N of La Cruzada) on road to Zaragoza, hills side forest above 
<normalizedToken id="ED0AC651403B029166F79F195CE62CE6" originalValue="río">rio</normalizedToken>
<normalizedToken id="9C477E6DB6ED8B2DFCB884E3C2BC5F3A" originalValue="Poconé">Pocone</normalizedToken>
, 
<geoCoordinate id="3D302C0AC27071C2F768920BEE3231FE" degrees="07" direction="north" minutes="12" orientation="latitude" precision="925" value="7.2">07°12'N</geoCoordinate>
, 
<geoCoordinate id="2F508718C8BA1FE5AE475803A771EEA4" degrees="074" direction="west" minutes="48" orientation="longitude" precision="925" value="-74.8">074°48'W</geoCoordinate>
, [not elev.], 20 Jul. 1987 (♂ fl), W.W. Thomas &amp; C.J. 
<normalizedToken id="B07A59166BCF437939C9314BD8B2C158" originalValue="Castaño">Castano</normalizedToken>
5501 (holotype: NY! [04206141]; isotypes: INPA-159329 [digital image!], UPCB-35245 [n.v.]).
</paragraph>
</subSubSection>
<subSubSection id="8804E76C38CA9E29E092C5A8A0C99701" pageId="0" pageNumber="147" type="diagnosis">
<paragraph id="1B63D0F6C68E40F8036F51E267B6A12B" pageId="0" pageNumber="147">Diagnosis.</paragraph>
<paragraph id="F5A5EF24B311F6B828BEB3C39DEE1F16" pageId="0" pageNumber="147">
<taxonomicName id="171B3CD28A9C0FF48775D6C3469E476F" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="F05F3F92874D272E5E723F3C65AD06AC" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="70172737749EDFE00786D8B105984A3F" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. cyclobasis" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="cyclobasis">
<emphasis id="99B63F6B2466E9EB7F03B22D8B5DBC2D" italics="true" pageId="0" pageNumber="147">O. cyclobasis</emphasis>
</taxonomicName>
from Ecuador. However, it differs in having smaller leaves (5.5-8.5 [-11.2] vs. 9-14 [-18] cm long) with an attenuate to acuminate apex (vs. cuspidate), inconspicuous marginal and secondary veins (vs. conspicuous), fewer lateral veins (4-7 vs. 14-17), straight axes of staminate inflorescences (vs. zig zag), a longer filament column ([1.3-] 1.5-1.6 vs. 1 mm long), larger fruits (1.8-2.1 vs. 1.5-2.5 mm long) with thinner pericarp (0.68-0.89 vs. 1-2 mm thick), and seeds that are gibbose at the apex (vs. near-basal and lateral gibba).
</paragraph>
</subSubSection>
<subSubSection id="E48ECE8D821ECC5626E411E27DCB41D8" pageId="0" pageNumber="147" type="description">
<paragraph id="8FC97B71A3EE90AC1A6027E4F8E240FD" pageId="0" pageNumber="147">Description.</paragraph>
<paragraph id="C1128EAA35B0BC92DD46DAF690BFC1C5" pageId="0" pageNumber="147">
<emphasis id="69F6FD4EFEA50F46B924AF53D5F250E9" bold="true" italics="true" pageId="0" pageNumber="147">Tree</emphasis>
(2-) 15-20 m tall 
<normalizedToken id="C7FC6EEC89CDC47FADACC9B159A08418" originalValue="×">x</normalizedToken>
45 cm diam., external and internal bark not described. 
<emphasis id="F241ECF7EB5EAB22736261C90A8FCFFC" bold="true" italics="true" pageId="0" pageNumber="147">Exudate</emphasis>
once described as red in the bark, transparent and without specifying from where, or without exudate. 
<emphasis id="7B7B3B71AE41108C5774850A2778252D" bold="true" italics="true" pageId="0" pageNumber="147">Twigs</emphasis>
0.88-0.1 cm thick, terete to slightly flattened laterally, the external bark brown to grayish, with malpighiaceous trichomes that are 0.2-0.3 mm long, brown to ferruginous or sometimes glabrescent. 
<emphasis id="300F4EF5578475BBAF1B5D85F6CCE69E" bold="true" italics="true" pageId="0" pageNumber="147">Young foliar bud</emphasis>
0.6-1.2 cm long, densely pubescent. 
<emphasis id="21EA071F5A352627EBD794A5E735C4D2" bold="true" italics="true" pageId="0" pageNumber="147">Leaves</emphasis>
: petiole 0.5-1.1 
<normalizedToken id="861157AEAB4FD0AF1B65056E34195AE1" originalValue="×">x</normalizedToken>
0.055-0.1 cm, canaliculate, not winged; lamina 5.5-8.5 (-11.2) 
<normalizedToken id="1F54DE7B9E67906CF13A410D653821A5" originalValue="×">x</normalizedToken>
1.7-2.5 (-3.7) cm, elliptic; adaxial side glabrous, drying blackish, sometimes brown in young leaves, the surface muricate; abaxial side drying brown to whitish grayish, the surface wrinkled to muricate, sparsely pubescent to glabrescent, with malpighiaceous trichomes that are 0.2-0.4 mm long, sessile, and ferruginous, and squamate trichomes that are ca. 0.08 mm diam. with the central portion dark and sides lighter (sometimes these sides appearing absent), with whitish crystals; vernation line absent; midvein flat to very slightly ribbed on adaxial side, the same color as the surface, abaxially 0.2-0.4 mm wide, slightly raised, darker than the surface; secondary veins brochidodromous, the loops 0.1-0.3 cm from the margin (sometimes the loops not visible), lateral veins 4-7 per side, 3-5 veins per 3 cm, on adaxial side flat and visible, on abaxial side flat and not very conspicuous, arcuate distally, the marginal vein not visible on adaxial side, slightly visible on abaxial side; tertiary veins indistinct; base attenuate, not revolute; margin entire, not revolute; apex attenuate to acuminate, the acumen 0.6-1 (-2) cm long. 
<emphasis id="64876A6FB3D0C94C9BA509ADD012F49F" bold="true" italics="true" pageId="0" pageNumber="147">Staminate inflorescence</emphasis>
: axillary (only very young twigs) or supraaxillary, with 1-2 (-3) main axes, spiciform, these axes 1.5-4.3 cm 
<normalizedToken id="9A7737C5D5545BAB922F8812034CF3F6" originalValue="×">x</normalizedToken>
0.41-0.1 mm, pubescent, the trichomes ferruginous to coppery, each axis compound with 2-3 (-6) fascicles of flowers, each fascicle with 1-5 (-8) flowers, alternate; bracts not seen; pedicel 3-5 mm long, pubescent; bracteoles absent. 
<emphasis id="0220FA1F5D9329EA4721DA2957A82CCB" bold="true" italics="true" pageId="0" pageNumber="147">Staminate flowers</emphasis>
: flower bud 1-2.5 
<normalizedToken id="DCA9EBF7A18A0A5D7A979FFF25AF5F63" originalValue="×">x</normalizedToken>
0.6-0.1 mm, elliptic to lanceolate; perianth 2-2.5 mm long, yellow or orange (in fresh material), sub-membranaceous, connate by 0.5-0.7 (-1) mm; lobes 3 (4), 1.5-1.8 
<normalizedToken id="7F1E333A4788908E535FFC960B199AB4" originalValue="×">x</normalizedToken>
0.8-1.1 mm, without resinous punctuations or lines, pubescent outside, the trichomes ferruginous to coppery, inside glabrous, the apex acute to obtuse, without inflexed-apiculo, the edges flat or slightly turned inwards distally, without a swollen lobed ring; filament column (1.3-) 1.5-1.6 mm long, bowling pin-shaped, fleshy, connate, glabrous; anthers 3, 0.2-0.4 mm long, free, lanceolate to oblong, apex slightly incurved. 
<emphasis id="DD02E952009A82AF5A0AAEFE84568C61" bold="true" italics="true" pageId="0" pageNumber="147">Pistillate inflorescence</emphasis>
and 
<emphasis id="CE02B48F25BD956077299AFF41CAFEB9" bold="true" italics="true" pageId="0" pageNumber="147">flowers</emphasis>
: not seen. 
<emphasis id="911ED6955A34116DD8A3181DE340F234" bold="true" italics="true" pageId="0" pageNumber="147">Infructescence</emphasis>
2-4 cm long, with a solitary fruit; pedicel 1.2-1.6 cm long. 
<emphasis id="513A978201FC0EFC9DA6F031EBBC76AE" bold="true" italics="true" pageId="0" pageNumber="147">Fruits</emphasis>
1.8-2.1 
<normalizedToken id="D97EEFE0B2089E511A00CA90DC4692D0" originalValue="×">x</normalizedToken>
1.5-1.7 cm, green when fresh, globose, surface glabrous, colliculate-rugose, sometimes with whitish lenticels (J. Brand 682), the line of dehiscence smooth, the base obtuse, apex apiculate, the acumen 0.1-0.3 cm long; pericarp 0.68-0.89 mm thick; seed ca. 1.6 
<normalizedToken id="A334CE6B0B6416FB9D22EB4BACBE2D4F" originalValue="×">x</normalizedToken>
1.5 cm, similar in shape to the fruit, brownish (in dry material), gibbose at the apex (ca. 0.3 mm wide), the testa ca. 0.4 mm thick; aril described once as red (E. Renteria 4680), brownish to yellowish translucent in dry material, dry and membranous in texture.
</paragraph>
<caption id="1CB3F91C11ABF216A2727958CD3CF169" doi="10.3897/phytokeys.178.64564.figure3" httpUri="https://binary.pensoft.net/fig/550342" pageId="0" pageNumber="147" start="Figure 3" startId="F3">
<paragraph id="E404334ED102A369267F1E4BA55618A0" pageId="0" pageNumber="147">
<emphasis id="06FDD4992410D43BE014F5A650790DFB" bold="true" pageId="0" pageNumber="147">Figure 3.</emphasis>
<taxonomicName id="2EA720CFB2826D881FC01B22F43589B1" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="B70C50A834955C5F86210BE003FE8E4A" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
<emphasis id="259AC219F82BB1CE432B15DAB4D896C6" bold="true" pageId="0" pageNumber="147">A</emphasis>
staminate flowering branch, with detail of leaf surface (detail below left) 
<emphasis id="93061FBFC369915EF300BF4FF3BEBD3C" bold="true" pageId="0" pageNumber="147">B</emphasis>
leaf base, petiole and basal portion of inflorescence 
<emphasis id="8994700906BE8D0E2B9E7EE421AFFF53" bold="true" pageId="0" pageNumber="147">C</emphasis>
detail of staminate inflorescence 
<emphasis id="A63CE71AC769B2E1AFD42D9AD3B53DCC" bold="true" pageId="0" pageNumber="147">D</emphasis>
flower bud 
<emphasis id="B0ABD1397F9125EAD058E515915035F3" bold="true" pageId="0" pageNumber="147">E</emphasis>
staminate flowers, entire (above right) and with one perianth lobe removed, showing the androecium (below center) and perianth lobe (above left) 
<emphasis id="9872E435EAAA1FD3FFB3CE49CC4B5C30" bold="true" pageId="0" pageNumber="147">F</emphasis>
androecium 
<emphasis id="3264C6E28B6CA2F9EED237B885BEC8C8" bold="true" pageId="0" pageNumber="147">G</emphasis>
stem with infructescence, including an immature, opening fruit 
<emphasis id="B5A464F8CB9F0C0767E55538F3B32A3E" bold="true" pageId="0" pageNumber="147">H</emphasis>
view of dehisced fruit (left, below G) showing the seed and aril (right) 
<taxonomicName id="10C93ECFCD92B8D48D873F0FB5DEF0DB" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba cyclobasis" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="cyclobasis">
<emphasis id="A62D56F38E5EE5416F699085B714D839" italics="true" pageId="0" pageNumber="147">Otoba cyclobasis</emphasis>
</taxonomicName>
<emphasis id="577A6658F6B4E327474A0E7C81888820" bold="true" pageId="0" pageNumber="147">I</emphasis>
staminate flowering branch showing leaves with brochidodromous venation, including detail of marginal vein 
<emphasis id="C6BE9ACFDCD347F7DCC741E284568436" bold="true" pageId="0" pageNumber="147">J</emphasis>
staminate flower with one perianth lobe removed, showing the androecium and perianth lobe (above left) 
<emphasis id="77A99114CF5B7F599B2330C32962C9C4" bold="true" pageId="0" pageNumber="147">K</emphasis>
androecium. Illustration by Bobbi Angell, W.W. Thomas &amp; C.J. 
<normalizedToken id="0BED5327CC387648B070D118E80B580D" originalValue="Castaño">Castano</normalizedToken>
5501 (NY) (
<emphasis id="797F85D919315DED8A2D95ECB9E636D1" bold="true" pageId="0" pageNumber="147">A-F</emphasis>
), J. Brand &amp; M. 
<normalizedToken id="31BCC6D4DEA7A554A422A9352DC05C3D" originalValue="Narváez">Narvaez</normalizedToken>
682 (MO) (
<emphasis id="B2593E786F1090254B382A1BAABA615D" bold="true" pageId="0" pageNumber="147">G, H</emphasis>
), E. 
<normalizedToken id="69AF04DB4DB454A0BF3925A4BB0C72C9" originalValue="Narvaéz">Narvaez</normalizedToken>
et al. 1072 (MO-2468293) (
<emphasis id="75199136529A5E270FBDAAF934A0A82D" bold="true" pageId="0" pageNumber="147">I-K</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9E2A4244934EE9BBB0C07BE1EB8F3EB3" pageId="0" pageNumber="147" type="description">
<paragraph id="7191EAE100A9CC2F079251640B804E9F" pageId="0" pageNumber="147">Distinctive characters.</paragraph>
<paragraph id="82FF313BB9200D615ADDBDD5CD5550D6" pageId="0" pageNumber="147">
<taxonomicName id="0A4C54BE79C434E67BB6BCB325561557" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="86F810D9A19FA8506D164244FAE76EEC" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
is recognized by a combination of leaf, inflorescence, and fruit traits. Its small leaf blades have long apices and thin petioles, and lack vernation lines on the abaxial surface. The staminate inflorescences are delicate with flowers on relatively thin, long pedicels; these flowers have sub-membranaceous perianth and lack a swollen-lobed ring, a bowling pin-shaped filament column, and anthers that are lanceolate to oblong. Finally, fruits are small with thin pericarp and membranous aril.
</paragraph>
</subSubSection>
<subSubSection id="21129B80D53603057B3F781E51CBCF27" pageId="0" pageNumber="147" type="etymology">
<paragraph id="6E5D1E78E22BF3C2524A48354D1E1BE6" pageId="0" pageNumber="147">Etymology.</paragraph>
<paragraph id="7DBD8D887C5102DD0595CCFF8593447D" pageId="0" pageNumber="147">
Is a pleasure to name this species after Dr. Scott A. Mori (1941-2020), a wonderful person and skilled botanist; a dedicated explorer of Central and South America humid forests (where this species occurs), especially in the Guianas and the Amazon basin; and an authority on Neotropical 
<taxonomicName id="39A228A220E70FB10A0245BF7F20D758" authorityName="H.Karsten" authorityYear="1882" baseAuthorityName="A.DC." class="Magnoliopsida" family="Lecythidaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="family">Lecythidaceae</taxonomicName>
. His taxonomic and ecological publications gave great inspiration to the first author, as did Dr. 
<normalizedToken id="E5CD270298A3AB73B5778656E4C5C712" originalValue="Mori’s">Mori's</normalizedToken>
personal support. For an account of Dr. 
<normalizedToken id="E1C9EBA42A5E006554873671D11DEC4E" originalValue="Mori’s">Mori's</normalizedToken>
legacy, see 
<bibRefCitation id="6B7CC143E28D553C08E72AC08F288AC5" DOI="https://doi.org/10.1007/s12229-020-09232-0" author="Boom, BM" journalOrPublisher="Botanical Review" pageId="0" pageNumber="147" pagination="149 - 179" refId="B6" refString="Boom, BM, 2020. Scott Alan Mori (1941-2020): An appreciation. Botanical Review 86 (3-4): 149 - 179, DOI: https://doi.org/10.1007/s12229-020-09232-0" title="Scott Alan Mori (1941 - 2020): An appreciation." url="https://doi.org/10.1007/s12229-020-09232-0" volume="86" year="2020">Boom (2020)</bibRefCitation>
, 
<bibRefCitation id="97CC1891B90E6E824CB417D922DBC12C" author="Forget, PM" journalOrPublisher="Ecotropica (Bonn)" pageId="0" pageNumber="147" pagination="1 - 4" publicationUrl="https://www.ecotropica.eu/index.php/ecotropica/article/view/72" refId="B13" refString="Forget, PM, 2020. The &quot;Adopt-a-tree&quot; strategy - a legacy of Scott A. Mori (1941-2020). Ecotropica (Bonn) 22: 1 - 4, https://www.ecotropica.eu/index.php/ecotropica/article/view/72" title="The &quot; Adopt-a-tree &quot; strategy - a legacy of Scott A. Mori (1941 - 2020)." url="https://www.ecotropica.eu/index.php/ecotropica/article/view/72" volume="22" year="2020">Forget (2020)</bibRefCitation>
, and 
<bibRefCitation id="A042B2FCC547EED8463E0FA1862D548C" DOI="https://doi.org/10.1111/btp.12920" author="Prance, G" journalOrPublisher="Biotropica" pageId="0" pageNumber="147" pagination="339 - 346" refId="B34" refString="Prance, G, Naczi, R, Lohmann, LG, 2021. In Remembrance of: Scott A. Mori (1941-2020), Tropical Botanist Extraordinaire. Biotropica 53 (1): 339 - 346, DOI: https://doi.org/10.1111/btp.12920" title="In Remembrance of: Scott A. Mori (1941 - 2020), Tropical Botanist Extraordinaire." url="https://doi.org/10.1111/btp.12920" volume="53" year="2021">Prance et al. (2021)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="575EB94AE9F20BB095DDCA765E430246" pageId="0" pageNumber="147" type="common name">
<paragraph id="13221B1F79E23D2FC9BF840692FF8211" pageId="0" pageNumber="147">Common name.</paragraph>
<paragraph id="C0D58BE82D017BE4EEF733EF8BBB5722" pageId="0" pageNumber="147">
<normalizedToken id="A5EA7CD9A53959D6EDF36F9794784911" originalValue="Cuángare">Cuangare</normalizedToken>
otobo (
<bibRefCitation id="A0F3E417CD4D243996F254F521424F67" author="Cogollo, A" editor="Garcia, N" journalOrPublisher="Instituto Alexander von Humboldt - CORANTIOQUIA - Jardin Botanico Joaquin Antonio Uribe de Medellin - Instituto de Ciencias Naturales de la Universidad Nacional de Colombia - Ministerio de Ambiente, Vivienda y Desarrollo Territorial" pageId="0" pageNumber="147" pagination="155 - 192" refId="B9" refString="Cogollo, A, Velasquez-Rua, C, Garcia, N, 2007. Las miristicaceas. In: Garcia, N, Ed., Libro rojo de plantas de Colombia. Las magnoliaceas, las miristicaceas y las podocarpaceas. Serie libros rojos de especies amenazadas de Colombia. Bogota, Colombia. Instituto Alexander von Humboldt - CORANTIOQUIA - Jardin Botanico Joaquin Antonio Uribe de Medellin - Instituto de Ciencias Naturales de la Universidad Nacional de Colombia - Ministerio de Ambiente, Vivienda y Desarrollo Territorial 5: 155 - 192" title="Las miristicaceas." volume="5" volumeTitle="Libro rojo de plantas de Colombia. Las magnoliaceas, las miristicaceas y las podocarpaceas. Serie libros rojos de especies amenazadas de Colombia. Bogota, Colombia." year="2007">Cogollo et al. 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2E4E7A9A6B78A815C271F25C000EB5D6" pageId="0" pageNumber="147" type="distribution">
<paragraph id="519CEBA053601A51656E680DE1EA1424" pageId="0" pageNumber="147">Distribution.</paragraph>
<paragraph id="28FD8AFE28A4AAF1818D5F86FA78DF66" pageId="0" pageNumber="147">
<taxonomicName id="1476B07F6974A43B05D8FC1B304787CE" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="592E71DD8BE15E3D26FB18D82E3F926A" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
is known only from the humid forests in the Department of Antioquia in northwestern Colombia in the Municipios of 
<normalizedToken id="5DDEF015ABA8A89B55BFDA7D111323D2" originalValue="Mutatá">Mutata</normalizedToken>
, Segovia, 
<normalizedToken id="F78E6F2E8EAC0A3CA7DD113B7D68CA6F" originalValue="Tarazá">Taraza</normalizedToken>
, Turbo, and Valdivia (Fig. 
<figureCitation id="2DE8A72BED0371677B72758F6C4B8A86" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution of Otoba scottmorii (black circle) and O. squamosa (black triangle). The shaded map area corresponds to the department of Antioquia." figureDoi="10.3897/phytokeys.178.64564.figure4" httpUri="https://binary.pensoft.net/fig/550343" pageId="0" pageNumber="147">4</figureCitation>
). Three collections were collected under 100 m elevation (20-80 m), and three others above 400 m elevation (410-730 m).
</paragraph>
<caption id="6886A9AC61E553BBE29BCEEDBC722796" doi="10.3897/phytokeys.178.64564.figure4" httpUri="https://binary.pensoft.net/fig/550343" pageId="0" pageNumber="147" start="Figure 4" startId="F4">
<paragraph id="546A2EFB7E5F68F96B0F081AD0B3ACE3" pageId="0" pageNumber="147">
<emphasis id="BF32D39A2AFE1413E3156C73734FFE50" bold="true" pageId="0" pageNumber="147">Figure 4.</emphasis>
Geographic distribution of 
<taxonomicName id="26E1849FE2B8E5CEAE23648F79B4890C" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="AC94FDAED3A3F6BF22D2BA06A4C62225" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
(black circle) and 
<taxonomicName id="BAF6CAA2641CEF86468F0532907D0ED2" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. squamosa" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="squamosa">
<emphasis id="151FACBAC86D67369D12CC44353CE935" italics="true" pageId="0" pageNumber="147">O. squamosa</emphasis>
</taxonomicName>
(black triangle). The shaded map area corresponds to the department of Antioquia.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0CFBAD2779A03417FCCF50E87A570A34" pageId="0" pageNumber="147" type="phenology">
<paragraph id="877289FA4D553BE54AA45FF95E428B92" pageId="0" pageNumber="147">Phenology.</paragraph>
<paragraph id="13F8AB7609C437623A595DE9CEFC9A92" pageId="0" pageNumber="147">
Herbarium specimens of 
<taxonomicName id="FB19B2298BDD5C6AC16A17A69BCE8C35" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="289398B2BA4415D07F2E20CF1E7A9143" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
have been collected with staminate flowers in May, July, and November, and with fruits in November. Pistillate flowers have yet to be observed.
</paragraph>
</subSubSection>
<subSubSection id="FB132BDBA5290C55F2643560CBE96DF9" pageId="0" pageNumber="147" type="preliminary conservation status">
<paragraph id="8CF1972A3BFCFF666A2A4CB9395C61EF" pageId="0" pageNumber="147">Preliminary conservation status.</paragraph>
<paragraph id="F18DB8F1FBB7098F4C74966549C42A7B" pageId="0" pageNumber="147">
<taxonomicName id="B7E2942CB59E07ED5BAFAC314D9B5240" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="C70B2EF8D7296A1ACCD8DE327607B13B" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
is Endangered following IUCN criteria B2a. Justifying this status, it is known from five localities, has an EOO of 5,341 km2, and an AOO of 20 km2. The regions where it occurs are threatened by high levels of deforestation (
<bibRefCitation id="AFAA7D48F37E1004DF80EA1662C50AE6" author="Gonzalez-Caro, S" editor="Quintero, E" journalOrPublisher="Jardin Botanico de Medellin, Medellin" pageId="0" pageNumber="147" pagination="63 - 80" refId="B18" refString="Gonzalez-Caro, S, Vasquez, A, 2017. Estado de los bosques de Antioquia entre 1990-2015. In: Quintero, E, Benavides, A, Moreno, N, Gonzalez-Caro, S, Eds., Bosques Andinos, estado actual y retos para su conservacion en Antioquia. Jardin Botanico de Medellin, Medellin: 63 - 80" title="Estado de los bosques de Antioquia entre 1990 - 2015." volumeTitle="Bosques Andinos, estado actual y retos para su conservacion en Antioquia." year="2017">
<normalizedToken id="FC267CCED322E37068C93D20418CC5F6" originalValue="González-Caro">Gonzalez-Caro</normalizedToken>
and 
<normalizedToken id="9BD32FB9CF2A4EBD0004C6EDEFF02B4D" originalValue="Vásquez">Vasquez</normalizedToken>
2017
</bibRefCitation>
). Of the relatively few specimens of this species that we were able to verify, the most recent specimen was collected in 1993.
</paragraph>
</subSubSection>
<subSubSection id="496F2C1E6A87F15093D79E7E92E65EE3" pageId="0" pageNumber="147" type="discussion">
<paragraph id="EF4BFE02E4292C7BE6820D6EF45262CD" pageId="0" pageNumber="147">Discussion.</paragraph>
<paragraph id="BB644136CD668EABF61C8304066D0FA7" pageId="0" pageNumber="147">
<taxonomicName id="5ED910BF62ACDD8CB70D32E2EBE093B5" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="A281B234610CCA9BD274D39CF2A927F7" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="BEB2AA4FE2411932A1D4D59816142356" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="B6386F2B9828565CC5415BBC5ED280F8" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
and 
<taxonomicName id="22C489D6897D107A86DB15B28A367400" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. vespertilio" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="vespertilio">
<emphasis id="81E07C0883EFE5AB26CE2C7EAB044EA6" italics="true" pageId="0" pageNumber="147">O. vespertilio</emphasis>
</taxonomicName>
D. Santam. &amp; J.E. 
<normalizedToken id="F308917AB94C2E6FFD3835CB7A3AE071" originalValue="Jiménez">Jimenez</normalizedToken>
, from Costa Rica and Panama, and 
<taxonomicName id="1CD01BC0BC2F94721DE7A7538AD04D50" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. cyclobasis" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="cyclobasis">
<emphasis id="737D686A1195D2BFB208F5F598133074" italics="true" pageId="0" pageNumber="147">O. cyclobasis</emphasis>
</taxonomicName>
T.S. Jaram. &amp; Balslev from Ecuador. They all have relatively small leaf blades of similar shape and thin petioles, and usually lack vernation lines (Figs 
<figureCitation id="8F84E210D0B68E8B637E9039B75CE018" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5</figureCitation>
, 
<figureCitation id="8450AF7A3A49FD24EBAE0AE973F9B313" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparations of herbarium specimens of Otoba acuminata (A L. Poveda et al. 3793, MO) O. cyclobasis (B P. Mendez et al. 156, MO) O. scottmorii (C W. W. Thomas &amp; C. J. Castano 5501, NY), and O. vespertilio (D R. Aguilar et al. 5024, MO)." figureDoi="10.3897/phytokeys.178.64564.figure6" httpUri="https://binary.pensoft.net/fig/550345" pageId="0" pageNumber="147">6</figureCitation>
); staminate flowers with short perianth and small anthers; and similarly sized fruit with thin pericarp (Fig. 
<figureCitation id="3B56E14A8839E656ED5E96AE2D536100" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5</figureCitation>
). The four species can be distinguished by the characteristics in Table 
<tableCitation id="14FD93433CF52FC4FC811A1FA9DC06B2" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of Otoba scottmorii with similar species (Fig. 6)." httpUri="http://table.plazi.org/id/ED3B54066976A8C722BF5968CC02ED0A" pageId="0" pageNumber="147" tableUuid="ED3B54066976A8C722BF5968CC02ED0A">2</tableCitation>
.
</paragraph>
<caption id="ED3B54066976A8C722BF5968CC02ED0A" ID-Table-UUID="ED3B54066976A8C722BF5968CC02ED0A" httpUri="http://table.plazi.org/id/ED3B54066976A8C722BF5968CC02ED0A" pageId="0" pageNumber="147" start="Table 2" startId="T2">
<paragraph id="195A403224AD4CA5E805A2621F357218" pageId="0" pageNumber="147">
<emphasis id="AA520DDB3F134AAA933A17E56A3704F7" bold="true" pageId="0" pageNumber="147">Table 2.</emphasis>
Comparison of 
<taxonomicName id="A4DEC3DD86E1EC3604594E7E405CA34C" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="849120D63523A8271283D56B63D7577E" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
with similar species (Fig. 
<figureCitation id="79586A3B2EA5BFDB8488F554155D4DDA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparations of herbarium specimens of Otoba acuminata (A L. Poveda et al. 3793, MO) O. cyclobasis (B P. Mendez et al. 156, MO) O. scottmorii (C W. W. Thomas &amp; C. J. Castano 5501, NY), and O. vespertilio (D R. Aguilar et al. 5024, MO)." figureDoi="10.3897/phytokeys.178.64564.figure6" httpUri="https://binary.pensoft.net/fig/550345" pageId="0" pageNumber="147">6</figureCitation>
).
</paragraph>
</caption>
<paragraph id="10E40D4B5A1B9DDF75C062915C6CB372" pageId="0" pageNumber="147">
<table id="C923C8353EC1E66D706DE1E7605EBA33" pageId="0" pageNumber="147">
<tr id="D4205F6533E422222EA36768B4F333D8" pageId="0" pageNumber="147">
<th id="EAB9463DCED4C6A5CBAD8E8A1D504CAF" colspan="1" pageId="0" pageNumber="147" rowspan="1">Taxon</th>
<th id="BD4CDB4030F1E21C97457C72A0C8ECD8" colspan="1" pageId="0" pageNumber="147" rowspan="1">
<taxonomicName id="50B8F956C05D71D7BAB137E925A94D23" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. scottmorii" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="scottmorii">
<emphasis id="53EB10BFA27BED02422B1C37AD37BCF5" italics="true" pageId="0" pageNumber="147">O. scottmorii</emphasis>
</taxonomicName>
</th>
<th id="7491892C92940DE45495AE5E9E807D87" colspan="1" pageId="0" pageNumber="147" rowspan="1">
<taxonomicName id="B972CE733411140A6681BE67B26C9F5D" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="5537C54056BA5B19E1C0747DDC4F194A" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
</th>
<th id="8A36049C87E0C1FA0671F3F405A3830C" colspan="1" pageId="0" pageNumber="147" rowspan="1">
<taxonomicName id="C868B3A9B334D2AD2AF7CB84DDE6B85C" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. cyclobasis" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="cyclobasis">
<emphasis id="6EE2840A7EE558E029D9B79505009B0D" italics="true" pageId="0" pageNumber="147">O. cyclobasis</emphasis>
</taxonomicName>
</th>
<th id="B93E3D7F5F7CC61350DA422E0F05CE65" colspan="1" pageId="0" pageNumber="147" rowspan="1">
<taxonomicName id="A07BB35298FC9C7F592135343B39C91A" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. vespertilio" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="vespertilio">
<emphasis id="EC05AC8500FFCC2B6C2A9A5CCB167FF2" italics="true" pageId="0" pageNumber="147">O. vespertilio</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="771D19DE219D558EC120FBABE7B8914E" pageId="0" pageNumber="147">
<td id="3D515F84419623174FFAB92708D1E2EC" colspan="1" pageId="0" pageNumber="147" rowspan="1">Petiole length (cm)</td>
<td id="41A584572AE117E496089F13F650B22F" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.5-1.1</td>
<td id="426A6B581198D72AF9F363424BD51E00" colspan="1" pageId="0" pageNumber="147" rowspan="1">(1.1-) 1.8-2.6</td>
<td id="CB1E5723D1E0CFD71DA7A4450A8C7768" colspan="1" pageId="0" pageNumber="147" rowspan="1">1-1.5‡</td>
<td id="117525910ADF5C98403E12E113957838" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.8-2</td>
</tr>
<tr id="CA5131473C2D75C85E2FAB6179626C8C" pageId="0" pageNumber="147">
<td id="EFE1465CB5CDB8F11B28E37D273F6369" colspan="1" pageId="0" pageNumber="147" rowspan="1">Leaf size (cm)</td>
<td id="9882E05E5B0794D0CB38CECA5282FA11" colspan="1" pageId="0" pageNumber="147" rowspan="1">
5.5-8.5 (-11.2) 
<normalizedToken id="E639865563222C4B1340E66167271D61" originalValue="×">x</normalizedToken>
1.7-2.5 (-3.7) (Fig. 
<figureCitation id="AC05995830AF9C084320C0F164FD7CC6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5B</figureCitation>
)
</td>
<td id="7B764AB96FA85BDB71F8E22594BEE712" colspan="1" pageId="0" pageNumber="147" rowspan="1">
(6.5-) 8.5-15.5 
<normalizedToken id="CF59C59681A414A0365E38CEBAAE86FD" originalValue="×">x</normalizedToken>
(2.7-) 4.4-6.1 (Fig. 
<figureCitation id="FD97E2305CC897B228837F97293CC4DA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5A</figureCitation>
)
</td>
<td id="B89755CE7750753DDCD99EDC826D14C4" colspan="1" pageId="0" pageNumber="147" rowspan="1">
9-14 (-18) 
<normalizedToken id="9E9B7297D55745B3D554318475CB015C" originalValue="×">x</normalizedToken>
3-5‡ (Fig. 
<figureCitation id="2FC6741CA7A6F871AB438082527513BE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5C</figureCitation>
)
</td>
<td id="024485F4F81729BA4F754B481EDB48A9" colspan="1" pageId="0" pageNumber="147" rowspan="1">
5.3-12.5 
<normalizedToken id="FE3AD816BD065F9BB7394E340FACBE70" originalValue="×">x</normalizedToken>
2-3.5 (-4.8) (Fig. 
<figureCitation id="C5F7D92B09B0641653790EC50A7E3E7D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5D</figureCitation>
)
</td>
</tr>
<tr id="103E5483E78894003BFCA95B1DB51A68" pageId="0" pageNumber="147">
<td id="0C2A26611304A3530BD6E2D2BE55A2B1" colspan="1" pageId="0" pageNumber="147" rowspan="1">Lateral vein number</td>
<td id="743F7938A3F6E99612F439EFCD7F5A6B" colspan="1" pageId="0" pageNumber="147" rowspan="1">4-7</td>
<td id="C3FCD9B01F913B4C9EFD78ECED7C0D4D" colspan="1" pageId="0" pageNumber="147" rowspan="1">8-12</td>
<td id="FBAFA3F7399F1E44CE65219EC609861F" colspan="1" pageId="0" pageNumber="147" rowspan="1">14-17‡</td>
<td id="5F535EC00B77BC0A78BA27ACB9CE31A4" colspan="1" pageId="0" pageNumber="147" rowspan="1">6-8</td>
</tr>
<tr id="2F8263DE6C8D66E2D9653C3263EE1EF4" pageId="0" pageNumber="147">
<td id="79AE978900AEE8F66C25239D60E0259D" colspan="1" pageId="0" pageNumber="147" rowspan="1">Staminate flower pedicel length (mm)</td>
<td id="C33838A9403B60AB35092BAC5A10F90D" colspan="1" pageId="0" pageNumber="147" rowspan="1">3-5</td>
<td id="811305C736ED20387FA3DCE507E0981B" colspan="1" pageId="0" pageNumber="147" rowspan="1">5-8</td>
<td id="04000B71F601E1F7FBBB6FD8E21ED6A5" colspan="1" pageId="0" pageNumber="147" rowspan="1">4-5‡</td>
<td id="BFA8723DBED6A612FFCB640D82FCB6A2" colspan="1" pageId="0" pageNumber="147" rowspan="1">2-3.6</td>
</tr>
<tr id="99038BB5BB46B973A3F2D37A551CE525" pageId="0" pageNumber="147">
<td id="36EC2ECFB168B92F5A38C2D2014AF722" colspan="1" pageId="0" pageNumber="147" rowspan="1">Staminate perianth length (mm)</td>
<td id="25CA2DC5806AE0C4A34FBC1034EBF0F4" colspan="1" pageId="0" pageNumber="147" rowspan="1">2-2.5</td>
<td id="FD97506E5CB294E3812E7611C6393B00" colspan="1" pageId="0" pageNumber="147" rowspan="1">4.5-5.5</td>
<td id="9FD4C770B185E7A553202921EFDC64F1" colspan="1" pageId="0" pageNumber="147" rowspan="1">ca. 2.5‡</td>
<td id="11187DDA46CEC65ED93875C4597C0054" colspan="1" pageId="0" pageNumber="147" rowspan="1">2-3</td>
</tr>
<tr id="68490815C9DC9C743741702AE7446AB1" pageId="0" pageNumber="147">
<td id="CBBE3033201B79DBF1957815D6899652" colspan="1" pageId="0" pageNumber="147" rowspan="1">Perianth lobe width (mm)</td>
<td id="7ACE48FFFB253738086B0218820EC05C" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.8-1.1</td>
<td id="70848267FDBFA4B7889A5C82448989ED" colspan="1" pageId="0" pageNumber="147" rowspan="1">1.5-2</td>
<td id="4288731961848C1822D9C75554601954" colspan="1" pageId="0" pageNumber="147" rowspan="1">ca. 1 §</td>
<td id="7D79EC6D756931879CEFEC191FDCBFB1" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.5-1</td>
</tr>
<tr id="E635F6F57D530A433A64DF71A5311E7D" pageId="0" pageNumber="147">
<td id="B5D88753735F931B8CB05BC47E19EFCA" colspan="1" pageId="0" pageNumber="147" rowspan="1">Ring</td>
<td id="1C741425EBB08C5474ECD155DAF36346" colspan="1" pageId="0" pageNumber="147" rowspan="1">Absent</td>
<td id="14BBB727E3C6637883783DAA75917944" colspan="1" pageId="0" pageNumber="147" rowspan="1">Absent</td>
<td id="D92CECEB96C73F214C3A79129836E460" colspan="1" pageId="0" pageNumber="147" rowspan="1">Present‡</td>
<td id="197557AE8833F3C4B849CEA9B15CC17C" colspan="1" pageId="0" pageNumber="147" rowspan="1">Absent</td>
</tr>
<tr id="6519233DD6E82406D839403E9158E529" pageId="0" pageNumber="147">
<td id="1E93C8EF9280F22B3C5BCCDC5AE594C6" colspan="1" pageId="0" pageNumber="147" rowspan="1">Filament column length (mm) and shape</td>
<td id="B07CFC193D5BF39E62996DEC643240D0" colspan="1" pageId="0" pageNumber="147" rowspan="1">(1.3-) 1.5-1.6, bowling pin</td>
<td id="AC797896DDE78AAF5BD24438C1FC3D4A" colspan="1" pageId="0" pageNumber="147" rowspan="1">2-2.7, cylindrical, ovoid, to pyriform</td>
<td id="F1EC463401FD65F364D29B2FFACB8425" colspan="1" pageId="0" pageNumber="147" rowspan="1">1, cylindrical</td>
<td id="F8AEC52ADF5BA7F6D151FE426C694510" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.6-1.5, cylindrical</td>
</tr>
<tr id="E25380F34C9B2521518F993942126524" pageId="0" pageNumber="147">
<td id="02C05813A62A39054D7981D2760C1843" colspan="1" pageId="0" pageNumber="147" rowspan="1">Anther length (mm) and shape</td>
<td id="270600C3D31BEF6ECA78560B3D986D2F" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.2-0.4 lanceolate to oblong</td>
<td id="FFD8537885E3A0F01344CFEC6A72FCA5" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.45-0.8, oblong</td>
<td id="BB080DBE855AADD423688CB4C14DEB02" colspan="1" pageId="0" pageNumber="147" rowspan="1">ca. 0.3, globose‡</td>
<td id="05E7F0EF427CB585AA4E4AB100FE1340" colspan="1" pageId="0" pageNumber="147" rowspan="1">ca. 0.2-0.3, globose to subglobose</td>
</tr>
<tr id="686B39A53FDD840253FF4E325A4A8155" pageId="0" pageNumber="147">
<td id="277E6E04D7BFF1DDE7F4735E648A593C" colspan="1" pageId="0" pageNumber="147" rowspan="1">Ovary vestiture</td>
<td id="A2DB6A30BE3D1ECEB899AFD42F92E732" colspan="1" pageId="0" pageNumber="147" rowspan="1">Unknown</td>
<td id="714B639034FDC1F0D0666C62EF92DCF6" colspan="1" pageId="0" pageNumber="147" rowspan="1">Pubescent</td>
<td id="544854B0465BC61E43B02C3FE3E2AF07" colspan="1" pageId="0" pageNumber="147" rowspan="1">Glabrous‡</td>
<td id="7A32E4E686010C1510936C8A85E49C46" colspan="1" pageId="0" pageNumber="147" rowspan="1">Glabrous</td>
</tr>
<tr id="6CB76806C555F742009A8403C17E2961" pageId="0" pageNumber="147">
<td id="771203D3E2465D7311FD903CF4CCC0F8" colspan="1" pageId="0" pageNumber="147" rowspan="1">Fruit length and width (cm)</td>
<td id="463761EF57418CC108AD2ABEB906B863" colspan="1" pageId="0" pageNumber="147" rowspan="1">
1.8-2.1 
<normalizedToken id="7F42A0BABDC0FC09A22E7D490E341CF8" originalValue="×">x</normalizedToken>
1.5-1.7 (Fig. 
<figureCitation id="032A96767B785E70E96FB5CC02CE307D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5</figureCitation>
1, 2)
</td>
<td id="2DF70C9F72ADB8A14BCB8DDCFC4D28EE" colspan="1" pageId="0" pageNumber="147" rowspan="1">
1.9-2.5 (-2.8) 
<normalizedToken id="D3A4A8D85344F895914FBBB7EFFDA309" originalValue="×">x</normalizedToken>
1.7-2.2 (-2.4) (Fig. 
<figureCitation id="0E245BE6A433740CA19851DE17DFA3C9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5A</figureCitation>
1)
</td>
<td id="A383E1BA30DE561DCD4FC1E63E35E67C" colspan="1" pageId="0" pageNumber="147" rowspan="1">
1.5-2.5 
<normalizedToken id="A01A2EDEEC1FC98CEA94DA01EE51BEC4" originalValue="×">x</normalizedToken>
1.5-2‡ (Fig. 
<figureCitation id="A0C3DCAE4805B331A28C68E3A722BC6C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5C</figureCitation>
1, 2)
</td>
<td id="1F258F966BDB57CC36596365E08D1EB3" colspan="1" pageId="0" pageNumber="147" rowspan="1">
2.2-2.7 
<normalizedToken id="7E8DB8EE4D0A5E5AD7674EC7B9CDFEF2" originalValue="×">x</normalizedToken>
1.6-1.7 (Fig. 
<figureCitation id="6933A421ADBC8AFB37251AA5C328CBDE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5D</figureCitation>
1)
</td>
</tr>
<tr id="70669DD6AB355747DE8BB85898F629A6" pageId="0" pageNumber="147">
<td id="4004AF7C9CEBCAEE714C6CEA85B841F8" colspan="1" pageId="0" pageNumber="147" rowspan="1">Pericarp thickness (mm)</td>
<td id="05BBAE5C18968B37AC1B5F4355CBCBD0" colspan="1" pageId="0" pageNumber="147" rowspan="1">0.68-0.89</td>
<td id="16BD2FE13FC2EA26EE2142293417F7AE" colspan="1" pageId="0" pageNumber="147" rowspan="1">1-2.1</td>
<td id="AE4E1788F512A8E06C5418D06C0C26BE" colspan="1" pageId="0" pageNumber="147" rowspan="1">1-2‡</td>
<td id="0120FC225A1FB4793B0DA95348D8CAEC" colspan="1" pageId="0" pageNumber="147" rowspan="1">1.3-1.8</td>
</tr>
<tr id="A6EBD4D248437618857678FD574D6D74" pageId="0" pageNumber="147">
<td id="4A094F61AEE6C289DCB5F3B4CC2840BA" colspan="1" pageId="0" pageNumber="147" rowspan="1">Seed length and width (cm)</td>
<td id="031C0AB3C21B10BC22FA8CEA94A9B179" colspan="1" pageId="0" pageNumber="147" rowspan="1">
1.6 
<normalizedToken id="60147F99B76C5BFF9D32CA4A389D24A1" originalValue="×">x</normalizedToken>
1.5
</td>
<td id="C4F6B18AAC9E61CF38CE3DE1A3AC2282" colspan="1" pageId="0" pageNumber="147" rowspan="1">
ca. 2.2 
<normalizedToken id="9BB98EB816FFD540952873A17985C5F9" originalValue="×">x</normalizedToken>
1.9
</td>
<td id="C3599AE5088A308F3D3FC22F61BD5DA8" colspan="1" pageId="0" pageNumber="147" rowspan="1">
1-2 
<normalizedToken id="9D9F16DE2AF633722D065599832439EE" originalValue="×">x</normalizedToken>
1-1.5‡
</td>
<td id="CD302696BA54D19AE3DC58C0C2D24B97" colspan="1" pageId="0" pageNumber="147" rowspan="1">
1.8-1.9 
<normalizedToken id="582C9AD9172FB60E2C9B867F632F2B76" originalValue="×">x</normalizedToken>
ca. 1.7 cm
</td>
</tr>
<tr id="296985F2E9DEE8921D919CD1689E33A7" pageId="0" pageNumber="147">
<td id="A39701A9BDBCF9FD13CF3FA164B31BEB" colspan="1" pageId="0" pageNumber="147" rowspan="1">Distribution and elevation</td>
<td id="1BE217222CE9ACCF3F9F08C5A1DEE118" colspan="1" pageId="0" pageNumber="147" rowspan="1">Colombia, 20-80 m and 410-730 m</td>
<td id="1C18F905B39C01B73884886DB87744F7" colspan="1" pageId="0" pageNumber="147" rowspan="1">Costa Rica, 100-900 m; Panama, 500-1000</td>
<td id="A9C917B0F7ECA19DB14D5404F58E8181" colspan="1" pageId="0" pageNumber="147" rowspan="1">Ecuador, 150-300 m‡</td>
<td id="CBC157CE7377680BFB337AE7A933E5DD" colspan="1" pageId="0" pageNumber="147" rowspan="1">Costa Rica 0-200 m, Panama 100-800 m</td>
</tr>
</table>
</paragraph>
<paragraph id="FF5E86590744794C51F11E396441A1DE" pageId="0" pageNumber="147">
<tableNote id="287C6AA2F77F878D25B0BBFFAA5B2BB6" pageId="0" pageNumber="147">
‡Information from 
<bibRefCitation id="5D0790422CE0655091EACB04C11E9AF4" DOI="https://doi.org/10.11646/phytotaxa.441.2.3" author="Jaramillo-Vivanco, T" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="147" pagination="143 - 175" refId="B25" refString="Jaramillo-Vivanco, T, Balslev, H, 2020. Revision of Otoba (Myristicaceae). Phytotaxa 441 (2): 143 - 175, DOI: https://doi.org/10.11646/phytotaxa.441.2.3" title="Revision of Otoba (Myristicaceae)." url="https://doi.org/10.11646/phytotaxa.441.2.3" volume="441" year="2020">Jaramillo-Vivanco and Balslev 2020</bibRefCitation>
; §from E. 
<normalizedToken id="4F54BE3C0A4DCA81563C0349E5848272" originalValue="Narvaéz">Narvaez</normalizedToken>
et al. 1072, MO-2 sheets.
</tableNote>
</paragraph>
<caption id="E160C82DD176DFDE93A743C945EE574B" doi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147" start="Figure 5" startId="F5">
<paragraph id="4B6A8405C1CAA6E732A3BA43B1A9EB1B" pageId="0" pageNumber="147">
<emphasis id="46EA35A637C6FC8787984854C792E7DE" bold="true" pageId="0" pageNumber="147">Figure 5.</emphasis>
Comparison of leaf blades and fruits of 
<taxonomicName id="413B9B6D0CFC4A4F99F21280D73BBFFC" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="376FD904707B52F9CF09161A8A2DD9E9" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
(
<emphasis id="DD076072B4AFDA6C2A9519088AFC0D8D" bold="true" pageId="0" pageNumber="147">A</emphasis>
L. Poveda et al. 3793, MO 
<emphasis id="F7150D931CF7C4BCADB9B39B222F9CFD" bold="true" pageId="0" pageNumber="147">A1</emphasis>
from H.W. Churchill &amp; G.C. de Nevers 4976 MO; immature fruit) 
<taxonomicName id="A752A6488F9D9AA8E25C84A95E0C33D7" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. scottmorii" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="scottmorii">
<emphasis id="EE7B2F1CF51891B519C37F6DD8F5E930" italics="true" pageId="0" pageNumber="147">O. scottmorii</emphasis>
</taxonomicName>
(
<emphasis id="83374925AE3D4830C56E1AE19B16D279" bold="true" pageId="0" pageNumber="147">B</emphasis>
R. Callejas et al. 3440 MO 
<emphasis id="139FFA65BA37B201DBEEF3BBF9102714" bold="true" pageId="0" pageNumber="147">B1-B2</emphasis>
from R. Callejas et al. 5789, MO) 
<taxonomicName id="A085778647F3309E2FD6A139A2834024" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. cyclobasis" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="cyclobasis">
<emphasis id="81361A6EA31BF3AB355A1C429A24F8C1" italics="true" pageId="0" pageNumber="147">O. cyclobasis</emphasis>
</taxonomicName>
(
<emphasis id="6AEAB6770DDBBDBECB4AE90BE7CD84A8" bold="true" pageId="0" pageNumber="147">C</emphasis>
E. 
<normalizedToken id="543C7AF8ADB980D05A950BD7A75A476F" originalValue="Narvaéz">Narvaez</normalizedToken>
et al. 1072, MO 
<emphasis id="AC85B8BB60C72BDE4B03747F6BF9B0BE" bold="true" pageId="0" pageNumber="147">C1-C2</emphasis>
from W.A. Palacios &amp; M. Tirado 11318 MO) and 
<taxonomicName id="CC35724697E457057598C481539A1431" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. vespertilio" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="vespertilio">
<emphasis id="A340912150FAE00E82F80B3F480064E1" italics="true" pageId="0" pageNumber="147">O. vespertilio</emphasis>
</taxonomicName>
(
<emphasis id="100EBA526981D38735760F475645A56C" bold="true" pageId="0" pageNumber="147">D</emphasis>
G. McPherson 13597, MO, juvenile leaf, 
<emphasis id="E802292E024511701F431BCD824EFFB9" bold="true" pageId="0" pageNumber="147">D1</emphasis>
G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm.
</paragraph>
</caption>
<caption id="DDB08FD43B7AC271A5A812AFEAC0FB72" doi="10.3897/phytokeys.178.64564.figure6" httpUri="https://binary.pensoft.net/fig/550345" pageId="0" pageNumber="147" start="Figure 6" startId="F6">
<paragraph id="56C7A1A065CA3B668614DC8AEFAFA33A" pageId="0" pageNumber="147">
<emphasis id="829E3FD33BFA693C9A0B02BCE931482F" bold="true" pageId="0" pageNumber="147">Figure 6.</emphasis>
Comparations of herbarium specimens of 
<taxonomicName id="7A7DB0B2C09D535EA0B3E24495EDF3B3" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba acuminata" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="acuminata">
<emphasis id="9BA166523CEF26104E5A99050A2F53C1" italics="true" pageId="0" pageNumber="147">Otoba acuminata</emphasis>
</taxonomicName>
(
<emphasis id="DE8D00C8FB58A30C7193DB0E12E1974E" bold="true" pageId="0" pageNumber="147">A</emphasis>
L. Poveda et al. 3793, MO) 
<taxonomicName id="74C2B5A2DE353E1F75CE51873EA7C78C" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. cyclobasis" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="cyclobasis">
<emphasis id="A62B45B60B366C870153DECD158B1522" italics="true" pageId="0" pageNumber="147">O. cyclobasis</emphasis>
</taxonomicName>
(
<emphasis id="0D1903C86FABD2DBD9F14FF04B136F51" bold="true" pageId="0" pageNumber="147">B</emphasis>
P. 
<normalizedToken id="CDB56D66E4F58D0A02610B4ECEF3C6EF" originalValue="Méndez">Mendez</normalizedToken>
et al. 156, MO) 
<taxonomicName id="3DA05BA5EFC0CF85218B2D93AF0A63B2" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. scottmorii" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="scottmorii">
<emphasis id="D4078ECC23D4C8371A6D933FFA62F408" italics="true" pageId="0" pageNumber="147">O. scottmorii</emphasis>
</taxonomicName>
(
<emphasis id="41C151E86B6059003BCFF1CCB4521CAC" bold="true" pageId="0" pageNumber="147">C</emphasis>
W.W. Thomas &amp; C.J. 
<normalizedToken id="B852B75BF9BC98A96300BCB68C847FE3" originalValue="Castaño">Castano</normalizedToken>
5501, NY), and 
<taxonomicName id="E2603E26D3B1ECFE1B15D95BD2F2FFAC" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. vespertilio" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="vespertilio">
<emphasis id="F4CF0D89317B07FD6365EA9DB43E9701" italics="true" pageId="0" pageNumber="147">O. vespertilio</emphasis>
</taxonomicName>
(
<emphasis id="9F818F9DA7AE9B3FCF1200440D60E1E2" bold="true" pageId="0" pageNumber="147">D</emphasis>
R. Aguilar et al. 5024, MO).
</paragraph>
</caption>
<paragraph id="6E589752A31C3641A51E33B9A41BABFC" pageId="0" pageNumber="147">
As mentioned in the introduction, 
<taxonomicName id="4948EDE575838468B7C12C1402492A20" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="7AB44A7FE70AD4050DDE6EC25D2C2734" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
was confused with the Mesoamerican species 
<taxonomicName id="DBF05A58C36B3C46B6C417F9CC802E7A" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="989E6A10BA9360B494ACB8BE37F30531" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
, which we now consider to be endemic to the Caribbean slope of Costa Rica and Panama. The specimens referred to as 
<taxonomicName id="E4C9AF91B986B627CDB45FFD6735A824" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="CC5324C6E954253AB7036841E0B279D3" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
from Colombia in recent floras or checklists (e.g., 
<bibRefCitation id="D55F7778C675988B15018124A7DB5F7D" author="Cogollo, A" editor="Garcia, N" journalOrPublisher="Instituto Alexander von Humboldt - CORANTIOQUIA - Jardin Botanico Joaquin Antonio Uribe de Medellin - Instituto de Ciencias Naturales de la Universidad Nacional de Colombia - Ministerio de Ambiente, Vivienda y Desarrollo Territorial" pageId="0" pageNumber="147" pagination="155 - 192" refId="B9" refString="Cogollo, A, Velasquez-Rua, C, Garcia, N, 2007. Las miristicaceas. In: Garcia, N, Ed., Libro rojo de plantas de Colombia. Las magnoliaceas, las miristicaceas y las podocarpaceas. Serie libros rojos de especies amenazadas de Colombia. Bogota, Colombia. Instituto Alexander von Humboldt - CORANTIOQUIA - Jardin Botanico Joaquin Antonio Uribe de Medellin - Instituto de Ciencias Naturales de la Universidad Nacional de Colombia - Ministerio de Ambiente, Vivienda y Desarrollo Territorial 5: 155 - 192" title="Las miristicaceas." volume="5" volumeTitle="Libro rojo de plantas de Colombia. Las magnoliaceas, las miristicaceas y las podocarpaceas. Serie libros rojos de especies amenazadas de Colombia. Bogota, Colombia." year="2007">Cogollo et al. 2007</bibRefCitation>
; 
<bibRefCitation id="9A0E01058F20AFA3FEA5576F52F97C00" author="Cogollo, A" editor="Idarraga-Piedrahita, A" journalOrPublisher="Universidad de Antioquia, Medellin" pageId="0" pageNumber="147" pagination="641 - 644" refId="B8" refString="Cogollo, A, 2011. Myristicaceae. In: Idarraga-Piedrahita, A, Ortiz, RDC, Callejas Posada, R, Merello, M, Eds., Flora de Antioquia: catalogo de las plantas vasculares, listado de las plantas vasculares del departamento de Antioquia. Universidad de Antioquia, Medellin: 641 - 644" title="Myristicaceae." volumeTitle="Flora de Antioquia: catalogo de las plantas vasculares, listado de las plantas vasculares del departamento de Antioquia." year="2011">Cogollo 2011</bibRefCitation>
; 
<bibRefCitation id="C7A8BF9D8D877A37BE24BA2849BD6683" DOI="https://doi.org/10.1126/science.aao0398" author="Ulloa Ulloa, C" journalOrPublisher="Science" pageId="0" pageNumber="147" pagination="1614 - 1617" refId="B44" refString="Ulloa Ulloa, C, Acevedo-Rodriguez, P, Beck, S, Belgrano, MJ, Bernal, R, Berry, PE, Brako, L, Celis, M, Davidse, G, Forzza, RC, Gradstein, SR, Hokche, O, Leon, B, Leon-Yanez, S, Magill, RE, Neill, DA, Nee, M, Raven, PH, Stimmel, H, Strong, MT, Villasenor, JL, Zarucchi, JL, Zuloaga, FO, Jorgensen, PM, 2017. An integrated assessment of the vascular plant species of the Americas. Science 358 (6370): 1614 - 1617, DOI: https://doi.org/10.1126/science.aao0398" title="An integrated assessment of the vascular plant species of the Americas." url="https://doi.org/10.1126/science.aao0398" volume="358" year="2017">Ulloa Ulloa et al. 2017</bibRefCitation>
; 
<bibRefCitation id="2223FBEE71463BB6B1BB82AD6C0C50CF" DOI="https://doi.org/10.11646/phytotaxa.441.2.3" author="Jaramillo-Vivanco, T" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="147" pagination="143 - 175" refId="B25" refString="Jaramillo-Vivanco, T, Balslev, H, 2020. Revision of Otoba (Myristicaceae). Phytotaxa 441 (2): 143 - 175, DOI: https://doi.org/10.11646/phytotaxa.441.2.3" title="Revision of Otoba (Myristicaceae)." url="https://doi.org/10.11646/phytotaxa.441.2.3" volume="441" year="2020">Jaramillo-Vivanco and Balslev 2020</bibRefCitation>
) correspond to 
<taxonomicName id="22CDD4DC18F175C5A9F2037708287927" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. scottmorii" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="scottmorii">
<emphasis id="0AEB11A4755C6A19C8CA96A2B237B730" italics="true" pageId="0" pageNumber="147">O. scottmorii</emphasis>
</taxonomicName>
. The confusion between these two species is likely driven by the similar morphology of the leaf blades (i.e., size and shape, usually without vernation lines, and thin petioles). However, 
<taxonomicName id="416F894CCC71FAF01EDF3C0E9D59A370" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. scottmorii" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="scottmorii">
<emphasis id="7FFD11845CC5AC7D4CEBEF134B1693D1" italics="true" pageId="0" pageNumber="147">O. scottmorii</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="67336EF1D42C29CA253ED1FC8ED38B02" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="70BDD52E8ACF9D1F3BBC0CA637837278" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
in having narrower leaf blades (Fig. 
<figureCitation id="78B660A98A1488E767C41B1A8A737FB4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of leaf blades and fruits of O. acuminata (A L. Poveda et al. 3793, MO A 1 from H. W. Churchill &amp; G. C. de Nevers 4976 MO; immature fruit) O. scottmorii (B R. Callejas et al. 3440 MO B 1 - B 2 from R. Callejas et al. 5789, MO) O. cyclobasis (C E. Narvaez et al. 1072, MO C 1 - C 2 from W. A. Palacios &amp; M. Tirado 11318 MO) and O. vespertilio (D G. McPherson 13597, MO, juvenile leaf, D 1 G. McPherson 12543, MO). Scale bars: leaves: 2 cm; fruits: 0.75 cm." figureDoi="10.3897/phytokeys.178.64564.figure5" httpUri="https://binary.pensoft.net/fig/550344" pageId="0" pageNumber="147">5B, A</figureCitation>
respectively), staminate flowers with short perianth and narrow perianth lobes, a longer column of filaments, and fruits with a smooth surface and thin pericarp (Table 
<tableCitation id="B0B5182BB510F527000C39CF26FE30D5" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of Otoba scottmorii with similar species (Fig. 6)." httpUri="http://table.plazi.org/id/ED3B54066976A8C722BF5968CC02ED0A" pageId="0" pageNumber="147" tableUuid="ED3B54066976A8C722BF5968CC02ED0A">2</tableCitation>
). Although we have not observed the pistillate flowers of 
<taxonomicName id="FD6D75E48EE88784F03D78C7DC8756DC" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. scottmorii" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="scottmorii">
<emphasis id="E1DB4FC5460141DC5306806B13F93808" italics="true" pageId="0" pageNumber="147">O. scottmorii</emphasis>
</taxonomicName>
, it is likely that the ovary is glabrous as the fruits do not have traces of trichomes, while in 
<taxonomicName id="5D8E0F65E040A584E52A4CEAD45E03B3" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="88F9F60A1EDD19802DCA42F35177A887" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
the ovary is pubescent; see for example: A. 
<normalizedToken id="5F21B41BB2D52252ED034971AD4E5873" originalValue="Rodríguez">Rodriguez</normalizedToken>
et al. 1559 (CR-2 sheets, MO); A. Estrada et al. 4829 (CR); B. Hammel &amp; M. Grayum 14288 (CR, INPA [digital image], MEXU [digital image], MO). For a list of specimens that correspond to 
<taxonomicName id="91AD3C93A95B0641DD870C820264F677" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="12C4EA441566000A48712383A1D41584" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
, see 
<bibRefCitation id="A4D6AF8A48343AF86A11889B59399811" DOI="https://doi.org/10.1007/s12228-019-09585-w" author="Santamaria-Aguilar, D" journalOrPublisher="Brittonia" pageId="0" pageNumber="147" pagination="369 - 380" refId="B35" refString="Santamaria-Aguilar, D, Jimenez, JE, Aguilar, R, 2019. Otoba vespertilio (Myristicaceae), una especie nueva de Mesoamerica. Brittonia 71 (4): 369 - 380, DOI: https://doi.org/10.1007/s12228-019-09585-w" title="Otoba vespertilio (Myristicaceae), una especie nueva de Mesoamerica." url="https://doi.org/10.1007/s12228-019-09585-w" volume="71" year="2019">
<normalizedToken id="2C425853FB92CD95260B9424F6082B59" originalValue="Santamaría-Aguilar">Santamaria-Aguilar</normalizedToken>
et al. (2019)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B196F30791DCD6319E1D6AF42063AD77" pageId="0" pageNumber="147" type="notes">
<paragraph id="84A1974D3B8BB05D260A7C08B8B4EBCD" pageId="0" pageNumber="147">Notes.</paragraph>
<paragraph id="96478D693078D5CB12279600B5E2F978" pageId="0" pageNumber="147">
The type specimen of 
<taxonomicName id="7C1967106E199C76D41C76B4BCA5F529" class="Magnoliopsida" family="Myristicaceae" genus="Otoba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Otoba scottmorii" order="Magnoliales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="scottmorii">
<emphasis id="37BAB63D0F86B3231509C9F2D8FCCD6C" italics="true" pageId="0" pageNumber="147">Otoba scottmorii</emphasis>
</taxonomicName>
and other collections here mentioned have been previously identified in herbaria and cited in literature as 
<taxonomicName id="C50DAE8E71F28A48213D4FB987E9842B" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. acuminata" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="acuminata">
<emphasis id="8EA4259D2FD9A8E3F8DDB9AE195263C1" italics="true" pageId="0" pageNumber="147">O. acuminata</emphasis>
</taxonomicName>
and 
<taxonomicName id="0C4FD8CB349EC75AE0E4A2FC84526452" class="Magnoliopsida" family="Myristicaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="O. gracilipes" order="Rosales" pageId="0" pageNumber="147" phylum="Magnoliophyta" rank="species" species="gracilipes">
<emphasis id="383C7A68BD73C53EFD1E529CF3FFD3FA" italics="true" pageId="0" pageNumber="147">O. gracilipes</emphasis>
</taxonomicName>
, and duplicates may have been distributed under these names.
</paragraph>
</subSubSection>
<subSubSection id="185A7A7330FA081065460B6FB00BC884" pageId="0" pageNumber="147" type="materials_examined">
<paragraph id="BDC9FB58057F3B4C26416AEF9559A8B4" pageId="0" pageNumber="147">Specimens examined.</paragraph>
<paragraph id="0B39496DFD4B0A607F93FF4A10428F97" pageId="0" pageNumber="147">
<materialsCitation id="BBE908D7059680C4B82C38CE82E7FF3B" collectingDate="1986-03-13" collectingDateMax="1993-11-29" collectingDateMin="1986-03-13" collectorName="R. Callejas, J. Brand, M. Narvaez, Paso de la Reina, E. Renteria, de Puerto Valdivia" country="Colombia" county="Mutata" elevation="55" latitude="7.3333335" location="Rio Surambay" longLatPrecision="1303" longitude="-75.333336" municipality="Sitio Rio Surambay" specimenCount="♂" stateProvince="Antioquia">
<emphasis id="568E052F8464D1081F4B36DB436B36F5" bold="true" pageId="0" pageNumber="147">
<collectingCountry id="141D7BABB8D482B839312360A13BA0B2" name="Colombia">Colombia</collectingCountry>
. 
<collectingRegion id="95EE73CF3DFA63D9E2267AFF87F1C118" country="Colombia" name="Antioquia">Antioquia</collectingRegion>
</emphasis>
: 
<normalizedToken id="D6E0A25CB26FCFA3E9B1C026AE26FF56" originalValue="Mutatá">
<collectingCounty id="A07895B4FCADA3AA3BF1111B6D84E034">Mutata</collectingCounty>
</normalizedToken>
, 
<collectingMunicipality id="C3FC6382D4E7E956004C2BA7F3EC76E6">
Sitio 
<location id="039092545FEF28E5F72909C6DAEFD81D" LSID="urn:lsid:plazi:treatment:B510B805C1405B1081A1B04ACA258316:039092545FEF28E5F72909C6DAEFD81D" country="Colombia" county="Mutata" latitude="7.3333335" longLatPrecision="1303" longitude="-75.333336" municipality="Sitio Rio Surambay" name="Rio Surambay" stateProvince="Antioquia">Rio Surambay</location>
</collectingMunicipality>
, 
<locationDeviation id="975BDECF2F0F6AD63BF421308432B26A" location="Mutata">
<quantity id="285078040A76480357A19EA0765E2615" metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
N de Mutata
</locationDeviation>
, 
<geoCoordinate id="249FEEECBFC33C45BBFC17E210F611B2" degrees="07" direction="north" minutes="20" orientation="latitude" precision="925" value="7.3333335">07°20'N</geoCoordinate>
, 
<geoCoordinate id="446D68F071B6894B00A547E4FD302CD6" degrees="076" direction="west" minutes="30" orientation="longitude" precision="925" value="-76.5">076°30'W</geoCoordinate>
, 
<quantity id="185D67C7A3DFBC16B5363FDE47224822" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" unit="m" value="55.0" valueMax="80.0" valueMin="30.0">
<elevation id="AE14A5FA111B946978371F3C58BDA8BE" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" unit="m" value="55.0" valueMax="80.0" valueMin="30.0">30-80 m</elevation>
</quantity>
, 
<collectingDate id="C1A58F45B972353B72B07D7CB6CD94B2" value="1987-11-21">21 Nov. 1987</collectingDate>
(
<specimenCount id="0F846C73A45CEC03B7397050DEB6ED30">♂</specimenCount>
fl), 
<collectorName id="3F5A5F528EB2B119AF2631687D022AF7">R. Callejas</collectorName>
et al. 5752 (INPA digital image, MO, NY); ibid, 
<collectingDate id="A0E6D2C29083EB836FD7C56002D40DFF" value="1987-11-21">21 Nov. 1987</collectingDate>
(fr), 
<collectorName id="A3076D15576873EC42C9D9FC679FD5F7">R. Callejas</collectorName>
et al. 5789 (INPA digital image, MO, NY); Turbo, 
<location id="19881B98367AD6E1FB85BED4879540E9" LSID="urn:lsid:plazi:treatment:B510B805C1405B1081A1B04ACA258316:19881B98367AD6E1FB85BED4879540E9" country="Colombia" county="Mutata" latitude="7.3333335" longLatPrecision="1303" longitude="-75.333336" municipality="Sitio Rio Surambay" name="Carretera Tapon del Darien" stateProvince="Antioquia">
Carretera 
<normalizedToken id="A251F07DED40191A1F3B4AA5C18F2192" originalValue="Tapón">Tapon</normalizedToken>
del 
<normalizedToken id="58586BF381DAF857EC61B7F322D5EAEB" originalValue="Darién">Darien</normalizedToken>
</location>
, Sector 
<location id="94BD3267E95AB548291DBA4DBE2EEDDC" LSID="urn:lsid:plazi:treatment:B510B805C1405B1081A1B04ACA258316:94BD3267E95AB548291DBA4DBE2EEDDC" country="Colombia" county="Mutata" latitude="7.3333335" longLatPrecision="1303" longitude="-75.333336" municipality="Sitio Rio Surambay" name="Rio Leon-Lomas Aisladas" stateProvince="Antioquia">
<normalizedToken id="26D8F8D8EA3932F48F65ECC9A4062F7D" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="8F654457AB81005F353AC869509636F1" originalValue="León-Lomas">Leon-Lomas</normalizedToken>
Aisladas
</location>
, km 37, [
<geoCoordinate id="E4DF18A5906FDA7E0F42EB5AB8F75D32" degrees="07" direction="north" minutes="39" orientation="latitude" precision="15" seconds="11" value="7.6530557">07°39'11&quot;N</geoCoordinate>
, 
<geoCoordinate id="F64CEA04C84741CE8411FFFB626BA7AD" degrees="076" direction="west" minutes="58" orientation="longitude" precision="15" seconds="02" value="-76.967224">076°58'02&quot;W</geoCoordinate>
], 
<quantity id="F7D731CD487354CA4A9A3B940A34CB4E" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<elevation id="BC23AA755192611140B4FBDD52A13694" metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</elevation>
</quantity>
, 
<collectingDate id="5C55951B6444D81D0989D06BD70B3C58" value="1993-11-29">29 Nov. 1993</collectingDate>
(fr), 
<collectorName id="B2EB141D4D091A6200E80679FC2A1DED">J. Brand</collectorName>
&amp; 
<collectorName id="975D87CB4D3724F5E07B40FD4EDE076E">
M. 
<normalizedToken id="25909BAB3F5EE2B3528879D76F16A358" originalValue="Narváez">Narvaez</normalizedToken>
</collectorName>
682 (COL [n.v.], JAUM digital image, MO); 
<location id="943D947A7CDC49FA383BB8C400102D3E" LSID="urn:lsid:plazi:treatment:B510B805C1405B1081A1B04ACA258316:943D947A7CDC49FA383BB8C400102D3E" country="Colombia" county="Mutata" latitude="7.3333335" longLatPrecision="1303" longitude="-75.333336" municipality="Sitio Rio Surambay" name="rio Cianura" stateProvince="Antioquia">
<normalizedToken id="EED42FDC1FA6163940BD57B9B59DDBA5" originalValue="río">rio</normalizedToken>
<normalizedToken id="D5804B44FC3E33492289B77DE8256603" originalValue="Cianurá">Cianura</normalizedToken>
</location>
, 
<collectorName id="8028457527AD2DB6E35A29FED5F766DC">Paso de la Reina</collectorName>
, 
<quantity id="0A4A5ED13B3C14D91988E17F3850C19A" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">
<elevation id="FB1C3A9F1E0174CB5F25ABB87F38034C" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">730 m</elevation>
</quantity>
, 
<collectingDate id="90131EF495BA84E06B565B8CF39A70C2" value="1986-03-13">13 Mar 1986</collectingDate>
(fr), 
<collectorName id="35A361474B38D231331A7A1D319E5024">E. Renteria</collectorName>
et al. 4680 (JAUM digital image); 
<normalizedToken id="C606C73C53A6A95B5D5C5DD40C3C1D9A" originalValue="Tarazá">Taraza</normalizedToken>
, Corregimiento El 12, 210 kms, 
<locationDeviation id="7C4F36E55BF84CF4D5E9512D1AD7E341" location="Medellin">
NE de 
<normalizedToken id="BC8EC1899EA5ED9347DCA7A999D3568F" originalValue="Medellín">Medellin</normalizedToken>
</locationDeviation>
, 
<normalizedToken id="3EB22A6C00069213076878259502D23C" originalValue="vía">via</normalizedToken>
El 12-Barroblanco, km 3, 
<geoCoordinate id="3A648B1BFF49E3ABD64A748E6C0C473E" degrees="07" direction="north" minutes="30" orientation="latitude" precision="925" value="7.5">07°30'N</geoCoordinate>
, 
<geoCoordinate id="66DA86755B706E1CBF18BA2CBDA64204" degrees="075" direction="west" minutes="20" orientation="longitude" precision="925" value="-75.333336">075°20'W</geoCoordinate>
, [
<quantity id="BDB61E630F1137E477B19415784C2B42" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">
<elevation id="A06317E2FD6FA1FAD2D251FE7B60E0A2" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</elevation>
</quantity>
], 
<collectingDate id="C978F23B13766486320E832F96E0905D" value="1987-11-09">09 Nov. 1987</collectingDate>
(
<specimenCount id="6E85FF1EAB9A9DC0CAC51F9EA5BD291F">♂</specimenCount>
fl), 
<collectorName id="B9C0E48BA0674CF38CBFCDBC628BC981">R. Callejas</collectorName>
et al. 5509 (INPA digital image, NY); Valdivia, corregimiento Puerto Valdivia, km 5 
<collectorName id="EB56AF5583148F3D8F9A74081FA0B28E">de Puerto Valdivia</collectorName>
hacia &quot;El 12&quot;, colecciones a lo largo del 
<location id="9841DC6774828FB3F0BC432302D5B4DB" LSID="urn:lsid:plazi:treatment:B510B805C1405B1081A1B04ACA258316:9841DC6774828FB3F0BC432302D5B4DB" country="Colombia" county="Mutata" latitude="7.3333335" longLatPrecision="1303" longitude="-75.333336" municipality="Sitio Rio Surambay" name="Rio Pescado" stateProvince="Antioquia">
<normalizedToken id="54E752D2775CB016EA0CADC3F66AE923" originalValue="Río">Rio</normalizedToken>
Pescado
</location>
, 
<geoCoordinate id="C9902932AAA9CA39E9F02E93A91AA929" degrees="07" direction="north" minutes="20" orientation="latitude" precision="925" value="7.3333335">07°20'N</geoCoordinate>
, 
<geoCoordinate id="0298A0E68FC262AA44AD3342F5229EE6" degrees="075" direction="west" minutes="20" orientation="longitude" precision="925" value="-75.333336">075°20'W</geoCoordinate>
, 
<quantity id="B7F348AC0D9F4213ECBB2327C3F3FC8A" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">
<elevation id="9FDA7276745C95DEA566F601FB67DDD2" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</elevation>
</quantity>
, 
<collectingDate id="FC420BCEB5D687E959CE6FF0A69018F3" value="1987-05-14">14 May. 1987</collectingDate>
(
<specimenCount id="7369079FA8B1858970EB17485689A9C7">♂</specimenCount>
fl), 
<collectorName id="6420B9101C0F56EEEE0338810EF1F022">R. Callejas</collectorName>
et al. 3440 (MO-2 sheets, NY)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>