treatments-xml/data/B2/4A/49/B24A496CFFC6066CFC7FE862FA4DFAD6.xml

<document id="1F704C7A554CC306548BBA064CB66992" ID-DOI="10.3897/zoologia.35.e21351" ID-ISSN="1984-4689" ID-ZooBank="2961994A-B275-4653-86DB-80B5952C0D42" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722589191538" checkinUser="felipe" docAuthor="Pinto, Ângelo Parise &amp; Kompier, Tom" docDate="2018" docId="B24A496CFFC6066CFC7FE862FA4DFAD6" docLanguage="en" docName="Zoologia.35.e21351.pdf" docOrigin="Zoologia (e 21351) 35" docSource="http://dx.doi.org/10.3897/zoologia.35.e21351" docStyle="DocumentStyle:9D87517C92C8E430512793D922E147D4.1:Zoologia.2017-2020.journal_article" docStyleId="9D87517C92C8E430512793D922E147D4" docStyleName="Zoologia.2017-2020.journal_article" docStyleVersion="1" docTitle="Forcepsioneura serrabonita Pinto &amp; Kompier, 2018, sp. nov." docType="treatment" docUuid="9D6A447E-4F57-4DF3-AA37-BB59DA7F7829" docUuidSource="ZooBank" docVersion="1" lastPageNumber="12" masterDocId="4E733114FFC00667FFE6EE58FFD8FFA1" masterDocTitle="In honor of conservation of the Brazilian Atlantic Forest: description of two new damselflies of the genus Forcepsioneura discovered in private protected areas (Odonata: Coenagrionidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="7" updateTime="1722625001857" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="187CDE0A9409F1BFE0346A6D55E8955A">In honor of conservation of the Brazilian Atlantic Forest: description of two new damselflies of the genus Forcepsioneura discovered in private protected areas (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="645D24BF4561B6D72C5B83B80817496A">Pinto, Ângelo Parise</mods:namePart>
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<mods:namePart id="579EA53D91B6AE252E71B0EDEA467F6B">Kompier, Tom</mods:namePart>
<mods:affiliation id="0C2C866C770834A77201F10A27C1AD78">Schoutenstraat 69, 2596 SK, Den Haag, The Netherlands.</mods:affiliation>
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<mods:title id="5A3DC0F6D05CC0D871B475532DAAD2C6">Zoologia</mods:title>
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<mods:date id="6EBDC44E4F563C89332FB3B7C1D1C3A8">2018</mods:date>
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<taxonomicName id="FDE383F9FFC60661FC7FE862FB14F9F0" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[921,1228,1593,1617]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC60661FC7FE862FB14F9F0" box="[921,1228,1593,1617]" italics="true" pageId="6" pageNumber="7">Forcepsioneura serrabonita</emphasis>
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<taxonomicNameLabel id="13A49913FFC60661FB35E862FAECF9F0" box="[1235,1332,1594,1617]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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http://zoobank.org/ 
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<paragraph id="3A5CF87AFFC60661FC63E8DAFA9FF937" blockId="6.[816,1437,1640,1686]" box="[901,1351,1666,1686]" pageId="6" pageNumber="7">
<figureCitation id="A2D8E4FFFFC60661FC63E8DAFC3AF937" box="[901,994,1666,1686]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="6" pageNumber="7">Figs 7–12</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC60661FC0DE8DAFBF2F937" box="[1003,1066,1666,1686]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="6" pageNumber="7">13–16</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC60661FBD5E8DAFBAAF937" box="[1075,1138,1666,1686]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="6" pageNumber="7">21–23</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC60661FB9DE8DAFB62F937" box="[1147,1210,1666,1686]" captionStart="Figures 27–30" captionStartId="11.[151,218,1125,1145]" captionTargetBox="[151,1428,197,1105]" captionTargetId="figure-1@11.[151,1428,197,1105]" captionTargetPageId="11" captionText="Figures 27–30. Head and prothorax in dorsal view of females of Forcepsioneura: (27–28) F. regua sp. nov., paratype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (29–30) F. serrabonita sp. nov., paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars = 1 mm." pageId="6" pageNumber="7">29–30</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC60661FB25E8DAFAD9F937" box="[1219,1281,1666,1686]" captionStart="Figures 31–34" captionStartId="12.[159,223,1739,1759]" captionTargetBox="[244,1350,197,1720]" captionTargetId="figure-0@12.[244,1350,197,1720]" captionTargetPageId="12" captionText="Figures 31–34. Abdominal segments seven to ten (S7–10) of females of Forcepsioneura: (31–32)F.regua sp.nov., paratype (Brazil.RJ:Cachoeiras de Macacu, DZRJ); (33–34) F. serrabonita sp. nov., paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars for Figs 31–32 = 1 mm." pageId="6" pageNumber="7">33–34</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC60661FAEDE8DAFA9FF937" box="[1291,1351,1666,1686]" captionStart="Figures 35–40" captionStartId="13.[151,218,1575,1595]" captionTargetBox="[151,1428,212,1555]" captionTargetId="figure-105@13.[151,1428,197,1555]" captionTargetPageId="13" captionText="Figures 35–40. Habitus and habitat of species of Forcepsioneura: (35) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (36) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050), (37) F. regua sp. nov., live individual at type locality at Reserva Ecológica de Guapiaçu (REGUA); (38) overview of type locality at RPPN Serra Bonita; (39) shallow muddy and slow stream at Itatiaia National Park, typical habitat of small light blue species; (40) seepage covered by hornworts at Três Picos State Park, typical habitat of large orange-black species. Scale bars = 1 mm. Photos: (37) TMFK; (38) RA Carvalho; (39–40) APP." pageId="6" pageNumber="7">36, 38</figureCitation>
</paragraph>
<paragraph id="3A5CF87AFFC6066EFC8BE8EEFE7DFE2D" blockId="6.[817,1437,1718,1768]" lastBlockId="9.[149,775,198,1770]" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
Diagnosis. A medium, dorsally black and lateroventrally yellowish protoneurid species; frons angulated; rear of head pale; posterolateral margin of median lobe of prothorax with well-defined tubercle-like process; CuP&amp;AA indistinct; Ax space 1 as long as Ax space 3; GL with long inner fold and distal lateral lobe prolonged into a flagellum; cercus forcipate with mediobasal (MBP) and ventrobasal process (VBP), all characters agreeing with the definition of 
<taxonomicName id="FDE383F9FFC9066EFEDDEF04FE1BFECF" authorityName="Lencioni" authorityYear="1999" box="[315,451,348,366]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="08972468FFC9066EFEDDEF04FE1BFECF" box="[315,451,348,366]" italics="true" pageId="9" pageNumber="10">Forcepsioneura</emphasis>
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as recently diagnosed (Machado 2009, 
<bibRefCitation id="5E72858BFFC9066EFF35EF21FE43FE2D" author="Garrison RW &amp; von Ellenrieder N &amp; Louton JA" box="[211,411,377,397]" pageId="9" pageNumber="10" refId="ref13131" refString="Garrison RW, von Ellenrieder N, Louton JA (2010) Damselfly genera of the New World. An illustrated and annotated key to the Zygoptera. The John Hopkins University Press, Baltimore, 490 pp." type="book" year="2010">Garrison et al. 2010</bibRefCitation>
).
</paragraph>
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<paragraph id="3A5CF87AFFC70660FF71E8F5FA4DF95C" blockId="7.[150,1430,1709,1789]" pageId="7" pageNumber="8">
Figures 13–20. Prothorax and genital ligula of males of 
<taxonomicName id="FDE383F9FFC70660FD75E8F5FCC0F960" authorityName="Lencioni" authorityYear="1999" box="[659,792,1709,1729]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="08972468FFC70660FD75E8F5FCC0F960" box="[659,792,1709,1729]" italics="true" pageId="7" pageNumber="8">Forcepsioneura</emphasis>
</taxonomicName>
: (13–16) 
<taxonomicName id="FDE383F9FFC70660FC94E8F5FC32F960" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[882,1002,1709,1729]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC70660FC94E8F5FC32F960" box="[882,1002,1709,1729]" italics="true" pageId="7" pageNumber="8">F. serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC70660FC16E8F5FBE4F960" box="[1008,1084,1709,1729]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) 
<taxonomicName id="FDE383F9FFC70660FECDE893FEA9F97E" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[299,369,1739,1759]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="regua" status="sp. nov.">
<emphasis id="08972468FFC70660FECDE893FEA9F97E" box="[299,369,1739,1759]" italics="true" pageId="7" pageNumber="8">F. regua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC70660FE9EE893FE1DF97E" box="[376,453,1739,1759]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm.
</paragraph>
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<paragraph id="3A5CF87AFFC8066FFF79E82EFDE0F964" blockId="8.[159,1438,1653,1733]" pageId="8" pageNumber="9">
Figures 21–26. Caudal appendages of males of 
<taxonomicName id="FDE383F9FFC8066FFD97E82DFD2EF928" authorityName="Lencioni" authorityYear="1999" box="[625,758,1653,1673]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="08972468FFC8066FFD97E82DFD2EF928" box="[625,758,1653,1673]" italics="true" pageId="8" pageNumber="9">Forcepsioneura</emphasis>
</taxonomicName>
: (21–23) 
<taxonomicName id="FDE383F9FFC8066FFCBFE82DFC0DF928" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[857,981,1653,1673]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC8066FFCBFE82DFC0DF928" box="[857,981,1653,1673]" italics="true" pageId="8" pageNumber="9">F. serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC8066FFC38E82EFBF6F92B" box="[990,1070,1654,1674]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) 
<taxonomicName id="FDE383F9FFC8066FFEDFE8CBFE59F909" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[313,385,1683,1704]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="regua" status="sp. nov.">
<emphasis id="08972468FFC8066FFEDFE8CBFE59F909" box="[313,385,1683,1704]" italics="true" pageId="8" pageNumber="9">F. regua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC8066FFE6EE8CCFE0FF909" box="[392,471,1684,1704]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="3A5CF87AFFC9066EFF32EFCFFDE2FCCB" blockId="9.[149,775,198,1770]" pageId="9" pageNumber="10">
Based on general color and shape of male cercus, 
<taxonomicName id="FDE383F9FFC9066EFD5CEFC0FF0AFE69" authorityName="Pinto &amp; Kompier" authorityYear="2018" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC9066EFD5CEFC0FF0AFE69" italics="true" pageId="9" pageNumber="10">F. serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC9066EFF3EEFEDFEFDFE69" box="[216,293,437,456]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
resembles 
<taxonomicName id="FDE383F9FFC9066EFE69EFEEFDD0FE69" authorityName="Machado" authorityYear="2001" box="[399,520,438,456]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grossiorum">
<emphasis id="08972468FFC9066EFE69EFEEFDD0FE69" box="[399,520,438,456]" italics="true" pageId="9" pageNumber="10">F. grossiorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="FDE383F9FFC9066EFDDEEFEEFDA2FE69" authorityName="Machado" authorityYear="2000" box="[568,634,437,456]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lucia">
<emphasis id="08972468FFC9066EFDDEEFEEFDA2FE69" box="[568,634,437,456]" italics="true" pageId="9" pageNumber="10">F. lucia</emphasis>
</taxonomicName>
. Males of this orange-black group of large species with robust cercus have a short VBP (as long as≤ 0.40 of dorsal branch of cercus in lateral view), while it is longer (as long as ≥ 0.55 of dorsal branch) in 
<taxonomicName id="FDE383F9FFC9066EFF7EEC75FF25FD9E" authorityName="Lencioni" authorityYear="1999" box="[152,253,557,575]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="garrisoni">
<emphasis id="08972468FFC9066EFF7EEC75FF25FD9E" box="[152,253,557,575]" italics="true" pageId="9" pageNumber="10">F. garrisoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="FDE383F9FFC9066EFEEFEC75FEBCFD9E" authorityName="Machado" authorityYear="2001" box="[265,356,556,575]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="haerteli">
<emphasis id="08972468FFC9066EFEEFEC75FEBCFD9E" box="[265,356,556,575]" italics="true" pageId="9" pageNumber="10">F. haerteli</emphasis>
</taxonomicName>
, 
<taxonomicName id="FDE383F9FFC9066EFE96EC75FE0BFD9E" baseAuthorityName="Santos" baseAuthorityYear="1970" box="[368,467,557,575]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="itatiaiae">
<emphasis id="08972468FFC9066EFE96EC75FE0BFD9E" box="[368,467,557,575]" italics="true" pageId="9" pageNumber="10">F. itatiaiae</emphasis>
</taxonomicName>
, 
<taxonomicName id="FDE383F9FFC9066EFE39EC75FDFEFD9E" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[479,550,557,575]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="regua" status="sp. nov.">
<emphasis id="08972468FFC9066EFE39EC75FDFEFD9E" box="[479,550,557,575]" italics="true" pageId="9" pageNumber="10">F. regua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC9066EFDCAEC75FDA2FDE1" box="[556,634,557,576]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="FDE383F9FFC9066EFD4AEC75FD25FD9E" box="[684,765,557,575]" class="Insecta" family="Protoneuridae" genus="Phasmoneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sancta">
<emphasis id="08972468FFC9066EFD4AEC75FD25FD9E" box="[684,765,557,575]" italics="true" pageId="9" pageNumber="10">F. sancta</emphasis>
</taxonomicName>
. 
<taxonomicName id="FDE383F9FFC9066EFF71EC13FE53FDFC" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[151,395,586,605]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC9066EFF71EC13FE53FDFC" box="[151,395,586,605]" italics="true" pageId="9" pageNumber="10">Forcepsioneura serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC9066EFE77EC13FE06FDFF" box="[401,478,587,606]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from the first two species by having posterior lobe of prothorax regularly rounded (
<figureCitation id="A2D8E4FFFFC9066EFF1EECDEFEEBFD38" box="[248,307,646,666]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 9</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFED8ECDEFE80FD38" box="[318,344,646,665]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="9" pageNumber="10">13</figureCitation>
; sinuous in 
<taxonomicName id="FDE383F9FFC9066EFE37ECDFFD93FD38" authorityName="Machado" authorityYear="2001" box="[465,587,647,665]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grossiorum">
<emphasis id="08972468FFC9066EFE37ECDFFD93FD38" box="[465,587,647,665]" italics="true" pageId="9" pageNumber="10">F. grossiorum</emphasis>
</taxonomicName>
); Pt quadrangular, ratio between costal length and proximal length 
<quantity id="FD1B559FFFC9066EFD46ECFCFD32FD16" box="[672,746,676,695]" metricMagnitude="-2" metricUnit="m" metricValue="1.3716" pageId="9" pageNumber="10" unit="in" value="0.54">0.54 in</quantity>
<taxonomicName id="FDE383F9FFC9066EFD12ECFCFF26FD74" authorityName="Pinto &amp; Kompier" authorityYear="2018" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC9066EFD12ECFCFF26FD74" italics="true" pageId="9" pageNumber="10">F. serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC9066EFEE3EC9AFE8DFD74" box="[261,341,706,725]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, being similar in size to 
<taxonomicName id="FDE383F9FFC9066EFDA0EC9AFD50FD74" authorityName="Machado" authorityYear="2000" box="[582,648,706,725]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lucia">
<emphasis id="08972468FFC9066EFDA0EC9AFD50FD74" box="[582,648,706,725]" italics="true" pageId="9" pageNumber="10">F. lucia</emphasis>
</taxonomicName>
(ratio ≅ 0.6), but rectangular and distinctly longer in 
<taxonomicName id="FDE383F9FFC9066EFDD2ECB8FD68FD53" authorityName="Machado" authorityYear="2001" box="[564,688,736,754]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grossiorum">
<emphasis id="08972468FFC9066EFDD2ECB8FD68FD53" box="[564,688,736,754]" italics="true" pageId="9" pageNumber="10">F. grossiorum</emphasis>
</taxonomicName>
(ratio = 0.8); cercus slender, with anteromesal margin of MBP process rounded in dorsal view (
<figureCitation id="A2D8E4FFFFC9066EFE6CED43FE09FC8F" box="[394,465,795,815]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 10</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFE3AED43FE2FFC8F" box="[476,503,795,814]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="9" pageNumber="10">22</figureCitation>
; acute in both 
<taxonomicName id="FDE383F9FFC9066EFD6CED44FCDBFC8F" authorityName="Machado" authorityYear="2001" box="[650,771,796,814]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grossiorum">
<emphasis id="08972468FFC9066EFD6CED44FCDBFC8F" box="[650,771,796,814]" italics="true" pageId="9" pageNumber="10">F. grossiorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="FDE383F9FFC9066EFF24ED62FEDCFCED" authorityName="Machado" authorityYear="2000" box="[194,260,825,844]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lucia">
<emphasis id="08972468FFC9066EFF24ED62FEDCFCED" box="[194,260,825,844]" italics="true" pageId="9" pageNumber="10">F. lucia</emphasis>
</taxonomicName>
), MBP almost not visible in lateral view (
<figureCitation id="A2D8E4FFFFC9066EFD72ED61FD03FCED" box="[660,731,825,845]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 11</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFD02ED61FD26FCED" box="[740,766,825,844]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="9" pageNumber="10">21</figureCitation>
; clearly visible in 
<taxonomicName id="FDE383F9FFC9066EFED9ED00FE60FCCB" authorityName="Machado" authorityYear="2001" box="[319,440,856,874]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grossiorum">
<emphasis id="08972468FFC9066EFED9ED00FE60FCCB" box="[319,440,856,874]" italics="true" pageId="9" pageNumber="10">F. grossiorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="FDE383F9FFC9066EFE0CED00FDF4FCCB" authorityName="Machado" authorityYear="2000" box="[490,556,855,874]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lucia">
<emphasis id="08972468FFC9066EFE0CED00FDF4FCCB" box="[490,556,855,874]" italics="true" pageId="9" pageNumber="10">F. lucia</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="3A5CF87AFFC9066EFF32ED2DFCD9FBF8" blockId="9.[149,775,198,1770]" pageId="9" pageNumber="10">
In view of the large size of specimens of 
<taxonomicName id="FDE383F9FFC9066EFD83ED2EFD39FC29" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[613,737,885,904]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC9066EFD83ED2EFD39FC29" box="[613,737,885,904]" italics="true" pageId="9" pageNumber="10">F. serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC9066EFD01ED2DFF1DFC07" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, they can be confused with 
<taxonomicName id="FDE383F9FFC9066EFE3DEDCBFDE5FC04" baseAuthorityName="Santos" baseAuthorityYear="1970" box="[475,573,915,933]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="itatiaiae">
<emphasis id="08972468FFC9066EFE3DEDCBFDE5FC04" box="[475,573,915,933]" italics="true" pageId="9" pageNumber="10">F. itatiaiae</emphasis>
</taxonomicName>
, from which, in addition to the shorter VBP mentioned above, it can be separated by the shape of the male cercus with strongly cylindrical VBP (
<figureCitation id="A2D8E4FFFFC9066EFF78EDB4FF3EFBA1" box="[158,230,1004,1024]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 11</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFF17EDB5FED3FBA1" box="[241,267,1005,1024]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="9" pageNumber="10">21</figureCitation>
; blade-like and truncated apex in 
<taxonomicName id="FDE383F9FFC9066EFDB9EDB5FD19FC5E" baseAuthorityName="Santos" baseAuthorityYear="1970" box="[607,705,1005,1023]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="itatiaiae">
<emphasis id="08972468FFC9066EFDB9EDB5FD19FC5E" box="[607,705,1005,1023]" italics="true" pageId="9" pageNumber="10">F. itatiaiae</emphasis>
</taxonomicName>
), MBP largely rounded dorsally and almost not visible in lateral view (
<figureCitation id="A2D8E4FFFFC9066EFF78EA70FED6FB9A" box="[158,270,1064,1084]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 10–11</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFEFDEA70FE82FB9A" box="[283,346,1064,1083]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="9" pageNumber="10">21–22</figureCitation>
; angulated plate as an axe with concave blade dorsally, and clearly visible in lateral view in 
<taxonomicName id="FDE383F9FFC9066EFD74EA1FFD2CFBF8" baseAuthorityName="Santos" baseAuthorityYear="1970" box="[658,756,1095,1113]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="itatiaiae">
<emphasis id="08972468FFC9066EFD74EA1FFD2CFBF8" box="[658,756,1095,1113]" italics="true" pageId="9" pageNumber="10">F. itatiaiae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="3A5CF87AFFC9066EFF32EA3BFD72FB71" blockId="9.[149,775,198,1770]" pageId="9" pageNumber="10">
Additionally, the distinctive shape of the GL (
<figureCitation id="A2D8E4FFFFC9066EFD68EA3BFD20FBD6" box="[654,760,1123,1143]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="9" pageNumber="10">Figs 14–16</figureCitation>
), with a large and upright internal fold (obliquely toward anteriorly in the other species), will separate 
<taxonomicName id="FDE383F9FFC9066EFDD7EAF8FD77FB12" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[561,687,1184,1203]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFC9066EFDD7EAF8FD77FB12" box="[561,687,1184,1203]" italics="true" pageId="9" pageNumber="10">F. serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFC9066EFD53EAF8FCDCFB12" box="[693,772,1184,1203]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
from all species for which the GL has been described.
</paragraph>
<paragraph id="3A5CF87AFFC9066EFF32EA83FE9DFAE9" blockId="9.[149,775,198,1770]" pageId="9" pageNumber="10">
Females of several 
<taxonomicName id="FDE383F9FFC9066EFE67EA84FD97FB4F" box="[385,591,1243,1262]" pageId="9" pageNumber="10">
<emphasis id="08972468FFC9066EFE67EA84FDDEFB4F" box="[385,518,1244,1262]" italics="true" pageId="9" pageNumber="10">Forcepsioneura</emphasis>
species
</taxonomicName>
are poorly known, which hampers a useful diagnosis. Putative females should be identified by comparison with descriptions, figures and association with males.
</paragraph>
<paragraph id="3A5CF87AFFC9066EFF32EB0BFBD6FEB4" blockId="9.[149,775,198,1770]" lastBlockId="9.[807,1431,198,1777]" pageId="9" pageNumber="10">
Description of male 
<typeStatus id="E55846D8FFC9066EFE44EB0AFE26FAC7" box="[418,510,1362,1382]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
. Head (
<figureCitation id="A2D8E4FFFFC9066EFDACEB0AFD47FAC7" box="[586,671,1362,1382]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 7–8</figureCitation>
): Labium, visible parts of maxillae and mandibles (except apex) ivory yellow (
<figureCitation id="A2D8E4FFFFC9066EFF2FEBD6FED9FA03" box="[201,257,1422,1442]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Fig. 8</figureCitation>
). Genae yellowish, darkening gradually posteriorly. Labrum orange-brown (possibly light orange in life) with two lateral and one mesal rounded dark brown spots from clypeolabral suture to 0.5 ventral, pale areas among them darkened. Anteclypeus with lateral and mesal membranous ventral part yellowish, remainder part forming a semicircular dark brown plate, postclypeus almost entirely shining dark brown with ill-defined orange-brown spots. Antefrons dark brown with ill-defined pale areas, postfrons and remainder of epicranium opaque black with a weak copper luster. Antenna, antennifer and scape black, posterior surface of pedicel light dark brown, all parts with laterofrontal surface and distal apex of antennifer and scape ivory yellow; flagellum lost. Posterior region of the cranium (“postgena” plus “occiput”) pale brown (
<figureCitation id="A2D8E4FFFFC9066EFAF1EEBBFA96FF56" box="[1303,1358,227,247]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Fig. 7</figureCitation>
), probably light ivory yellow.
</paragraph>
<paragraph id="3A5CF87AFFC9066EFC83EF7AFBD4FBDB" blockId="9.[807,1431,198,1777]" pageId="9" pageNumber="10">
Thorax (
<figureCitation id="A2D8E4FFFFC9066EFC5DEF79FC20FE94" box="[955,1016,289,309]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 9</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFBE3EF7AFBF8FE94" box="[1029,1056,290,309]" captionStart="Figures 35–40" captionStartId="13.[151,218,1575,1595]" captionTargetBox="[151,1428,212,1555]" captionTargetId="figure-105@13.[151,1428,197,1555]" captionTargetPageId="13" captionText="Figures 35–40. Habitus and habitat of species of Forcepsioneura: (35) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (36) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050), (37) F. regua sp. nov., live individual at type locality at Reserva Ecológica de Guapiaçu (REGUA); (38) overview of type locality at RPPN Serra Bonita; (39) shallow muddy and slow stream at Itatiaia National Park, typical habitat of small light blue species; (40) seepage covered by hornworts at Três Picos State Park, typical habitat of large orange-black species. Scale bars = 1 mm. Photos: (37) TMFK; (38) RA Carvalho; (39–40) APP." pageId="9" pageNumber="10">36</figureCitation>
): Prothorax dorsally black, gradually lightening ventrally to yellow at pleural sclerites; anterior margin of anterior lobe ivory yellow; posterolateral margin of median lobe with well-defined tubercle-like process; posterior lobe rounded convex, narrower than median (
<figureCitation id="A2D8E4FFFFC9066EFB09EFC1FAF1FE0D" box="[1263,1321,409,429]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 9</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFAD5EFC1FA95FE0D" box="[1331,1357,409,428]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="9" pageNumber="10">13</figureCitation>
); sternellum (furcasternum sensu 
<bibRefCitation id="5E72858BFFC9066EFBD4EFEFFB60FE6B" author="Asahina S" box="[1074,1208,439,458]" pageId="9" pageNumber="10" refId="ref12455" refString="Asahina S (1954) A morphological study of a relic dragonfly Epiophlebia superstes Selys (Odonata, Anisozygoptera). The Japan Society for Promotion of Science, Tokyo, 153 pp." type="book" year="1954">Asahina 1954</bibRefCitation>
) almost entirely black. Synthorax (
<figureCitation id="A2D8E4FFFFC9066EFC7EEF8DFC01FE49" box="[920,985,469,489]" captionStart="Figures 35–40" captionStartId="13.[151,218,1575,1595]" captionTargetBox="[151,1428,212,1555]" captionTargetId="figure-105@13.[151,1428,197,1555]" captionTargetPageId="13" captionText="Figures 35–40. Habitus and habitat of species of Forcepsioneura: (35) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (36) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050), (37) F. regua sp. nov., live individual at type locality at Reserva Ecológica de Guapiaçu (REGUA); (38) overview of type locality at RPPN Serra Bonita; (39) shallow muddy and slow stream at Itatiaia National Park, typical habitat of small light blue species; (40) seepage covered by hornworts at Três Picos State Park, typical habitat of large orange-black species. Scale bars = 1 mm. Photos: (37) TMFK; (38) RA Carvalho; (39–40) APP." pageId="9" pageNumber="10">Fig. 36</figureCitation>
) dorsally dark, gradually lightening ventrally; mesepisternum from mesostigmal region to antealar sinus entirely dark brown to black with greenish-copper metallic reflections, except light brown line running longitudinal to mesopleural suture; mesepimeron dorsally light brown, narrowed ventrally by an ivory yellow stripe occupying 0.4 of the lateroventral apex of the sclerite, over mesinfraepisternum it occupies 0.5 ventral; remainder of synthorax ivory yellow, except poorly defined light brown stripe with about 0.3 of width along metapleural suture ending at spiracle level and a small black spot at ventro-distal apex of metapostepimeron. Legs ivory yellow with irregular dark brown to black areas in articulation femur-tibia, dorsal surface of metathoracic femur and apex of pretarsal claws (pretarsus); spurs black except scale-like ivory yellow proximal femoral spurs and tibial comb of prothoracic leg; anteroventral surface of femora armed with long and robust spurs, 
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pro- and mesothoracic and 
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metathoracic femora, posteroventral surface with short- er and thinner spurs, 
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pro- and mesothoracic femora to 
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metathoracic femur; anteroventral surface of tibiae armed with 9 spurs in pro- (4 of tibial comb), 
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meso-, 
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metathoracic tibiae, usually longer than intervening spaces; pretarsal claws with distinct acute supplementary inferior tooth at ca 0.50 of their length in all legs.
</paragraph>
<paragraph id="3A5CF87AFFC9066EFC83EADEFA4DFAEC" blockId="9.[807,1431,198,1777]" pageId="9" pageNumber="10">
Wings (
<figureCitation id="A2D8E4FFFFC9066EFC49EADEFC2BFB38" box="[943,1011,1158,1178]" captionStart="Figures 35–40" captionStartId="13.[151,218,1575,1595]" captionTargetBox="[151,1428,212,1555]" captionTargetId="figure-105@13.[151,1428,197,1555]" captionTargetPageId="13" captionText="Figures 35–40. Habitus and habitat of species of Forcepsioneura: (35) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (36) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050), (37) F. regua sp. nov., live individual at type locality at Reserva Ecológica de Guapiaçu (REGUA); (38) overview of type locality at RPPN Serra Bonita; (39) shallow muddy and slow stream at Itatiaia National Park, typical habitat of small light blue species; (40) seepage covered by hornworts at Três Picos State Park, typical habitat of large orange-black species. Scale bars = 1 mm. Photos: (37) TMFK; (38) RA Carvalho; (39–40) APP." pageId="9" pageNumber="10">Fig. 36</figureCitation>
): Membrane hyaline; venation dark brown to black; Pt brown, with a narrow internal pale line surrounding the enclosing veins, quadrangular. MP reaches anal margin slightly distal of vein descending from subnodus, covering 2 cells in FW, distal to vein descending from subnodus covering ca 0.5 of a third cell in HW. Px in Fw 12; Hw 11; RP2 originating at or slightly distal to Px 
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Fw, at or slightly proximal to Px 
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Hw.
</paragraph>
<paragraph id="3A5CF87AFFC9066DFC83EB02FE41FC40" blockId="9.[807,1431,198,1777]" lastBlockId="10.[157,781,198,1771]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Abdomen (
<figureCitation id="A2D8E4FFFFC9066EFC3CEB01FB95FACC" box="[986,1101,1369,1389]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="9" pageNumber="10">Figs 10–12</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFBBCEB02FB43FACC" box="[1114,1179,1370,1389]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="9" pageNumber="10">14–16</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFB4EEB02FB30FACC" box="[1192,1256,1370,1389]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="9" pageNumber="10">21–23</figureCitation>
, 
<figureCitation id="A2D8E4FFFFC9066EFB10EB02FACAFACC" box="[1270,1298,1370,1389]" captionStart="Figures 35–40" captionStartId="13.[151,218,1575,1595]" captionTargetBox="[151,1428,212,1555]" captionTargetId="figure-105@13.[151,1428,197,1555]" captionTargetPageId="13" captionText="Figures 35–40. Habitus and habitat of species of Forcepsioneura: (35) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (36) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050), (37) F. regua sp. nov., live individual at type locality at Reserva Ecológica de Guapiaçu (REGUA); (38) overview of type locality at RPPN Serra Bonita; (39) shallow muddy and slow stream at Itatiaia National Park, typical habitat of small light blue species; (40) seepage covered by hornworts at Três Picos State Park, typical habitat of large orange-black species. Scale bars = 1 mm. Photos: (37) TMFK; (38) RA Carvalho; (39–40) APP." pageId="9" pageNumber="10">36</figureCitation>
): Tergites of S1–10 dark brown to black dorsally, yellow ivory lateroventrally, darker in S8–10; sternites similar in color to adjacent areas of tergite, yellow ivory in S1, remainder segments light yellow to orange-yellow with irregular dark areas; pale longitudinal stripe occupying 0.5 ventral of S1–2 tergites laterally, gradually narrowing to ca 
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S8, S9 with two ill-defined small pale spots, one close to posterior field of S8 and other close to posterior carina, S10 almost entirely black; S3–7 with an narrow anterior pale ring ≤0.1 of total segment length, dorsally separated by a thin line; S1–7 cylindrical, S8–10 distinctly wider than previous segments, S7–8 slightly smashed preventing an accurate measurement of abdominal width in these segments; posterior margin of S10 with a shallow and wide concavity. Secondary genitalia (
<figureCitation id="A2D8E4FFFFC9066EFAC5E885FA54F950" box="[1315,1420,1757,1777]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="9" pageNumber="10">Figs 14–16</figureCitation>
) typical of Coenagrionoidea, anterior lamina with a deep and acute incision, anterior hamule quadrangular, with anteroventral angle acutely projected, posterior hamule almost entirely internalized with a curved thumb-shape; VS longer than wide, maximum width 0.4 of total length in ventral view. Genital ligula (as in 
<figureCitation id="A2D8E4FFFFCA066DFF31EF03FEE6FECF" box="[215,318,347,367]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="10" pageNumber="11">Figs 14–16</figureCitation>
) rectangular in ectal view; L1 smooth, without any kind of special setae; L2 with posterolateral portion of flexure projected distally beyond median region, mesally slightly projected in ectal view, in lateral view this projection similar to terminal fold (
<figureCitation id="A2D8E4FFFFCA066DFECAEF8AFEA8FE47" box="[300,368,466,486]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="10" pageNumber="11">Fig. 15</figureCitation>
); distal margin of L2 (tip of ligula) almost straight in ectal view; distal lateral lobe prolonged into a gradually thinner, twisted, flagellum; internal fold proximal to flexure, upright, large, ca 0.4 of L2 total length in lateral view (
<figureCitation id="A2D8E4FFFFCA066DFD5FEC74FD25FD9E" box="[697,765,556,576]" captionStart="Figures 13–20" captionStartId="7.[151,218,1709,1729]" captionTargetBox="[217,1365,200,1683]" captionTargetId="figure-1@7.[214,1365,197,1690]" captionTargetPageId="7" captionText="Figures 13–20. Prothorax and genital ligula of males of Forcepsioneura: (13–16) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (17–20) F. regua sp. nov., (17) holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071), remainder from paratype of same locality; (13, 17) prothorax in dorsal view; (14–16, 18–20) genital ligula in ventral (14, 18) and lateral (15–16, 19–20) views. Scale bars = 0.5 mm." pageId="10" pageNumber="11">Fig. 16</figureCitation>
); no sclerotized tubercle at level of flexure. Epiproct reduced to a membranous-like plate. Cercus (
<figureCitation id="A2D8E4FFFFCA066DFE30EC30FDE7FDDA" box="[470,575,616,636]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="10" pageNumber="11">Figs 10–12</figureCitation>
, 
<figureCitation id="A2D8E4FFFFCA066DFDA1EC30FD5BFDDA" box="[583,643,616,635]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="10" pageNumber="11">21–23</figureCitation>
) ivory yellow, with apex, ventrobasal (VBP) and mediobasal (MBP) processes dark brown to black; in lateral view strongly directed obliquely dorsad; VBP in opposite direction, short, apex rounded, distal edge at half distance from ventral margin of S10; MBP visible as a rounded tubercle between cercus and VBP; tip of cercus blunt; in dorsal view forcipate, proximally wide, slender distally, lateral margin gently curved, internal margin slightly more curved, apices converging (
<figureCitation id="A2D8E4FFFFCA066DFEDCED0FFE58FCCB" box="[314,384,855,875]" captionStart="Figures 7–12" captionStartId="4.[159,224,1564,1584]" captionTargetBox="[164,1431,197,1544]" captionTargetId="figure-0@4.[164,1431,197,1544]" captionTargetPageId="4" captionText="Figures 7–12. Forcepsioneura serrabonita sp. nov., holotype male (Brazil. BA: Camacan, DZRJ 0050): (7–8) head in dorsal (7) and ventral (8) views;(9) prothorax in dorsolateral view; (10–12) caudal appendages in dorsal (10), lateral (11) and dorsolateral (12) views. Scale bars = 1 mm." pageId="10" pageNumber="11">Figs 10</figureCitation>
, 
<figureCitation id="A2D8E4FFFFCA066DFE6FED0FFE7CFCCB" box="[393,420,855,874]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="10" pageNumber="11">22</figureCitation>
); MBP as a rounded plate, positioned at 0.33 basal; in posterior view obliquely directed ventrally and projected from a dilated area of cercus; apex of VBP process acute and slightly curved in posterior view (
<figureCitation id="A2D8E4FFFFCA066DFDB3EDE8FD43FC65" box="[597,667,944,964]" captionStart="Figures 21–26" captionStartId="8.[159,226,1654,1674]" captionTargetBox="[167,1426,199,1631]" captionTargetId="figure-0@8.[159,1436,197,1634]" captionTargetPageId="8" captionText="Figures 21–26. Caudal appendages of males of Forcepsioneura: (21–23) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050); (24–26) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (21, 24) lateral view; (22, 25) dorsal view; (23, 26) posterior view. Scale bars = 1 mm." pageId="10" pageNumber="11">Fig. 23</figureCitation>
). Paraproct orange-yellow, plate-like.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFF3DEDB5FEBAFB71" blockId="10.[157,781,198,1771]" pageId="10" pageNumber="11">
Measurements (mm). Total length (incl. caudal appendages) 38.7; abdomen length (excluding caudal appendages) 33.0; head maximum width 3.4; Fw length 21.8; Hw length 20.3; Fw maximum width 3.8, 
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Hw; Pt length 
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Fw, 
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Hw; length of metathoracic femur 2.0; metathoracic tibia 2.1; length of S9+ 
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lateral view 1.46; total length of cercus (dorsal branch) in lateral view 0.88; length of ventrobasal process in lateral view 0.36.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFF3DEA83FEBDFAAD" blockId="10.[157,781,198,1771]" pageId="10" pageNumber="11">
Female 
<typeStatus id="E55846D8FFCA066DFEC2EA84FEA4FB4E" box="[292,380,1244,1263]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
. Similar to 
<typeStatus id="E55846D8FFCA066DFE03EA83FD98FB4E" box="[485,576,1243,1263]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
. The only differences are described below.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFF3DEB4FFDBFFA1E" blockId="10.[157,781,198,1771]" pageId="10" pageNumber="11">
Head (
<figureCitation id="A2D8E4FFFFCA066DFEFDEB4FFE51FA8B" box="[283,393,1303,1323]" captionStart="Figures 27–30" captionStartId="11.[151,218,1125,1145]" captionTargetBox="[151,1428,197,1105]" captionTargetId="figure-1@11.[151,1428,197,1105]" captionTargetPageId="11" captionText="Figures 27–30. Head and prothorax in dorsal view of females of Forcepsioneura: (27–28) F. regua sp. nov., paratype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (29–30) F. serrabonita sp. nov., paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars = 1 mm." pageId="10" pageNumber="11">Figs 29–30</figureCitation>
): Labrum light orange with two lateral and a mesal rounded dark brown spots from clypeolabral suture to 0.5 ventral. Anteclypeus yellowish. Antefrons ivory yellow. Scape articulation and posterior surface of pedicel ivory yellow. Posterior region of the cranium (“postgena” plus “occiput”) yellowish, brownish dorsally close to foramen
</paragraph>
<paragraph id="3A5CF87AFFCA066DFF3DEB92FDFFF930" blockId="10.[157,781,198,1771]" pageId="10" pageNumber="11">
Thorax (
<figureCitation id="A2D8E4FFFFCA066DFECAEB92FEA9FA7C" box="[300,369,1482,1502]" captionStart="Figures 27–30" captionStartId="11.[151,218,1125,1145]" captionTargetBox="[151,1428,197,1105]" captionTargetId="figure-1@11.[151,1428,197,1105]" captionTargetPageId="11" captionText="Figures 27–30. Head and prothorax in dorsal view of females of Forcepsioneura: (27–28) F. regua sp. nov., paratype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (29–30) F. serrabonita sp. nov., paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars = 1 mm." pageId="10" pageNumber="11">Figs 30</figureCitation>
, 
<figureCitation id="A2D8E4FFFFCA066DFE9FEB92FE4CFA7C" box="[377,404,1482,1501]" captionStart="Figures 35–40" captionStartId="13.[151,218,1575,1595]" captionTargetBox="[151,1428,212,1555]" captionTargetId="figure-105@13.[151,1428,197,1555]" captionTargetPageId="13" captionText="Figures 35–40. Habitus and habitat of species of Forcepsioneura: (35) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (36) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050), (37) F. regua sp. nov., live individual at type locality at Reserva Ecológica de Guapiaçu (REGUA); (38) overview of type locality at RPPN Serra Bonita; (39) shallow muddy and slow stream at Itatiaia National Park, typical habitat of small light blue species; (40) seepage covered by hornworts at Três Picos State Park, typical habitat of large orange-black species. Scale bars = 1 mm. Photos: (37) TMFK; (38) RA Carvalho; (39–40) APP." pageId="10" pageNumber="11">36</figureCitation>
): Leg spurs black; anteroventral surface of femora armed with long and robust spurs, 
<quantity id="FD1B559FFFCA066DFDACEBB0FDB6FA5A" box="[586,622,1512,1531]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="11" unit="in" value="5.0">5 in</quantity>
pro-, 
<quantity id="FD1B559FFFCA066DFD43EBB0FD12FA5A" box="[677,714,1512,1531]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="11" unit="in" value="4.0">4 in</quantity>
meso- and metathoracic femora, posteroventral surface with shorter and thinner spurs, 
<quantity id="FD1B559FFFCA066DFEB8E87CFE5EF996" box="[350,390,1572,1591]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="11" unit="in" value="3.0">3 in</quantity>
pro-, 
<quantity id="FD1B559FFFCA066DFE27E87CFE31F996" box="[449,489,1572,1591]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="11" unit="in" value="4.0">4 in</quantity>
meso- and 
<quantity id="FD1B559FFFCA066DFDBAE87CFD5CF996" box="[604,644,1572,1591]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="11" unit="in" value="6.0">6 in</quantity>
metathoracic femora; anteroventral surface of tibiae armed with 10 spurs in pro- (5 of tibial comb), 
<quantity id="FD1B559FFFCA066DFE40E838FE17F9D2" box="[422,463,1632,1651]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="11" unit="in" value="6.0">6 in</quantity>
meso- and metathoracic tibiae, usually longer than intervening spaces.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFF3DE8C3FAF6FF78" blockId="10.[157,781,198,1771]" lastBlockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">
Wings: MP reaches anal margin slightly distal of the vein descending from subnodus, covering 2 cells and ca 0.25 of a third cell in FW and 
<quantity id="FD1B559FFFCA066DFED5E88FFEB4F94B" box="[307,364,1751,1770]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="11" unit="in" value="0.5">0.5 in</quantity>
HW. Px in Fw 13; Hw 11; RP2 originating at or slightly proximal to Px 
<quantity id="FD1B559FFFCA066DFBA8EE9EFBADFF78" box="[1102,1141,198,217]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="11" unit="in" value="6.0">6 in</quantity>
Fw, at Px 
<quantity id="FD1B559FFFCA066DFB3FEE9EFAD8FF78" box="[1241,1280,198,217]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="11" unit="in" value="4.0">4 in</quantity>
Hw.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BEEBCFC76FE47" blockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">
Abdomen (
<figureCitation id="A2D8E4FFFFCA066DFC3CEEBBFB9DFF56" box="[986,1093,227,247]" captionStart="Figures 31–34" captionStartId="12.[159,223,1739,1759]" captionTargetBox="[244,1350,197,1720]" captionTargetId="figure-0@12.[244,1350,197,1720]" captionTargetPageId="12" captionText="Figures 31–34. Abdominal segments seven to ten (S7–10) of females of Forcepsioneura: (31–32)F.regua sp.nov., paratype (Brazil.RJ:Cachoeiras de Macacu, DZRJ); (33–34) F. serrabonita sp. nov., paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars for Figs 31–32 = 1 mm." pageId="10" pageNumber="11">Figs 33–34</figureCitation>
): Coloration very similar to that of male 
<typeStatus id="E55846D8FFCA066DFC80EF59FC67FEB4" box="[870,959,257,277]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
except for the yellow ivory posterior field of S8 and dorsal longitudinal stripe on S9–10, forming an irregular T-spot dorsally on S9. Sternites light yellow to orange-yellow. Ovipositor (
<figureCitation id="A2D8E4FFFFCA066DFC4CEF03FC37FECF" box="[938,1007,347,367]" captionStart="Figures 31–34" captionStartId="12.[159,223,1739,1759]" captionTargetBox="[244,1350,197,1720]" captionTargetId="figure-0@12.[244,1350,197,1720]" captionTargetPageId="12" captionText="Figures 31–34. Abdominal segments seven to ten (S7–10) of females of Forcepsioneura: (31–32)F.regua sp.nov., paratype (Brazil.RJ:Cachoeiras de Macacu, DZRJ); (33–34) F. serrabonita sp. nov., paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars for Figs 31–32 = 1 mm." pageId="10" pageNumber="11">Fig. 34</figureCitation>
) reaching distally level of cercus in lateral view, yellowish-orange, dorsal lobe of third valve (gonoplac of S9) slightly paler, ventral surface smooth; stylus dark brown, tip yellow. Cercus, epiproct and paraproct dark brown to black (
<figureCitation id="A2D8E4FFFFCA066DFCD1EF8AFC7BFE47" box="[823,931,466,486]" captionStart="Figures 31–34" captionStartId="12.[159,223,1739,1759]" captionTargetBox="[244,1350,197,1720]" captionTargetId="figure-0@12.[244,1350,197,1720]" captionTargetPageId="12" captionText="Figures 31–34. Abdominal segments seven to ten (S7–10) of females of Forcepsioneura: (31–32)F.regua sp.nov., paratype (Brazil.RJ:Cachoeiras de Macacu, DZRJ); (33–34) F. serrabonita sp. nov., paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars for Figs 31–32 = 1 mm." pageId="10" pageNumber="11">Figs 33–34</figureCitation>
).
</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BEFA9FAEDFD16" blockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">
Measurements (mm). Total length (incl. caudal appendages) 37.3; abdomen length (excluding caudal appendages) 31.0; head maximum width 3.1; Fw length 21.2; Hw length 19.8; Fw maximum width 3.8; Hw maximum width 3.9; Pt length in Fw 0.54; Pt length in Hw 0.59; length of metathoracic femur 2.2; metathoracic tibia 2.4; length of S9+ 
<quantity id="FD1B559FFFCA066DFB7FECDEFB12FD38" box="[1177,1226,646,665]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="11" unit="in" value="10.0">10 in</quantity>
lateral view 1.0; total length of cercus (dorsal branch) in lateral view 0.32.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BEC9AFBB1FCB1" blockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">
Variation in male 
<typeStatus id="E55846D8FFCA066DFBCFEC9AFB53FD74" box="[1065,1163,706,725]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
. The four 
<typeStatus id="E55846D8FFCA066DFB11EC9AFA45FD74" box="[1271,1437,706,725]" isOtherTypeStatus="true" pageId="10" pageNumber="11">other specimens</typeStatus>
(excluding the teneral one) are very similar to 
<typeStatus id="E55846D8FFCA066DFB1CEC87FA8EFD52" box="[1274,1366,735,755]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
, minor differences are described below.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BED43FB10FC07" blockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">Head: Labrum brownish to black. Postclypeus shining dark or with ill-defined mesal orange-brown spot. Frons and remainder of epicranium opaque black with a weak copper luster. Antennal flagellum brown to black. Posterior region of cranium (“postgena” plus “occiput”) ivory yellow.</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BEDE9FACAFBF8" blockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">
Thorax: Anteroventral surface of femora armed with long and robust spurs, 
<quantity id="FD1B559FFFCA066DFC38ED96FBDCFC40" box="[990,1028,974,993]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="11" unit="in" value="5.0">5 in</quantity>
pro-, 
<quantity id="FD1B559FFFCA066DFBD8ED97FBA4FC40" box="[1086,1148,974,994]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="10" pageNumber="11" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
meso- and 
<quantity id="FD1B559FFFCA066DFB0AED96FACBFC40" box="[1260,1299,974,993]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="11" unit="in" value="5.0">5 in</quantity>
metathoracic femora, posteroventral surface with shorter and thinner spurs, 
<quantity id="FD1B559FFFCA066DFA76EDB5FC9CFBBC" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="11" unit="in" value="3.0">3 in</quantity>
pro-, 
<quantity id="FD1B559FFFCA066DFC9DEA52FC6FFBBC" box="[891,951,1034,1053]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="10" pageNumber="11" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5 in</quantity>
meso-, 
<quantity id="FD1B559FFFCA066DFBE6EA52FBE4FBBC" box="[1024,1084,1034,1053]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="10" pageNumber="11" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
metathoracic femora; anteroventral surface of tibiae armed with 9–10 spurs in pro- (4–5 of tibial comb), 
<quantity id="FD1B559FFFCA066DFC9AEA1EFC62FBF8" box="[892,954,1094,1113]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="10" pageNumber="11" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
meso-, 
<quantity id="FD1B559FFFCA066DFBE1EA1EFB9EFBF8" box="[1031,1094,1094,1113]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="10" pageNumber="11" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
metathoracic tibiae.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BEA3BFBD6FB70" blockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">
Wings: MP reaches anal margin at or slightly distally to vein descending from subnodus, covering from 0.33 to 0.5 of a third cell in HW. Px in Fw 12–15; Hw 10–13; RP2 originating between Px 
<quantity id="FD1B559FFFCA066DFC4EEAE5FC3FFB71" box="[936,999,1213,1232]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="10" pageNumber="11" unit="in" value="5.5" valueMax="6.0" valueMin="5.0">5–6 in</quantity>
Fw.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BEA84FBA0FA7C" blockId="10.[815,1438,198,1771]" pageId="10" pageNumber="11">
Measurements (mm). Total length (incl. caudal appendages) 37.5–40.5; abdomen length (excluding caudal appendages) 31.7–33.7; head maximum width 2.9–3.3; Fw length 20.3–22.3; Hw length 19.5–21.3; Fw maximum width 3.5–3.8; Hw maximum width 3.7–3.8; Pt length 
<quantity id="FD1B559FFFCA066DFBB0EB0BFB16FAC7" box="[1110,1230,1363,1382]" metricMagnitude="-2" metricUnit="m" metricValue="1.4732" metricValueMax="1.524" metricValueMin="1.4224" pageId="10" pageNumber="11" unit="in" value="0.58" valueMax="0.6" valueMin="0.56">0.56–0.60 in</quantity>
Fw; 
<quantity id="FD1B559FFFCA066DFB1EEB0BFAA8FAC7" box="[1272,1392,1363,1382]" metricMagnitude="-2" metricUnit="m" metricValue="1.5493999999999999" metricValueMax="1.6256000000000002" metricValueMin="1.4731999999999998" pageId="10" pageNumber="11" unit="in" value="0.61" valueMax="0.64" valueMin="0.58">0.58–0.64 in</quantity>
Hw; length of metathoracic femur 1.9–2.1; metathoracic tibia 2.1–2.2; length of S9+ 
<quantity id="FD1B559FFFCA066DFC51EBD7FC33FA00" box="[951,1003,1422,1442]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="11" unit="in" value="10.0">10 in</quantity>
lateral view 1.3–1.4; total length of cercus (dorsal branch) in lateral view 0.74–0.86; length of ventrobasal process in lateral view 0.12–0.22.
</paragraph>
<paragraph id="3A5CF87AFFCA066DFC8BEBB1FBC3FA5A" blockId="10.[815,1438,198,1771]" box="[877,1051,1512,1532]" pageId="10" pageNumber="11">Larva. Unknown.</paragraph>
<paragraph id="3A5CF87AFFCA066CFC8BE85EFE30FA4F" blockId="10.[815,1438,198,1771]" lastBlockId="11.[149,772,1260,1787]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<materialsCitation id="8A8BF227FFCA066CFC8BE85EFE30FA4F" collectingDate="2012-02-27" collectingDateMax="2015-02-26" collectingDateMin="2012-02-27" collectionCode="RPPN, USNM, MZSP, MNRJ" collectorName="R. A. Carvalho &amp; C. M. Flint &amp; O. S. Flint Jr. &amp; T. W. Donnelly &amp; O. S. Flint &amp; Flint, Jr." country="Brazil" county="Material" elevation="810" lastPageId="11" lastPageNumber="12" latitude="-15.383889" location="Reserva Particular do Patrimonio Natural" longLatPrecision="21" longitude="-39.571945" municipality="Camacan" pageId="10" pageNumber="11" specimenCode="DZRJ 0050, ENT2857" specimenCount="13" specimenCount-female="2" specimenCount-male="11" stateProvince="Bahia" typeStatus="holotype">
<collectingCounty id="D33D80F6FFCA066DFC8BE85EFC67F9B8" box="[877,959,1542,1561]" pageId="10" pageNumber="11">Material</collectingCounty>
examined (
<specimenCount id="2CE533F3FFCA066DFBDFE85EFB51F9B8" box="[1081,1161,1542,1561]" count="6" pageId="10" pageNumber="11" type="male">6 males</specimenCount>
, 
<specimenCount id="2CE533F3FFCA066DFB73E85EFB37F9B8" box="[1173,1263,1542,1561]" count="1" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
). 
<typeStatus id="E55846D8FFCA066DFAE7E85EFA85F9BB" box="[1281,1373,1542,1562]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
male, 
<collectingCountry id="42F4B8EAFFCA066DFCD6E87CFCA5F996" box="[816,893,1572,1591]" name="Brazil" pageId="10" pageNumber="11">BRAZIL</collectingCountry>
. 
<collectingRegion id="F8273698FFCA066DFC60E87CFC2FF996" box="[902,1015,1572,1591]" country="Brazil" name="Bahia" pageId="10" pageNumber="11">Bahia State</collectingRegion>
: 
<collectingMunicipality id="DA386200FFCA066DFBE6E87CFB85F996" box="[1024,1117,1572,1591]" pageId="10" pageNumber="11">Camacan</collectingMunicipality>
municipality, 
<location id="3F3CAEA1FFCA066DFB0BE87CFBC0F9F4" LSID="urn:lsid:plazi:treatment:B24A496CFFC6066CFC7FE862FA4DFAD6:3F3CAEA1FFCA066DFB0BE87CFBC0F9F4" country="Brazil" county="Material" latitude="-15.383889" longLatPrecision="21" longitude="-39.571945" municipality="Camacan" name="Reserva Particular do Patrimonio Natural" pageId="10" pageNumber="11" stateProvince="Bahia">Reserva Particular do Patrimônio Natural</location>
(
<collectionCode id="5CF260BFFFCA066DFBC1E81AFBB9F9F4" box="[1063,1121,1602,1621]" pageId="10" pageNumber="11">RPPN</collectionCode>
) 
<location id="3F3CAEA1FFCA066DFB96E81AFB36F9F4" LSID="urn:lsid:plazi:treatment:B24A496CFFC6066CFC7FE862FA4DFAD6:3F3CAEA1FFCA066DFB96E81AFB36F9F4" box="[1136,1262,1602,1621]" country="Brazil" county="Material" latitude="-15.383889" longLatPrecision="21" longitude="-39.571945" municipality="Camacan" name="Serra Bonita" pageId="10" pageNumber="11" stateProvince="Bahia">Serra Bonita</location>
, collecting point “Alojamento” (BA 2012-001; 
<geoCoordinate id="5FD79EBDFFCA066DFBBCE807FB11F9D3" box="[1114,1225,1631,1650]" degrees="15" direction="south" minutes="23" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="16" value="-15.387777">15°23’16”S</geoCoordinate>
, 
<geoCoordinate id="5FD79EBDFFCA066DFB30E807FA96F9D3" box="[1238,1358,1631,1650]" degrees="39" direction="west" minutes="33" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="58" value="-39.56611">39°33’58”W</geoCoordinate>
, 
<elevation id="B1CE1F49FFCA066DFABDE838FC85F931" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="10" pageNumber="11" unit="m" value="810.0">
<quantity id="FD1B559FFFCA066DFABDE838FA44F9D2" box="[1371,1436,1632,1651]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="10" pageNumber="11" unit="m" value="810.0">810 m</quantity>
a.s.l.
</elevation>
), 
<date id="4E5DDEBAFFCA066DFC92E826FC26F930" box="[884,1022,1662,1681]" pageId="10" pageNumber="11" value="2012-02-27" valueMax="2012-02-29" valueMin="2012-02-27">
<collectingDate id="5E192752FFCA066DFC92E826FC26F930" box="[884,1022,1662,1681]" pageId="10" pageNumber="11" value="2012-02-27" valueMax="2012-02-29" valueMin="2012-02-27">27–29.II.2012</collectingDate>
</date>
, 
<collectorName id="97169DACFFCA066DFBEDE826FB4FF931" box="[1035,1175,1661,1681]" pageId="10" pageNumber="11">R.A. Carvalho</collectorName>
leg. (
<specimenCode id="6A455001FFCA066DFB34E826FA9BF931" box="[1234,1347,1661,1681]" collectionCode="DZRJ" pageId="10" pageNumber="11">DZRJ 0050</specimenCode>
); 
<specimenCount id="2CE533F3FFCA066DFAB1E826FA44F931" box="[1367,1436,1661,1681]" count="1" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
<typeStatus id="E55846D8FFCA066DFCD6E8C4FC51F90E" box="[816,905,1692,1711]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
, same data but first trickle on road beyond managers house (
<geoCoordinate id="5FD79EBDFFCA066DFC9FE8E1FC3FF96D" box="[889,999,1721,1740]" degrees="15" direction="south" minutes="23" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="19" value="-15.388611">15°23’19”S</geoCoordinate>
, 
<geoCoordinate id="5FD79EBDFFCA066DFC16E8E1FBB1F96D" box="[1008,1129,1721,1740]" degrees="39" direction="west" minutes="33" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="57" value="-39.565834">39°33’57”W</geoCoordinate>
, 
<elevation id="B1CE1F49FFCA066DFB94E8E1FB3AF96D" box="[1138,1250,1721,1741]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="10" pageNumber="11" unit="m" value="710.0">
<quantity id="FD1B559FFFCA066DFB94E8E1FB68F96C" box="[1138,1200,1721,1741]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="10" pageNumber="11" unit="m" value="710.0">710 m</quantity>
a.s.l.
</elevation>
[
<quantity id="FD1B559FFFCA066DFB0BE8E1FAF7F96D" box="[1261,1327,1721,1740]" metricMagnitude="2" metricUnit="m" metricValue="8.22" pageId="10" pageNumber="11" unit="m" value="822.0">
<elevation id="B1CE1F49FFCA066DFB0BE8E1FAF7F96D" box="[1261,1327,1721,1740]" metricMagnitude="2" metricUnit="m" metricValue="8.22" pageId="10" pageNumber="11" unit="m" value="822.0">822 m</elevation>
</quantity>
]), 
<date id="4E5DDEBAFFCA066DFAA0E8E1FA41F96D" box="[1350,1433,1721,1740]" pageId="10" pageNumber="11" value="2013-01-05">
<collectingDate id="5E192752FFCA066DFAA0E8E1FA41F96D" box="[1350,1433,1721,1740]" pageId="10" pageNumber="11" value="2013-01-05">5.I.2013</collectingDate>
</date>
, 
<collectorName id="97169DACFFCA066DFCC9E88FFC4CF94B" box="[815,916,1751,1770]" pageId="10" pageNumber="11">C.M. Flint</collectorName>
&amp; 
<collectorName id="97169DACFFCA066DFC54E88FFBF3F94B" box="[946,1067,1751,1770]" pageId="10" pageNumber="11">O.S. Flint Jr.</collectorName>
leg. (
<collectionCode id="5CF260BFFFCA066DFBB9E88FFB7DF94B" box="[1119,1189,1751,1770]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
); 
<specimenCount id="2CE533F3FFCA066DFB53E88FFB2FF94B" box="[1205,1271,1751,1770]" count="2" pageId="10" pageNumber="11" type="male">2 male</specimenCount>
<typeStatus id="E55846D8FFCA066DFB18E88FFA87F94B" box="[1278,1375,1751,1770]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
, same data but quarry, 
<date id="4E5DDEBAFFCB066CFEABEAB4FE6EFB5E" box="[333,438,1260,1279]" pageId="11" pageNumber="12" value="2015-02-20">
<collectingDate id="5E192752FFCB066CFEABEAB4FE6EFB5E" box="[333,438,1260,1279]" pageId="11" pageNumber="12" value="2015-02-20">20.II.2015</collectingDate>
</date>
(
<collectionCode id="5CF260BFFFCB066CFE21EAB4FDD6FB5E" box="[455,526,1260,1279]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12" type="Museum">USNM</collectionCode>
; 
<collectionCode id="5CF260BFFFCB066CFDFDEAB4FD8DFB5E" box="[539,597,1260,1279]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="12" type="Museum">MZSP</collectionCode>
ODO-492, DNA voucher 
<specimenCode id="6A455001FFCB066CFF09EB52FE96FABC" box="[239,334,1290,1309]" collectionCode="ENT" country="Uganda" lsid="urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="11" pageNumber="12" type="Herbarium">ENT2857</specimenCode>
); 
<specimenCount id="2CE533F3FFCB066CFEB8EB52FE7AFABC" box="[350,418,1290,1309]" count="1" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
<typeStatus id="E55846D8FFCB066CFE4FEB52FDDAFABC" box="[425,514,1290,1309]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
, same data but 
<date id="4E5DDEBAFFCB066CFD78EB52FCDBFABC" box="[670,771,1290,1309]" pageId="11" pageNumber="12" value="2015-02-26">
<collectingDate id="5E192752FFCB066CFD78EB52FCDBFABC" box="[670,771,1290,1309]" pageId="11" pageNumber="12" value="2015-02-26">26.II.2015</collectingDate>
</date>
(
<collectionCode id="5CF260BFFFCB066CFF78EB70FF3CFA9A" box="[158,228,1320,1339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12" type="Museum">USNM</collectionCode>
); 
<specimenCount id="2CE533F3FFCB066CFF15EB70FE9EFA9A" box="[243,326,1320,1339]" count="1" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
<typeStatus id="E55846D8FFCB066CFEADEB70FE7CFA9A" box="[331,420,1320,1339]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
, same data but spring near telecommunication towers (
<geoCoordinate id="5FD79EBDFFCB066CFE87EB1EFE08FAF8" box="[353,464,1350,1369]" degrees="15" direction="south" minutes="23" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="02" value="-15.383889">15°23’02”S</geoCoordinate>
, 
<geoCoordinate id="5FD79EBDFFCB066CFE3CEB1EFD8CFAF8" box="[474,596,1350,1369]" degrees="39" direction="west" minutes="34" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="19" value="-39.571945">39°34’19”W</geoCoordinate>
; 
<elevation id="B1CE1F49FFCB066CFDB8EB1EFD09FAF8" box="[606,721,1350,1369]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="12" unit="m" value="850.0">
<quantity id="FD1B559FFFCB066CFDB8EB1EFD46FAF8" box="[606,670,1350,1369]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="12" unit="m" value="850.0">850 m</quantity>
a.s.l.
</elevation>
[
<quantity id="FD1B559FFFCB066CFD3BEB1EFF76FAD6" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="11" pageNumber="12" unit="m" value="880.0">
<elevation id="B1CE1F49FFCB066CFD3BEB1EFF76FAD6" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="11" pageNumber="12" unit="m" value="880.0">880 m</elevation>
</quantity>
]), 
<date id="4E5DDEBAFFCB066CFF22EB3BFEE2FAD7" box="[196,314,1379,1398]" pageId="11" pageNumber="12" value="2013-12-25">
<collectingDate id="5E192752FFCB066CFF22EB3BFEE2FAD7" box="[196,314,1379,1398]" pageId="11" pageNumber="12" value="2013-12-25">25.XII.2013</collectingDate>
</date>
, 
<collectorName id="97169DACFFCB066CFEA4EB3CFE15FAD6" box="[322,461,1379,1399]" pageId="11" pageNumber="12">T.W. Donnelly</collectorName>
leg. (
<collectionCode id="5CF260BFFFCB066CFE1BEB3CFDE0FAD6" box="[509,568,1380,1399]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ</collectionCode>
ODO-0001; ex-TWD collection). Additional specimen (excluded from type series). 
<specimenCount id="2CE533F3FFCB066CFD13EBD9FF1FFA13" count="1" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
(teneral), same data as 
<typeStatus id="E55846D8FFCB066CFE4EEBC7FDD9FA12" box="[424,513,1439,1459]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
but trickles at upper water tank (15°23’03”, 
<geoCoordinate id="5FD79EBDFFCB066CFEA7EBE5FE62FA71" box="[321,442,1469,1488]" degrees="39" direction="west" minutes="34" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="19" value="-39.571945">39°34’19”W</geoCoordinate>
, 
<elevation id="B1CE1F49FFCB066CFE23EBE5FDE1FA71" box="[453,569,1469,1489]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="11" pageNumber="12" unit="m" value="890.0">
<quantity id="FD1B559FFFCB066CFE23EBE5FDDDFA70" box="[453,517,1469,1489]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="11" pageNumber="12" unit="m" value="890.0">890 m</quantity>
a.s.l.
</elevation>
), 
<date id="4E5DDEBAFFCB066CFDA8EBE5FD1EFA71" box="[590,710,1469,1488]" pageId="11" pageNumber="12" value="2012-12-23">
<collectingDate id="5E192752FFCB066CFDA8EBE5FD1EFA71" box="[590,710,1469,1488]" pageId="11" pageNumber="12" value="2012-12-23">23.XII.2012</collectingDate>
</date>
, 
<collectorName id="97169DACFFCB066CFD37EBE5FF1FFA4F" pageId="11" pageNumber="12">C.M. Flint</collectorName>
&amp; 
<collectorName id="97169DACFFCB066CFF02EB83FE9CFA4F" box="[228,324,1499,1518]" pageId="11" pageNumber="12">O.S. Flint</collectorName>
, Jr. leg. (
<collectionCode id="5CF260BFFFCB066CFE71EB83FE05FA4F" box="[407,477,1499,1518]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="6E9CA8F2FFCB066CFF71EA3DFA9AFB36" pageId="11" pageNumber="12" startId="11.[151,218,1125,1145]" targetBox="[151,1428,197,1105]" targetPageId="11" targetType="figure">
<paragraph id="3A5CF87AFFCB066CFF71EA3DFA9AFB36" blockId="11.[151,1429,1124,1175]" pageId="11" pageNumber="12">
Figures 27–30. Head and prothorax in dorsal view of females of 
<taxonomicName id="FDE383F9FFCB066CFD09EA3CFCACFBD9" authorityName="Lencioni" authorityYear="1999" box="[751,884,1124,1144]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="08972468FFCB066CFD09EA3CFCACFBD9" box="[751,884,1124,1144]" italics="true" pageId="11" pageNumber="12">Forcepsioneura</emphasis>
</taxonomicName>
: (27–28) 
<taxonomicName id="FDE383F9FFCB066CFC37EA3CFBCFFBD8" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[977,1047,1124,1145]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="regua" status="sp. nov.">
<emphasis id="08972468FFCB066CFC37EA3CFBCFFBD8" box="[977,1047,1124,1145]" italics="true" pageId="11" pageNumber="12">F. regua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFCB066CFBF8EA3DFBB3FBD8" box="[1054,1131,1125,1145]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (29–30) 
<taxonomicName id="FDE383F9FFCB066CFE33EADAFD96FB37" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[469,590,1154,1174]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFCB066CFE33EADAFD96FB37" box="[469,590,1154,1174]" italics="true" pageId="11" pageNumber="12">F. serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFCB066CFDB2EADBFD7AFB36" box="[596,674,1155,1175]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (Brazil. BA: Camacan, MNRJ ODO-0001). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="3A5CF87AFFCB066CFF32EBA0FD67F922" blockId="11.[149,772,1260,1787]" pageId="11" pageNumber="12">
Etymology. Noun in apposition in reference to the 
<typeStatus id="E55846D8FFCB066CFD3EEBA1FCDBF9AD" box="[728,771,1529,1548]" pageId="11" pageNumber="12">type</typeStatus>
locality in the complex of RPPNs Serra Bonita, in honor of this protected area and its managers, who have taken positive steps towards the preservation of Atlantic Forest remnants in the southern portion of the biodiversity-rich state of 
<collectingRegion id="F8273698FFCB066CFD66E828FD63F922" box="[640,699,1648,1667]" country="Brazil" name="Bahia" pageId="11" pageNumber="12">Bahia</collectingRegion>
.
</paragraph>
<paragraph id="3A5CF87AFFCB066CFF32E8D6FA4DFAD6" blockId="11.[149,772,1260,1787]" lastBlockId="11.[807,1429,1260,1399]" pageId="11" pageNumber="12">
Biological and ecological data. Expected to be a submontane forest species, most likely occurring at seepages similar to the mesohabitat occupied by the large montane species from southeastern 
<collectingCountry id="42F4B8EAFFCB066CFEFBE8B0FE8CF95A" box="[285,340,1768,1787]" name="Brazil" pageId="11" pageNumber="12">Brazil</collectingCountry>
(e.g., 
<taxonomicName id="FDE383F9FFCB066CFE74E8B0FE2CF95B" baseAuthorityName="Santos" baseAuthorityYear="1970" box="[402,500,1768,1786]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="itatiaiae">
<emphasis id="08972468FFCB066CFE74E8B0FE2CF95B" box="[402,500,1768,1786]" italics="true" pageId="11" pageNumber="12">F. itatiaiae</emphasis>
</taxonomicName>
and 
<taxonomicName id="FDE383F9FFCB066CFDC1E8B0FD50F95A" authorityName="Machado" authorityYear="2000" box="[551,648,1768,1787]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" isUncertain="true" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lucia">
<emphasis id="08972468FFCB066CFDC1E8B0FDE2F95B" box="[551,570,1768,1786]" italics="true" pageId="11" pageNumber="12">F.</emphasis>
cf. 
<emphasis id="08972468FFCB066CFDBDE8B0FD50F95A" box="[603,648,1768,1787]" italics="true" pageId="11" pageNumber="12">lucia</emphasis>
</taxonomicName>
). The larvae possibly are madicolous and should inhabit shallow streams running over rocky substrates covered by ferns and hornworts/ liverworts forming small waterfalls (
<figureCitation id="A2D8E4FFFFCB066CFB99EB7FFB19FA9A" box="[1151,1217,1319,1339]" captionStart="Figures 35–40" captionStartId="13.[151,218,1575,1595]" captionTargetBox="[151,1428,212,1555]" captionTargetId="figure-105@13.[151,1428,197,1555]" captionTargetPageId="13" captionText="Figures 35–40. Habitus and habitat of species of Forcepsioneura: (35) F. regua sp. nov., holotype (Brazil. RJ: Cachoeiras de Macacu, DZRJ 2071); (36) F. serrabonita sp. nov., holotype (Brazil. BA: Camacan, DZRJ 0050), (37) F. regua sp. nov., live individual at type locality at Reserva Ecológica de Guapiaçu (REGUA); (38) overview of type locality at RPPN Serra Bonita; (39) shallow muddy and slow stream at Itatiaia National Park, typical habitat of small light blue species; (40) seepage covered by hornworts at Três Picos State Park, typical habitat of large orange-black species. Scale bars = 1 mm. Photos: (37) TMFK; (38) RA Carvalho; (39–40) APP." pageId="11" pageNumber="12">Fig.40</figureCitation>
). Though we have not observed adult behavior in the field it is plausible to assume that 
<taxonomicName id="FDE383F9FFCB066CFCC1EB3CFC47FAD7" authorityName="Pinto &amp; Kompier" authorityYear="2018" box="[807,927,1379,1398]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="serrabonita" status="sp. nov.">
<emphasis id="08972468FFCB066CFCC1EB3CFC47FAD7" box="[807,927,1379,1398]" italics="true" pageId="11" pageNumber="12">F.serrabonita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13A49913FFCB066CFC42EB3CFC37FAD6" box="[932,1007,1380,1399]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
is a forest damselfly like most 
<taxonomicName id="FDE383F9FFCB066CFAEFEB3CFA55FAD7" authorityName="Lencioni" authorityYear="1999" box="[1289,1421,1380,1398]" class="Insecta" family="Protoneuridae" genus="Forcepsioneura" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="08972468FFCB066CFAEFEB3CFA55FAD7" box="[1289,1421,1380,1398]" italics="true" pageId="11" pageNumber="12">Forcepsioneura</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>