155 lines
12 KiB
XML
155 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.715.15913" ID-GBIF-Dataset="894fe765-e2af-4695-9f90-fcf45b0c4a2c" ID-Pensoft-Pub="1313-2970-715-17" ID-ZBK="3587B83BD0D4494393CA09B16BBF231B" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-715-17" ModsDocOrigin="ZooKeys 715" ModsDocTitle="Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E.aipo-group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1510239869137" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sumoked, Bob & Balke, Michael" docDate="2017" docId="B28CAD0C4B7C1D3E3C53F021B5BD3B04" docLanguage="en" docName="ZooKeys 715: 17-37" docOrigin="ZooKeys 715" docSource="http://dx.doi.org/10.3897/zookeys.715.15913" docTitle="Exocelina okbapensis subsp. hajeki Shaverdo & Balke, ssp. n." docType="treatment" docUuid="56F86E0C-984E-4066-92FB-4739CD8DD7EC" docUuidSource="ZooBank" docVersion="6" lastPageNumber="26" masterDocId="FD28FFA11471A342006BFF903075FF81" masterDocTitle="Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E. aipo-group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="37" masterPageNumber="17" pageNumber="23" updateTime="1643478016727" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E. aipo-group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sumoked, Bob</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>715</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>17</mods:start>
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<mods:end>37</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.715.15913</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.715.15913</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-715-17</mods:identifier>
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<mods:identifier type="ZBK">3587B83BD0D4494393CA09B16BBF231B</mods:identifier>
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<mods:identifier type="ZooBank">3587B83BD0D4494393CA09B16BBF231B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137255797" LSID="urn:lsid:zoobank.org:act:56F86E0C-984E-4066-92FB-4739CD8DD7EC" httpUri="http://treatment.plazi.org/id/B28CAD0C4B7C1D3E3C53F021B5BD3B04" lastPageId="9" lastPageNumber="26" pageId="6" pageNumber="23">
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<subSubSection pageId="6" pageNumber="23" type="nomenclature">
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<paragraph pageId="6" pageNumber="23">
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1a.
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<taxonomicName LSID="http://zoobank.org/56F86E0C-984E-4066-92FB-4739CD8DD7EC" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina okbapensis" order="Coleoptera" pageId="6" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="okbapensis" subSpecies="hajeki">Exocelina okbapensis hajeki Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="6" pageNumber="23">ssp. n.</taxonomicNameLabel>
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Figs 3, 6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="24" type="reference_group">
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<paragraph pageId="7" pageNumber="24">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="7" pageNumber="24" phylum="Arthropoda" rank="genus">
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<pageBreakToken pageId="7" pageNumber="24" start="start">Exocelina</pageBreakToken>
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</taxonomicName>
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undescribed sp. MB0066:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="18" pageNumber="35" url="https://doi.org/10.1038/ncomms5001" year="2014">Toussaint et al. 2014</bibRefCitation>
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: Supplementary figs 1-4, tab. 2.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="24" type="type locality">
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<paragraph pageId="7" pageNumber="24">Type locality.</paragraph>
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<paragraph pageId="7" pageNumber="24">
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Papua: Jayawijaya Regency, Wamena,
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-4.06">04°03.6'S</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="92" value="139.03166">139°01.9'E</geoCoordinate>
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, 2050 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="24" type="type material">
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<paragraph pageId="7" pageNumber="24">Type material.</paragraph>
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<paragraph pageId="7" pageNumber="24">
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Holotype: male "IN, PA: Jayawijaya Regen., Baliem vall., 10km NE Wamena, forest above 'Baliem vall.
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<normalizedToken originalValue="Resort’">Resort'</normalizedToken>
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, 2050 m, 2-3.II.2015,
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-4.06">04°03.6'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="92" value="139.03166">139°01.9'E</geoCoordinate>
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; J.
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<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
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& J.
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<normalizedToken originalValue="Šumpich">Sumpich</normalizedToken>
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leg" (NMPC). Paratypes: 2 males and 7 females with the same label as the holotype, one female with an additional label "M. Balke 7372" (MZB, NHMW, NMPC, ZSM). 6 males, 11 females "INDONESIA, Papua: Jayawijaya Distr., Ballem valley, 10km NE of Wamena, forest above "Baliem valley resort",
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-4.06">04°03.6'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="92" value="139.03166">139°01.9'E</geoCoordinate>
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, 2050 m; 2-3.ii.2015 J.
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<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
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& J.
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<normalizedToken originalValue="Šumpich">Sumpich</normalizedToken>
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leg." (NHMW, NMPC, ZSM). 1 male "66 M. Balke" [green], "measured J. Parkin 43", "Indonesia: Irian Jaya, N Wamena, Cerny, M. Balke: MB 66" (ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="26" pageId="8" pageNumber="25" type="diagnosis">
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<paragraph pageId="8" pageNumber="25">
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<pageBreakToken pageId="8" pageNumber="25" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph lastPageId="9" lastPageNumber="26" pageId="8" pageNumber="25">
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Beetle medium-sized; dark brown to piceous, with reddish brown pronotal sides; submatt; pronotum with distinct lateral bead; male antennomeres 2-6 slightly, but evidently enlarged, female antennomeres 2-6 stout; male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, without
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<pageBreakToken pageId="9" pageNumber="26" start="start">concavity</pageBreakToken>
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, with anterior row of 26 and posterior row of eight relatively short setae; median lobe curved, with apex curved downwards and slightly rounded in lateral view. The subspecies differs from the nominative subspecies in the modified antennae, in the shape and setation of the paramere, and in the more striated abdominal ventrite 6; some beetles are also somehow slightly more matt due to more strongly impressed dorsal microreticulation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="26" type="description">
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<paragraph pageId="9" pageNumber="26">Description.</paragraph>
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<paragraph pageId="9" pageNumber="26">Size and shape: Beetle medium-sized (TL-H 3.75-4.5 mm, TL 4.1-4.9 mm, MW 2.0-2.3 mm), with oblong-oval habitus, broadest at elytral middle (Fig. 3). Coloration: As in nominative subspecies.</paragraph>
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<paragraph pageId="9" pageNumber="26">Surface sculpture: As in nominative subspecies apart from pronotum and elytra with evident, rather strongly impressed microreticulation, dorsal surface submatt.</paragraph>
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<paragraph pageId="9" pageNumber="26">Structures: As in nominative subspecies.</paragraph>
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<paragraph pageId="9" pageNumber="26">
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Male: Antennomeres 2-6 slightly, but evidently enlarged, antennomere 2 with slightly extended external upper angle (Fig. 3). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 26 and posterior row of eight relatively short setae (Fig. 6D). Median lobe curved, with apex curved downwards and slightly rounded in lateral view and evenly tapering to the slightly truncate apex in ventral view. Paramere with distinct notch on dorsal side and small, evidently separated subdistal part with a tuft of very dense, strong setae; proximal setae inconspicuous (Fig. 6
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Abdominal ventrite 6 with numerous (16-22) long lateral striae on each side.
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</paragraph>
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<caption pageId="9" pageNumber="26">
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<paragraph pageId="9" pageNumber="26">
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Figure 6.
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina okbapensis" order="Coleoptera" pageId="9" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="okbapensis" subSpecies="hajeki">Exocelina okbapensis hajeki</taxonomicName>
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ssp. n. A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4-5 in ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="9" pageNumber="26">Holotype: TL-H 4.0 mm, TL 4.4 mm, MW 2.05 mm.</paragraph>
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<paragraph pageId="9" pageNumber="26">Female: Antennomeres 2-6 stout, in some specimens only slightly more slender than in males, antennomere 2 with slightly extended external upper angle; pro- and mesotarsi not modified and abdominal ventrite 6 without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="26" type="variability">
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<paragraph pageId="9" pageNumber="26">Variability.</paragraph>
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<paragraph pageId="9" pageNumber="26">In some males, antennomeres 2-6 more strongly enlarged, in some others less strongly enlarged, similar to those of females.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="26" type="distribution">
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<paragraph pageId="9" pageNumber="26">Distribution.</paragraph>
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<paragraph pageId="9" pageNumber="26">Papua: Jayawijaya Regency. The subspecies is known only from the type locality (Fig. 12).</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="26" type="habitat">
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<paragraph pageId="9" pageNumber="26">Habitat.</paragraph>
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<paragraph pageId="9" pageNumber="26">Near Wamena, the species was collected from wet ground with weak water flow and forest puddles, which turn to a small creek during rain (Figs 13-14).</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="26" type="etymology">
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<paragraph pageId="9" pageNumber="26">Etymology.</paragraph>
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<paragraph pageId="9" pageNumber="26">
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The subspecies is named after our friend and colleague
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<normalizedToken originalValue="Jirí">Jiri</normalizedToken>
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<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
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who collected almost all the specimens. The name is a noun in the genitive case.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |