408 lines
46 KiB
XML
408 lines
46 KiB
XML
<document id="5EF945053848D022394F260052280141" ID-DOI="10.11646/zootaxa.5541.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14241026" ID-ZooBank="0E636780-A497-4048-9C6E-7D985BFCBA5A" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1732835163561" checkinUser="plazi" docAuthor="Li, Hua-Zhao, Liu, Zhe & Li, Jia-Ling" docDate="2024" docId="B40287DDFFFFFFF7A1BDF9F7FC8D79B6" docLanguage="en" docName="zootaxa.5541.3.6.pdf" docOrigin="Zootaxa 5541 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5541.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coladenia agni subsp. huanghaoi H. Z. Li & Z. Liu 2024, ssp. nov." docType="treatment" docVersion="3" lastPageNumber="362" masterDocId="483BFFA5FFFDFFF2A12AFFADFF9B7D6A" masterDocTitle="Reexamination of the taxonomic status of Coladenia agni (de Nicéville, [1884]) from Hainan, China reveals a new subspecies (Lepidoptera, Hesperiidae)" masterLastPageNumber="366" masterPageNumber="357" pageNumber="359" updateTime="1733426059084" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="6F6E69E72A5D2D1727158C151C0CED40">Reexamination of the taxonomic status of Coladenia agni (de Nicéville, [1884]) from Hainan, China reveals a new subspecies (Lepidoptera, Hesperiidae)</mods:title>
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<mods:namePart id="6220BF013E34E09B79D7A3C58DA98A44">Li, Hua-Zhao</mods:namePart>
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<mods:affiliation id="8717DCD0032BA576C8F46D543DEB833D">The People’s Government of Dusong Township, Jinchuan County, Aba, Sichuan, P. R. China</mods:affiliation>
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<mods:namePart id="FB0E53341F666F9CA2F51CDF9E3F6321">Liu, Zhe</mods:namePart>
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<mods:affiliation id="E698EC05E6E770BE8D50B77C2D74CC0A">Nanfanghangkongcheng, Sanya, Hainan, P. R. China.</mods:affiliation>
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<mods:namePart id="42DB2E654D1056520D5F7E3D27D01A0A">Li, Jia-Ling</mods:namePart>
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<mods:nameIdentifier id="C01C5702C5F2B9407A1D2ABC7CF9E286" type="email">19071300110003@hainanu.edu.cn</mods:nameIdentifier>
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<mods:title id="C719A9F6EB57FF9D1E2406968802368A">Zootaxa</mods:title>
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<mods:date id="DA9D561F80AF6C55EF29B787FC02EE68">2024</mods:date>
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<mods:number id="3EEE81BCAE438FBFBB1813CC9F08A7CD">2024-11-27</mods:number>
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<heading id="675C81A7FFFFFFF0A1BDF9F7FA937B1E" bold="true" box="[151,1288,1626,1653]" fontSize="11" level="1" pageId="2" pageNumber="359" reason="1">
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<emphasis id="0EDFEAD9FFFFFFF0A1BDF9F7FA937B1E" bold="true" box="[151,1288,1626,1653]" pageId="2" pageNumber="359">
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<taxonomicName id="FBAB4D48FFFFFFF0A1BDF9F7FD2F7B1E" authority="H.Z. Li & Z. Liu" authorityName="H. Z. Li & Z. Liu" authorityYear="2024" box="[151,692,1626,1653]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="subSpecies" species="agni" status="subsp. nov." subSpecies="huanghaoi">
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<emphasis id="0EDFEAD9FFFFFFF0A1BDF9F7FE477B1F" bold="true" box="[151,476,1626,1653]" italics="true" pageId="2" pageNumber="359">Coladenia agni huanghaoi</emphasis>
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H.Z. Li & Z. Liu
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<taxonomicNameLabel id="15EC57A2FFFFFFF0A396F9F6FCBE7B1F" box="[700,805,1627,1653]" pageId="2" pageNumber="359" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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<paragraph id="3C1436CBFFFFFFF0A206F9F7FA937B1E" blockId="2.[151,1288,1626,1725]" box="[812,1288,1626,1653]" pageId="2" pageNumber="359">(Chinese name: DZŧsṳxã南亚ª)</paragraph>
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</subSubSection>
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</emphasis>
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</heading>
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<subSubSection id="74B16540FFFFFFF0A1BDF9D3FCC57BD7" pageId="2" pageNumber="359" type="description">
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<uri id="483A3AC9FFFFFFF0A1BDF9D3FC5E7BF3" box="[151,965,1662,1689]" pageId="2" pageNumber="359">
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urn:lsid:zoobank.org:act:
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<paragraph id="3C1436CBFFFFFFF0A1BDF90EFCC57BD7" blockId="2.[151,1288,1626,1725]" box="[151,862,1698,1725]" pageId="2" pageNumber="359">
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(
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<figureCitation id="A4902A4EFFFFFFF0A1B5F90EFF767BD7" box="[159,237,1699,1725]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[161,1433,189,1520]" captionTargetId="figure-19@3.[151,1436,179,1521]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Coladenia agni and C. agnioides from China and adjacent areas." figureDoi="http://doi.org/10.5281/zenodo.14241028" httpUri="https://zenodo.org/record/14241028/files/figure.png" pageId="2" pageNumber="359">Figs. 1</figureCitation>
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,
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<figureCitation id="A4902A4EFFFFFFF0A1D1F90EFE907BD7" box="[251,267,1699,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,714,738]" captionTargetBox="[155,1430,415,687]" captionTargetId="figure-131@4.[151,1435,414,688]" captionTargetPageId="4" captionText="FIGURE 2. Magnified scales of some specialized structures.A: underside of the palpi, P1+2 = the first two segments of palpi, P3 = the third segment of palpi, P i = inside of P1+2, Po = outside of P1+2. B: inside of the hind tibiae and end of the abdomen. Scale for Fig. 2B." figureDoi="http://doi.org/10.5281/zenodo.14241030" httpUri="https://zenodo.org/record/14241030/files/figure.png" pageId="2" pageNumber="359">2</figureCitation>
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,
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<figureCitation id="A4902A4EFFFFFFF0A03DF90FFEBE7BD7" box="[279,293,1698,1725]" captionStart="FIGURE 5" captionStartId="5.[151,250,1875,1899]" captionTargetBox="[151,1436,1266,1850]" captionTargetId="figure-298@5.[151,1436,1266,1850]" captionTargetPageId="5" captionText="FIGURE 5. Coladenia agni huanghaoi ssp. nov. in nature and its type locality." figureDoi="http://doi.org/10.5281/zenodo.14241036" httpUri="https://zenodo.org/record/14241036/files/figure.png" pageId="2" pageNumber="359">5</figureCitation>
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for adults, 3 for the genitalia, 7 for the phylogeny)
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</paragraph>
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<subSubSection id="74B16540FFFFFFF0A1BDF948FE727A74" pageId="2" pageNumber="359" type="reference_group">
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<taxonomicName id="FBAB4D48FFFFFFF0A1BDF948FEDC7B95" ID-CoL="WWYS" baseAuthorityName="de Niceville" baseAuthorityYear="1884" box="[151,327,1765,1791]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="agni">
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<emphasis id="0EDFEAD9FFFFFFF0A1BDF948FEDC7B95" box="[151,327,1765,1791]" italics="true" pageId="2" pageNumber="359">Coladenia agni</emphasis>
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ssp.:
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<treatmentCitation id="BD0A10DAFFFFFFF0A0A7F94BFDFC7A6A" author="Huang, H." box="[397,615,1765,1792]" page="574" pageId="2" pageNumber="359" year="2021">
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<bibRefCitation id="583A4B3AFFFFFFF0A0A7F94BFDB47B95" author="Huang, H." box="[397,559,1765,1792]" pageId="2" pageNumber="359" pagination="569 - 620" refId="ref4940" refString="Huang, H. (2021) Taxonomy and morphology of Chinese butterflies 1 Hesperiidae: Pyrginae: Genera Coladenia Moore, [1881] and Pseudocoladenia Shirozu & Saigusa, 1962. Atalanta, 52 (4), 569 - 620." type="journal article" year="2021">Huang (2021)</bibRefCitation>
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: 574
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||
</treatmentCitation>
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for taxonomic notes + 587, fig. 2 for
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+ 594, 597–599, 602, figs. 34, 46–48, 51 for
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genitalia.
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</paragraph>
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<materialsCitation id="8CC33C96FFFFFFF0A1BDF8E1FA8A7A0C" box="[151,1297,1868,1895]" collectingDate="2023-10-31" collectionCode="CMNH" collectorName="Z. Liu" country="China" location="Jianfengling" pageId="2" pageNumber="359" specimenCode="CH1" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
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<typeStatus id="E3108869FFFFFFF0A1BDF8E1FE937A0C" box="[151,264,1868,1894]" pageId="2" pageNumber="359" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount id="2AADFD42FFFFFFF0A03EF8E0FEB07A0D" box="[276,299,1869,1895]" count="1" pageId="2" pageNumber="359" type="male">♂</specimenCount>
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,
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<location id="39746010FFFFFFF0A01DF8E1FE5F7A0C" LSID="urn:lsid:plazi:treatment:B40287DDFFFFFFF7A1BDF9F7FC8D79B6:39746010FFFFFFF0A01DF8E1FE5F7A0C" box="[311,452,1868,1894]" country="China" name="Jianfengling" pageId="2" pageNumber="359" stateProvince="Hainan">Jianfengling</location>
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,
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<collectingRegion id="FE6FF829FFFFFFF0A0FAF8E1FDBF7A0C" box="[464,548,1868,1894]" country="China" name="Hainan" pageId="2" pageNumber="359">Hainan</collectingRegion>
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,
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<collectingCountry id="44BC765BFFFFFFF0A305F8E1FDEE7A0C" box="[559,629,1868,1894]" name="China" pageId="2" pageNumber="359">China</collectingCountry>
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,
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<date id="4815100BFFFFFFF0A3AAF8E1FD627A0C" box="[640,761,1868,1894]" pageId="2" pageNumber="359" value="2023-10-31">
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<collectingDate id="5851E9E3FFFFFFF0A3AAF8E1FD627A0C" box="[640,761,1868,1894]" pageId="2" pageNumber="359" value="2023-10-31">31.X.2023</collectingDate>
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</date>
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,
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<collectorName id="915E531DFFFFFFF0A22EF8E0FCD17A0C" box="[772,842,1868,1895]" pageId="2" pageNumber="359">Z. Liu</collectorName>
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leg. (
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<specimenCode id="6C0D9EB0FFFFFFF0A2A0F8E1FC597A0C" box="[906,962,1868,1894]" pageId="2" pageNumber="359">CH1</specimenCode>
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; will be deposited at
|
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<collectionCode id="5ABAAE0EFFFFFFF0A59AF8E1FA927A0C" box="[1200,1289,1868,1894]" collectionName="USA, Pennsylvania, Pittsburgh, Carnegie Museum of Natural History" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="2" pageNumber="359" type="Museum">CMNH</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="8CC33C96FFFFFFF0A431F8E1FB957AE0" collectingDate="2023-11-16" collectingDateMax="2023-11-21" collectingDateMin="2023-11-16" collectionCode="CZL" collectorName="Z. Liu" country="China" location="Jianfengling" pageId="2" pageNumber="359" specimenCode="CH2, CH4" specimenCount="3" specimenCount-male="3" stateProvince="Hainan" typeStatus="paratype">
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<emphasis id="0EDFEAD9FFFFFFF0A431F8E1FA077A0C" bold="true" box="[1307,1436,1868,1894]" pageId="2" pageNumber="359">
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<typeStatus id="E3108869FFFFFFF0A431F8E1FA0C7A0C" box="[1307,1431,1868,1894]" pageId="2" pageNumber="359" type="paratype">Paratypes</typeStatus>
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:
|
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</emphasis>
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<specimenCount id="2AADFD42FFFFFFF0A1BDF8DDFF427AE1" box="[151,217,1904,1931]" count="3" pageId="2" pageNumber="359" type="male">3 ♂♂</specimenCount>
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, locality same as the HT,
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<date id="4815100BFFFFFFF0A32BF8DDFD367AE0" box="[513,685,1904,1930]" pageId="2" pageNumber="359" value="2023-11-16" valueMax="2023-11-21" valueMin="2023-11-16">
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<collectingDate id="5851E9E3FFFFFFF0A32BF8DDFD367AE0" box="[513,685,1904,1930]" pageId="2" pageNumber="359" value="2023-11-16" valueMax="2023-11-21" valueMin="2023-11-16">16–21.XI.2023</collectingDate>
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</date>
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,
|
||
<collectorName id="915E531DFFFFFFF0A390F8DCFC9A7AE0" box="[698,769,1904,1931]" pageId="2" pageNumber="359">Z. Liu</collectorName>
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leg. (
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<specimenCode id="6C0D9EB0FFFFFFF0A269F8DDFCE17AE0" box="[835,890,1904,1930]" pageId="2" pageNumber="359">CH2</specimenCode>
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–
|
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<specimenCode id="6C0D9EB0FFFFFFF0A2A3F8DDFC5B7AE0" box="[905,960,1904,1930]" pageId="2" pageNumber="359">CH4</specimenCode>
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;
|
||
<collectionCode id="5ABAAE0EFFFFFFF0A2E6F8DDFB9D7AE0" box="[972,1030,1904,1930]" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="2" pageNumber="359">CZL</collectionCode>
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)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="8CC33C96FFFFFFF0A531F8DDFCB17AC4" collectingDateMax="2023-06-28" collectingDateMin="2023-09-18" collectionCode="CZL" collectorName="Z. Liu" country="China" location="Mt. Wuzhi" pageId="2" pageNumber="359" specimenCode="CH5, CH6" specimenCount="2" specimenCount-female="2" stateProvince="Hainan" typeStatus="paratype">
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<specimenCount id="2AADFD42FFFFFFF0A531F8DDFBC07AE1" box="[1051,1115,1904,1931]" count="2" pageId="2" pageNumber="359" type="female">2 ♀♀</specimenCount>
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,
|
||
<location id="39746010FFFFFFF0A540F8DCFB7E7AE0" LSID="urn:lsid:plazi:treatment:B40287DDFFFFFFF7A1BDF9F7FC8D79B6:39746010FFFFFFF0A540F8DCFB7E7AE0" box="[1130,1253,1904,1931]" country="China" name="Mt. Wuzhi" pageId="2" pageNumber="359" stateProvince="Hainan">Mt. Wuzhi</location>
|
||
,
|
||
<collectingRegion id="FE6FF829FFFFFFF0A5D8F8DDFADD7AE0" box="[1266,1350,1904,1930]" country="China" name="Hainan" pageId="2" pageNumber="359">Hainan</collectingRegion>
|
||
,
|
||
<collectingCountry id="44BC765BFFFFFFF0A478F8DDFA037AE0" box="[1362,1432,1904,1930]" name="China" pageId="2" pageNumber="359">China</collectingCountry>
|
||
,
|
||
<date id="4815100BFFFFFFF0A1BDF839FE8D7AC4" box="[151,278,1940,1966]" pageId="2" pageNumber="359" value="2021-09-18">18.IX.2021</date>
|
||
and
|
||
<date id="4815100BFFFFFFF0A067F839FE547AC4" box="[333,463,1940,1966]" pageId="2" pageNumber="359" value="2023-06-28">28.VI.2023</date>
|
||
,
|
||
<collectorName id="915E531DFFFFFFF0A0F1F838FDBA7AC4" box="[475,545,1940,1967]" pageId="2" pageNumber="359">Z. Liu</collectorName>
|
||
leg. (
|
||
<specimenCode id="6C0D9EB0FFFFFFF0A34AF839FD0D7AC5" box="[608,662,1940,1967]" pageId="2" pageNumber="359">CH5</specimenCode>
|
||
–
|
||
<specimenCode id="6C0D9EB0FFFFFFF0A38CF839FD477AC5" box="[678,732,1940,1967]" pageId="2" pageNumber="359">CH6</specimenCode>
|
||
;
|
||
<collectionCode id="5ABAAE0EFFFFFFF0A3C2F839FCB97AC4" box="[744,802,1940,1966]" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="2" pageNumber="359">CZL</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="68D46643FFFEFFF1A1BDF9A6FB817B49" ID-DOI="http://doi.org/10.5281/zenodo.14241028" ID-Zenodo-Dep="14241028" box="[151,1050,1547,1571]" httpUri="https://zenodo.org/record/14241028/files/figure.png" pageId="3" pageNumber="360" startId="3.[151,250,1547,1571]" targetBox="[161,1433,189,1520]" targetPageId="3" targetType="figure">
|
||
<paragraph id="3C1436CBFFFEFFF1A1BDF9A6FB817B49" blockId="3.[151,1050,1547,1571]" box="[151,1050,1547,1571]" pageId="3" pageNumber="360">
|
||
<emphasis id="0EDFEAD9FFFEFFF1A1BDF9A6FE887B49" bold="true" box="[151,275,1547,1571]" pageId="3" pageNumber="360">FIGURE 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="FBAB4D48FFFEFFF1A0A0F9A6FDBC7B49" baseAuthorityName="de Niceville" baseAuthorityYear="1884" box="[394,551,1547,1571]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="agni">
|
||
<emphasis id="0EDFEAD9FFFEFFF1A0A0F9A6FDBC7B49" box="[394,551,1547,1571]" italics="true" pageId="3" pageNumber="360">Coladenia agni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FBAB4D48FFFEFFF1A372F9A1FD427B49" authorityName="Elwes & Edwards" authorityYear="1897" box="[600,729,1547,1571]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="agnioides">
|
||
<emphasis id="0EDFEAD9FFFEFFF1A372F9A1FD427B49" box="[600,729,1547,1571]" italics="true" pageId="3" pageNumber="360">C. agnioides</emphasis>
|
||
</taxonomicName>
|
||
from China and adjacent areas.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="74B16540FFFEFFF6A1EDF9D8FABE7CE3" lastPageId="4" lastPageNumber="361" pageId="3" pageNumber="360" type="description">
|
||
<paragraph id="3C1436CBFFFEFFF6A1EDF9D8FABE7CE3" blockId="3.[151,1437,1653,2040]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="361" pageId="3" pageNumber="360">
|
||
<emphasis id="0EDFEAD9FFFEFFF1A1EDF9D8FE3D7BFA" bold="true" box="[199,422,1653,1680]" pageId="3" pageNumber="360">Description. Male.</emphasis>
|
||
Length of forewing
|
||
<quantity id="FB539B2EFFFEFFF1A3BCF9D8FCCC7BFA" box="[662,855,1653,1680]" metricMagnitude="-2" metricUnit="m" metricValue="1.989" metricValueMax="2.055" metricValueMin="1.923" pageId="3" pageNumber="360" unit="mm" value="19.89" valueMax="20.55" valueMin="19.23">19.23–20.55 mm</quantity>
|
||
(
|
||
<quantity id="FB539B2EFFFEFFF1A242F9D8FC457BFA" box="[872,990,1653,1680]" metricMagnitude="-2" metricUnit="m" metricValue="2.055" pageId="3" pageNumber="360" unit="mm" value="20.55">20.55 mm</quantity>
|
||
in HT, n=4). Wings ground color dark brown. Forewing upperside: white hyaline spots situated in postdiscal area of spaces 1b to 3, subapical areas of space 6 to 8, subcostal area of space 11, and end of cell, spots in space 2 and cell larger. Hindwing upperside: covered with dense ochreous hairs at base, a series of postdiscal black spots present in spaces 1b to 7 and end of cell. Wings underside: generally similar to upperside but forewing submarginal areas lighter. Fringes: dark brown but paler in forewing spaces 1b and 7, and hindwing spaces 6 to 7.
|
||
<emphasis id="0EDFEAD9FFFEFFF1A2ADF887FBDB7A29" bold="true" box="[903,1088,1833,1860]" pageId="3" pageNumber="360">Male Genitalia.</emphasis>
|
||
Vinculum: slender, fourfold as long as saccus. Saccus: from the part connected to vinculum, continuously narrowed forward, ending with a sharp point. Tegumen: dorsally triangle, caudal part protruding. Gnathos: bilateral parts elongated ventrally, integrating a gourd-shaped terminal covered with dense and small spines. Uncus: abruptly tapered then elongated into a blunt end, of even width, with a small dorsal process. Valva: more or less rectangular; ampulla elliptic, forming a slightly sharp end; harpe tapered and bends upward with lower edge less protruding and with a long and blunt distal apex. Juxta: bilateral parts expansive and gradually narrowed posteriorly. Phallus: long and curved upwards, a pair of carinae present in middle of dorsal part.
|
||
<emphasis id="0EDFEAD9FFF9FFF6A345FF16FD567DBF" bold="true" box="[623,717,187,213]" pageId="4" pageNumber="361">Female.</emphasis>
|
||
Length of forewing
|
||
<quantity id="FB539B2EFFF9FFF6A2EAFF17FB197DBF" box="[960,1154,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.1595" metricValueMax="2.165" metricValueMin="2.154" pageId="4" pageNumber="361" unit="mm" value="21.595" valueMax="21.65" valueMin="21.54">21.54–21.65 mm</quantity>
|
||
(n=2). Larger, generally similar to male, but forewing central spots larger.
|
||
<emphasis id="0EDFEAD9FFF9FFF6A3FDFF72FC367D92" bold="true" box="[727,941,222,249]" pageId="4" pageNumber="361">Female Genitalia.</emphasis>
|
||
Lamella: Lamella antevaginalis cordiform, posterior margin sharply concave, basal part elongated and deviated to the right; Lamella postvaginalis rectangle with bilateral parts robust, anteriorly forming lateral shoulders; Antrum joined and integrated with basal lamella. Ductus bursae and corpus bursae: long, gradually expanding forward without signa. Lodix: more or less square, cephalic part protruding, posterior margin slightly concave. Papilla analis: smooth without serrate ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="68D46643FFF9FFF6A1BDFD67FE957E40" ID-DOI="http://doi.org/10.5281/zenodo.14241030" ID-Zenodo-Dep="14241030" httpUri="https://zenodo.org/record/14241030/files/figure.png" pageId="4" pageNumber="361" startId="4.[151,250,714,738]" targetBox="[155,1430,415,687]" targetPageId="4" targetType="figure">
|
||
<paragraph id="3C1436CBFFF9FFF6A1BDFD67FE957E40" blockId="4.[151,1436,714,810]" pageId="4" pageNumber="361">
|
||
<emphasis id="0EDFEAD9FFF9FFF6A1BDFD67FE887F88" bold="true" box="[151,275,714,738]" pageId="4" pageNumber="361">FIGURE 2.</emphasis>
|
||
Magnified scales of some specialized structures. A: underside of the palpi, P
|
||
<subScript id="A02F348EFFF9FFF6A530FD77FBAA7F82" attach="left" box="[1050,1073,730,744]" fontSize="6" pageId="4" pageNumber="361">1+2</subScript>
|
||
= the first two segments of palpi, P
|
||
<subScript id="A02F348EFFF9FFF6A4BFFD77FA077F82" attach="left" box="[1429,1436,730,744]" fontSize="6" pageId="4" pageNumber="361">3</subScript>
|
||
= the third segment of palpi, P
|
||
<subScript id="A02F348EFFF9FFF6A0E7FD52FE4A7E67" attach="left" box="[461,465,767,781]" fontSize="6" pageId="4" pageNumber="361">i</subScript>
|
||
= inside of P
|
||
<subScript id="A02F348EFFF9FFF6A370FD53FDEA7E66" attach="left" box="[602,625,766,780]" fontSize="6" pageId="4" pageNumber="361">1+2</subScript>
|
||
, P
|
||
<subScript id="A02F348EFFF9FFF6A3A6FD53FD087E66" attach="left" box="[652,659,766,780]" fontSize="6" pageId="4" pageNumber="361">o</subScript>
|
||
= outside of P
|
||
<subScript id="A02F348EFFF9FFF6A203FD53FCDB7E66" attach="left" box="[809,832,766,780]" fontSize="6" pageId="4" pageNumber="361">1+2</subScript>
|
||
. B: inside of the hind tibiae and end of the abdomen. Scale for Fig. 2B.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="68D46643FFF9FFF6A1BDF983FC387B00" ID-DOI="http://doi.org/10.5281/zenodo.14241032" ID-Zenodo-Dep="14241032" httpUri="https://zenodo.org/record/14241032/files/figure.png" pageId="4" pageNumber="361" startId="4.[151,250,1582,1606]" targetBox="[154,1431,872,1550]" targetPageId="4" targetType="figure">
|
||
<paragraph id="3C1436CBFFF9FFF6A1BDF983FC387B00" blockId="4.[151,1436,1582,1642]" pageId="4" pageNumber="361">
|
||
<emphasis id="0EDFEAD9FFF9FFF6A1BDF983FE8E7B2C" bold="true" box="[151,277,1582,1606]" pageId="4" pageNumber="361">FIGURE 3.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="FBAB4D48FFF9FFF6A0B5F983FD297B2C" authorityName="Li & Liu & Li" authorityYear="2024" box="[415,690,1582,1606]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="agni" status="subsp. nov." subSpecies="huanghaoi">
|
||
<emphasis id="0EDFEAD9FFF9FFF6A0B5F983FD297B2C" box="[415,690,1582,1606]" italics="true" pageId="4" pageNumber="361">Coladenia agni huanghaoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0EDFEAD9FFF9FFF6A391F983FC8C7B2C" bold="true" box="[699,791,1582,1606]" pageId="4" pageNumber="361">
|
||
<taxonomicNameLabel id="15EC57A2FFF9FFF6A391F983FC8C7B2C" box="[699,791,1582,1606]" pageId="4" pageNumber="361" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A: Male genitalia. B: Female genitalia. la = lateral view, lo = oblique lateral view, ds = dorsal view, vs = ventral view, po = posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="74B16540FFF9FFF6A1EDF911FE717AB9" pageId="4" pageNumber="361" type="diagnosis">
|
||
<paragraph id="3C1436CBFFF9FFF6A1EDF911FA937BBC" blockId="4.[151,1436,1724,2003]" box="[199,1288,1724,1751]" pageId="4" pageNumber="361">
|
||
<emphasis id="0EDFEAD9FFF9FFF6A1EDF911FED97BBC" bold="true" box="[199,322,1724,1750]" pageId="4" pageNumber="361">Diagnosis.</emphasis>
|
||
This new subspecies can be distinguished from
|
||
<taxonomicName id="FBAB4D48FFF9FFF6A274F910FB9C7BBC" authorityName="de Jong & Treadaway" authorityYear="1992" box="[862,1031,1724,1751]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="sundae">
|
||
<emphasis id="0EDFEAD9FFF9FFF6A274F910FB9C7BBC" box="[862,1031,1724,1751]" italics="true" pageId="4" pageNumber="361">C. agni sundae</emphasis>
|
||
</taxonomicName>
|
||
[Malayan] by (
|
||
<figureCitation id="A4902A4EFFF9FFF6A59FF911FB617BBC" box="[1205,1274,1724,1750]" captionStart="FIGURE 4" captionStartId="5.[151,250,500,524]" captionTargetBox="[164,1424,186,469]" captionTargetId="figure-19@5.[158,1430,179,473]" captionTargetPageId="5" captionText="FIGURE 4. Genital diagnosis of Coladenia agni huanghaoi ssp. nov. and other subspecies." figureDoi="http://doi.org/10.5281/zenodo.14241034" httpUri="https://zenodo.org/record/14241034/files/figure.png" pageId="4" pageNumber="361">Fig. 4</figureCitation>
|
||
):
|
||
</paragraph>
|
||
<paragraph id="3C1436CBFFF9FFF6A1EDF94DFD027B90" blockId="4.[151,1436,1724,2003]" box="[199,665,1760,1787]" pageId="4" pageNumber="361">(1) Much thinner distal part of the harpe.</paragraph>
|
||
<paragraph id="3C1436CBFFF9FFF6A1EDF8A9FE817A28" blockId="4.[151,1436,1724,2003]" pageId="4" pageNumber="361">
|
||
Whereas, it can be separated from
|
||
<taxonomicName id="FBAB4D48FFF9FFF6A360F8A8FD697A74" authorityName="de Jong & Treadaway" authorityYear="1992" box="[586,754,1796,1823]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="sundae">
|
||
<emphasis id="0EDFEAD9FFF9FFF6A360F8A8FD697A74" box="[586,754,1796,1823]" italics="true" pageId="4" pageNumber="361">C. agni sundae</emphasis>
|
||
</taxonomicName>
|
||
[Indo-Chinese] and
|
||
<taxonomicName id="FBAB4D48FFF9FFF6A2F3F8A8FBFF7A74" baseAuthorityName="Semper" baseAuthorityYear="1892" box="[985,1124,1797,1823]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="agni">
|
||
<emphasis id="0EDFEAD9FFF9FFF6A2F3F8A8FBFF7A74" box="[985,1124,1797,1823]" italics="true" pageId="4" pageNumber="361">C. agni agni</emphasis>
|
||
</taxonomicName>
|
||
by the following characters (
|
||
<figureCitation id="A4902A4EFFF9FFF6A1B5F885FF767A28" box="[159,237,1832,1858]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[161,1433,189,1520]" captionTargetId="figure-19@3.[151,1436,179,1521]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Coladenia agni and C. agnioides from China and adjacent areas." figureDoi="http://doi.org/10.5281/zenodo.14241028" httpUri="https://zenodo.org/record/14241028/files/figure.png" pageId="4" pageNumber="361">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="A4902A4EFFF9FFF6A1D1F885FE977A28" box="[251,268,1832,1858]" captionStart="FIGURE 4" captionStartId="5.[151,250,500,524]" captionTargetBox="[164,1424,186,469]" captionTargetId="figure-19@5.[158,1430,179,473]" captionTargetPageId="5" captionText="FIGURE 4. Genital diagnosis of Coladenia agni huanghaoi ssp. nov. and other subspecies." figureDoi="http://doi.org/10.5281/zenodo.14241034" httpUri="https://zenodo.org/record/14241034/files/figure.png" pageId="4" pageNumber="361">4</figureCitation>
|
||
):
|
||
</paragraph>
|
||
<paragraph id="3C1436CBFFF9FFF6A1EDF8E1FB057A0C" blockId="4.[151,1436,1724,2003]" box="[199,1182,1868,1895]" pageId="4" pageNumber="361">(2) Harpe is narrower with its outer edge less protruding and the tip longer and blunter.</paragraph>
|
||
<paragraph id="3C1436CBFFF9FFF6A1EDF8DDFB817AE0" blockId="4.[151,1436,1724,2003]" box="[199,1050,1904,1930]" pageId="4" pageNumber="361">(3) Phallus is strongly curved and slenderer, with carinae somewhat longer.</paragraph>
|
||
<paragraph id="3C1436CBFFF9FFF6A1EDF839FE717AB9" blockId="4.[151,1436,1724,2003]" pageId="4" pageNumber="361">
|
||
(4) Male forewing markings are similar to
|
||
<taxonomicName id="FBAB4D48FFF9FFF6A399F838FCAE7AC4" authorityName="de Jong & Treadaway" authorityYear="1992" box="[691,821,1940,1967]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="sundae">
|
||
ssp.
|
||
<emphasis id="0EDFEAD9FFF9FFF6A3CCF839FCAE7AC4" box="[742,821,1940,1966]" italics="true" pageId="4" pageNumber="361">sundae</emphasis>
|
||
</taxonomicName>
|
||
in having spots of the cell and spaces 2–3 thinner and more separated than
|
||
<taxonomicName id="FBAB4D48FFF9FFF6A0AAF814FE787AB8" baseAuthorityName="Semper" baseAuthorityYear="1892" box="[384,483,1977,2003]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="agni">
|
||
ssp.
|
||
<emphasis id="0EDFEAD9FFF9FFF6A09BF814FE787AB8" box="[433,483,1977,2002]" italics="true" pageId="4" pageNumber="361">agni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="68D46643FFF8FFF7A1BDFE59FBA77F66" ID-DOI="http://doi.org/10.5281/zenodo.14241034" ID-Zenodo-Dep="14241034" box="[151,1084,500,524]" httpUri="https://zenodo.org/record/14241034/files/figure.png" pageId="5" pageNumber="362" startId="5.[151,250,500,524]" targetBox="[164,1424,186,469]" targetPageId="5" targetType="figure">
|
||
<paragraph id="3C1436CBFFF8FFF7A1BDFE59FBA77F66" blockId="5.[151,1084,500,524]" box="[151,1084,500,524]" pageId="5" pageNumber="362">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1BDFE59FE887F66" bold="true" box="[151,275,500,524]" pageId="5" pageNumber="362">FIGURE 4.</emphasis>
|
||
Genital diagnosis of
|
||
<taxonomicName id="FBAB4D48FFF8FFF7A0C6FE59FD607F66" authorityName="Li & Liu & Li" authorityYear="2024" box="[492,763,500,524]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="362" phylum="Arthropoda" rank="subSpecies" species="agni" status="subsp. nov." subSpecies="huanghaoi">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A0C6FE59FD607F66" box="[492,763,500,524]" italics="true" pageId="5" pageNumber="362">Coladenia agni huanghaoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="0EDFEAD9FFF8FFF7A228FE59FCC77F66" bold="true" box="[770,860,500,524]" pageId="5" pageNumber="362">
|
||
<taxonomicNameLabel id="15EC57A2FFF8FFF7A228FE59FCC77F66" box="[770,860,500,524]" pageId="5" pageNumber="362" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and other subspecies.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="74B16540FFF8FFF7A1EDFDF3FCBB7FAA" pageId="5" pageNumber="362" type="description">
|
||
<paragraph id="3C1436CBFFF8FFF7A1EDFDF3FCBB7FAA" blockId="5.[151,1437,606,1245]" pageId="5" pageNumber="362">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1EDFDF3FEA47F12" bold="true" box="[199,319,606,632]" pageId="5" pageNumber="362">Variation.</emphasis>
|
||
The sizes of wing markings are slightly variable without taxonomic significance in differing
|
||
<taxonomicName id="FBAB4D48FFF8FFF7A45BFDF2FF7D7FF6" authorityName="de Jong & Treadaway" authorityYear="1992" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="362" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="sundae">
|
||
ssp.
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1BDFD2FFF7D7FF6" box="[151,230,642,668]" italics="true" pageId="5" pageNumber="362">sundae</emphasis>
|
||
</taxonomicName>
|
||
. Based on an individual with relatively small markings,
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A24EFD2EFB9A7FF6" author="Huang, H." box="[868,1025,642,669]" pageId="5" pageNumber="362" pagination="569 - 620" refId="ref4940" refString="Huang, H. (2021) Taxonomy and morphology of Chinese butterflies 1 Hesperiidae: Pyrginae: Genera Coladenia Moore, [1881] and Pseudocoladenia Shirozu & Saigusa, 1962. Atalanta, 52 (4), 569 - 620." type="journal article" year="2021">Huang (2021)</bibRefCitation>
|
||
considered the marking sizes of this new subspecies a diagnosis, which is denied here (
|
||
<figureCitation id="A4902A4EFFF8FFF7A3E7FD0BFC957FAA" box="[717,782,678,704]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[161,1433,189,1520]" captionTargetId="figure-19@3.[151,1436,179,1521]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Coladenia agni and C. agnioides from China and adjacent areas." figureDoi="http://doi.org/10.5281/zenodo.14241028" httpUri="https://zenodo.org/record/14241028/files/figure.png" pageId="5" pageNumber="362">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74B16540FFF8FFF7A1EDFD67FD217F8E" box="[199,698,714,741]" pageId="5" pageNumber="362" type="distribution">
|
||
<paragraph id="3C1436CBFFF8FFF7A1EDFD67FD217F8E" blockId="5.[151,1437,606,1245]" box="[199,698,714,741]" pageId="5" pageNumber="362">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1EDFD67FEFA7F8E" bold="true" box="[199,353,714,740]" pageId="5" pageNumber="362">Distribution.</emphasis>
|
||
Endemic to
|
||
<collectingRegion id="FE6FF829FFF8FFF7A0C5FD67FDDB7F8E" box="[495,576,714,740]" country="China" name="Hainan" pageId="5" pageNumber="362">Hainan</collectingRegion>
|
||
Is.,
|
||
<collectingCountry id="44BC765BFFF8FFF7A35AFD67FD2C7F8E" box="[624,695,714,740]" name="China" pageId="5" pageNumber="362">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74B16540FFF8FFF7A1EDFD43FC7F7E46" pageId="5" pageNumber="362" type="etymology">
|
||
<paragraph id="3C1436CBFFF8FFF7A1EDFD43FC7F7E46" blockId="5.[151,1437,606,1245]" pageId="5" pageNumber="362">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1EDFD43FED57E62" bold="true" box="[199,334,750,776]" pageId="5" pageNumber="362">Etymology.</emphasis>
|
||
The subspecific name is dedicated to Mr. Hao Huang, who first recognized the specificity of the
|
||
<collectingRegion id="FE6FF829FFF8FFF7A1BDFCBFFF737E46" box="[151,232,786,812]" country="China" name="Hainan" pageId="5" pageNumber="362">Hainan</collectingRegion>
|
||
population and frequently offered the first author taxonomical help.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74B16540FFF8FFF7A1EDFC9BFDA87E1E" pageId="5" pageNumber="362" type="biology_ecology">
|
||
<paragraph id="3C1436CBFFF8FFF7A1EDFC9BFDA87E1E" blockId="5.[151,1437,606,1245]" pageId="5" pageNumber="362">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1EDFC9BFED27E3A" bold="true" box="[199,329,822,848]" pageId="5" pageNumber="362">Phenology.</emphasis>
|
||
Multivoltine with at least two generations. The first brood is from April to June, and the second brood is from September to October.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="74B16540FFF8FFF7A1EDFCD3FC8D79B6" pageId="5" pageNumber="362" type="discussion">
|
||
<paragraph id="3C1436CBFFF8FFF7A1EDFCD3FC8D79B6" blockId="5.[151,1437,606,1245]" pageId="5" pageNumber="362">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1EDFCD3FEA07EF2" bold="true" box="[199,315,894,920]" pageId="5" pageNumber="362">Remarks.</emphasis>
|
||
(1) The records of this species from
|
||
<collectingRegion id="FE6FF829FFF8FFF7A3CFFCD3FCAD7EF2" box="[741,822,894,920]" country="China" name="Hainan" pageId="5" pageNumber="362">Hainan</collectingRegion>
|
||
in previous works, including
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A5A7FCD3FA0C7EF2" author="Joicey, J. J. & Talbot, G." box="[1165,1431,894,920]" pageId="5" pageNumber="362" pagination="257 - 262" refId="ref5024" refString="Joicey, J. J. & Talbot, G. (1932) A Catalogue of the Lepidoptera of Hainan. Bulletin of the Hill Museum, 4 (3), 257 - 262." type="journal article" year="1932">Joicey & Talbot (1932)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A1BDFC0EFEB57ED6" author="Evans, W. H." box="[151,302,930,957]" pageId="5" pageNumber="362" refId="ref4600" refString="Evans, W. H. (1949) A Catalogue of the Hesperiidae from Europe, Asia & Australia in the British Museum (Natural History). The British Museum, London, 502 pp. https: // doi. org / 10.5962 / bhl. title. 105941" type="book" year="1949">Evans (1949)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A012FC0FFDE17ED6" author="de Jong, R. & Treadaway, C. G." box="[312,634,930,957]" pageId="5" pageNumber="362" pagination="283 - 293" refId="ref4429" refString="de Jong, R. & Treadaway, C. G. (1992) Revisional notes on Coladenia Moore, 1881 (Lepidoptera: Hesperiidae). Zoologische Mededelingen, 66 (15), 283 - 293." type="journal article" year="1992">de Jong & Treadaway (1992)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A3AEFC0FFD637ED6" author="Gu, M. B." box="[644,760,930,956]" pageId="5" pageNumber="362" pagination="654 - 716" refId="ref4732" refString="Gu, M. B. (2002) Lepidoptera: Rhopalocera. In: Huang, F. S. (Ed.), Forest Insect of Hainan. Science Press, Beijing, pp. 654 - 716. [in Chinese]" type="book chapter" year="2002">Gu (2002)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A21AFC0FFC5E7ED6" author="Chiba, H." box="[816,965,930,956]" pageId="5" pageNumber="362" pagination="337 - 344" refId="ref4358" refString="Chiba, H. (2008) Skippers of Hainan (Lepidoptera: Hesperiidae). Report on Insect Inventory Project in Tropical Asia (TAIIV), 2008, 337 - 344." type="journal article" year="2008">Chiba (2008)</bibRefCitation>
|
||
, are difficult to verify since only checklists are available, and the photos of
|
||
<taxonomicName id="FBAB4D48FFF8FFF7A324FC6BFDF87E8A" baseAuthorityName="de Niceville" baseAuthorityYear="1884" box="[526,611,966,992]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="agni">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A324FC6BFDF87E8A" box="[526,611,966,992]" italics="true" pageId="5" pageNumber="362">C. agni</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A3A7FC6BFCF37E8A" author="Gu, M. B. & Chen, P. Z." box="[653,872,966,992]" pageId="5" pageNumber="362" refId="ref4697" refString="Gu, M. B. & Chen, P. Z. (1997) Butterflies in Hainan Island. China Forestry Publishing House, Beijing, 355 pp. [in Chinese]" type="book" year="1997">Gu & Chen (1997)</bibRefCitation>
|
||
are misidentifications of
|
||
<taxonomicName id="FBAB4D48FFF8FFF7A5B8FC6BFA027E8A" authority="(Chiba 2008)" baseAuthorityName="Chiba" baseAuthorityYear="2008" box="[1170,1433,966,992]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="laxmi">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A5B8FC6BFB6F7E8A" box="[1170,1268,966,992]" italics="true" pageId="5" pageNumber="362">C. laxmi</emphasis>
|
||
(
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A42FFC6BFA0A7E8A" author="Chiba, H." box="[1285,1425,966,992]" pageId="5" pageNumber="362" pagination="337 - 344" refId="ref4358" refString="Chiba, H. (2008) Skippers of Hainan (Lepidoptera: Hesperiidae). Report on Insect Inventory Project in Tropical Asia (TAIIV), 2008, 337 - 344." type="journal article" year="2008">Chiba 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Furthermore,
|
||
<taxonomicName id="FBAB4D48FFF8FFF7A019FC47FE58796E" authorityName="Elwes & Edwards" authorityYear="1897" box="[307,451,1002,1028]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="362" phylum="Arthropoda" rank="species" species="agnioides">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A019FC47FE58796E" box="[307,451,1002,1028]" italics="true" pageId="5" pageNumber="362">C. agnioides</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A0C1FC47FDE5796E" author="Chou, I." box="[491,638,1002,1028]" pageId="5" pageNumber="362" refId="ref4392" refString="Chou, I. (Ed.) (1994) Monographia Rhopalocerorum Sinensium 1 & 2. Henan Scientific and Technological Publishing House, Zhengzhou, 854 pp. [in Chinese with English abstract]" type="book" year="1994">Chou (1994)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A3A6FC47FCFC796E" author="Gu, M. B. & Chen, P. Z." box="[652,871,1002,1028]" pageId="5" pageNumber="362" refId="ref4697" refString="Gu, M. B. & Chen, P. Z. (1997) Butterflies in Hainan Island. China Forestry Publishing House, Beijing, 355 pp. [in Chinese]" type="book" year="1997">Gu & Chen (1997)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A28CFC46FB1A796E" author="Wang M. & Liu J. & Fu C. R." box="[934,1153,1002,1029]" pageId="5" pageNumber="362" refId="ref6063" refString="Wang M., Liu J. & Fu C. R. (2020) A Photographic Guide to the Butterflies of Yinggeling, Hainan. Nan Hai Publishing Company, Haikou, 369 pp. [in Chinese]" type="book" year="2020">
|
||
Wang
|
||
<emphasis id="0EDFEAD9FFF8FFF7A2DAFC46FBB0796E" box="[1008,1067,1002,1028]" italics="true" pageId="5" pageNumber="362">et al.</emphasis>
|
||
(2020)
|
||
</bibRefCitation>
|
||
, are not easily identified without examination of the genitalia, due to its sympatric distribution with and superficial resemblance to other
|
||
<taxonomicName id="FBAB4D48FFF8FFF7A1BDFB9FFE977926" authorityName="Moore" authorityYear="1881" box="[151,268,1074,1100]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="362" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0EDFEAD9FFF8FFF7A1BDFB9FFE977926" box="[151,268,1074,1100]" italics="true" pageId="5" pageNumber="362">Coladenia</emphasis>
|
||
</taxonomicName>
|
||
members. (2) The female individual from Nonggang, a reserve between Daxin and Longzhou, given in
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A19CFBFAFECF791A" author="Huang, H." box="[182,340,1110,1137]" pageId="5" pageNumber="362" pagination="569 - 620" refId="ref4940" refString="Huang, H. (2021) Taxonomy and morphology of Chinese butterflies 1 Hesperiidae: Pyrginae: Genera Coladenia Moore, [1881] and Pseudocoladenia Shirozu & Saigusa, 1962. Atalanta, 52 (4), 569 - 620." type="journal article" year="2021">Huang (2021)</bibRefCitation>
|
||
probably belongs to
|
||
<taxonomicName id="FBAB4D48FFF8FFF7A362FBFAFD50791A" authorityName="de Jong & Treadaway" authorityYear="1992" box="[584,715,1110,1137]" class="Insecta" family="Hesperiidae" genus="Coladenia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="362" phylum="Arthropoda" rank="subSpecies" species="agni" subSpecies="sundae">
|
||
ssp.
|
||
<emphasis id="0EDFEAD9FFF8FFF7A356FBFBFD50791A" box="[636,715,1110,1136]" italics="true" pageId="5" pageNumber="362">sundae</emphasis>
|
||
</taxonomicName>
|
||
[Indo-Chinese] rather than the nominotypical one. Meanwhile, it is noteworthy that the female genitalia of the new subspecies are not entirely consistent with those from the mainland, especially from
|
||
<collectingRegion id="FE6FF829FFF8FFF7A0E2FB32FDBA79D3" box="[456,545,1183,1209]" country="China" name="Yunnan" pageId="5" pageNumber="362">Yunnan</collectingRegion>
|
||
,
|
||
<collectingCountry id="44BC765BFFF8FFF7A304FB33FDEA79D2" box="[558,625,1182,1208]" name="China" pageId="5" pageNumber="362">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="583A4B3AFFF8FFF7A3A8FB33FC8A79D2" author="Huang, H." box="[642,785,1182,1208]" pageId="5" pageNumber="362" pagination="569 - 620" refId="ref4940" refString="Huang, H. (2021) Taxonomy and morphology of Chinese butterflies 1 Hesperiidae: Pyrginae: Genera Coladenia Moore, [1881] and Pseudocoladenia Shirozu & Saigusa, 1962. Atalanta, 52 (4), 569 - 620." type="journal article" year="2021">Huang 2021</bibRefCitation>
|
||
) (
|
||
<figureCitation id="A4902A4EFFF8FFF7A201FB33FCE979D2" box="[811,882,1182,1208]" captionStart="FIGURE 3" captionStartId="4.[151,250,1582,1606]" captionTargetBox="[154,1431,872,1550]" captionTargetId="figure-195@4.[151,1435,868,1556]" captionTargetPageId="4" captionText="FIGURE 3. Genitalia of Coladenia agni huanghaoi ssp. nov. A: Male genitalia. B: Female genitalia. la = lateral view, lo = oblique lateral view, ds = dorsal view, vs = ventral view, po = posterior view." figureDoi="http://doi.org/10.5281/zenodo.14241032" httpUri="https://zenodo.org/record/14241032/files/figure.png" pageId="5" pageNumber="362">Fig. 3</figureCitation>
|
||
). Further studies are needed to confirm whether these differences are individual or subspecific variations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |