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<document id="09A564EFFEC58A55BB9218555A764ED7" ID-CLB-Dataset="30156" ID-DOI="10.11646/zootaxa.4415.1.6" ID-GBIF-Dataset="c7b9429a-8520-49c7-b8f0-f62ccac35ad9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1241858" ID-ZooBank="FE0C98F6-E952-4FE9-82DE-37F13BD1C703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525599103729" checkinUser="plazi" docAuthor="Soulier-Perkins, Adeline, Navasero, Mario, Yap, Sheryl &amp; Wilson, Michael R." docDate="2018" docId="B20A87ECFFCDFFD6D7858584FB38FE85" docLanguage="en" docName="zootaxa.4415.1.6.pdf" docOrigin="Zootaxa 4415 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Virgilia cocovora Soulier-Perkins &amp; Navasero &amp; Yap &amp; Wilson 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="140" masterDocId="4E33FF94FFCFFFD3D7128720FF93FF98" masterDocTitle="New species for Virgilia Stål, 1870 (Hemiptera, Lophopidae), genus review and key to species" masterLastPageNumber="149" masterPageNumber="135" pageNumber="137" updateTime="1698541229410" updateUser="ExternalLinkService">
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<mods:title id="AA842C4828909DFAA7E71AF1A25A0B93">New species for Virgilia Stål, 1870 (Hemiptera, Lophopidae), genus review and key to species</mods:title>
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<mods:namePart id="D55787B5E4295BC4B55EA3913220CF95">Soulier-Perkins, Adeline</mods:namePart>
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<mods:namePart id="7FBFF3A0861AC400141742CDF1F7221D">Navasero, Mario</mods:namePart>
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<mods:namePart id="7E79F7521CED5EB0AA2A551D89E91020">Yap, Sheryl</mods:namePart>
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<mods:namePart id="BBF361A9134B82A6DFCD9D6FD042CD05">Wilson, Michael R.</mods:namePart>
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<mods:title id="21794C0076395832B127542AF965EE80">Zootaxa</mods:title>
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<mods:date id="370468FEA8CCFC78CD8D5CAF7D59D58B">2018</mods:date>
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<mods:number id="5D5B360BE04C1CDB6D10DF2FBCADF95D">2018-04-27</mods:number>
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<treatment id="B20A87ECFFCDFFD6D7858584FB38FE85" ID-DOI="http://doi.org/10.5281/zenodo.5946530" ID-GBIF-Taxon="143811240" ID-Zenodo-Dep="5946530" LSID="urn:lsid:plazi:treatment:B20A87ECFFCDFFD6D7858584FB38FE85" httpUri="http://treatment.plazi.org/id/B20A87ECFFCDFFD6D7858584FB38FE85" lastPageId="5" lastPageNumber="140" pageId="2" pageNumber="137">
<subSubSection id="72B96571FFCDFFD1D7858584FE8AFD78" pageId="2" pageNumber="137" type="nomenclature">
<paragraph id="3A1C36FAFFCDFFD1D7858584FD06FD27" blockId="2.[151,661,676,736]" box="[151,661,676,703]" pageId="2" pageNumber="137">
<heading id="61548196FFCDFFD1D7858584FD06FD27" bold="true" box="[151,661,676,703]" fontSize="11" level="1" pageId="2" pageNumber="137" reason="1">
<emphasis id="08D7EAE8FFCDFFD1D7858584FD06FD27" bold="true" box="[151,661,676,703]" pageId="2" pageNumber="137">
<taxonomicName id="FDA34D79FFCDFFD1D7858584FDA3FD27" ID-CoL="5BJSL" authority="Soulier-Perkins" authorityName="Soulier-Perkins &amp; Navasero &amp; Yap &amp; Wilson" authorityYear="2018" box="[151,560,676,703]" class="Insecta" family="Lophopidae" genus="Virgilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="cocovora" status="sp. nov.">
<emphasis id="08D7EAE8FFCDFFD1D7858584FEF5FD26" bold="true" box="[151,358,676,702]" italics="true" pageId="2" pageNumber="137">Virgilia cocovora</emphasis>
Soulier-Perkins
</taxonomicName>
<taxonomicNameLabel id="13E45793FFCDFFD1D5258585FD06FD27" box="[567,661,677,703]" pageId="2" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3A1C36FAFFCDFFD1D78585E7FE8AFD78" blockId="2.[151,661,676,736]" box="[151,281,711,736]" pageId="2" pageNumber="137">
(
<figureCitation id="A2982A7FFFCDFFD1D78D85E7FE82FD78" box="[159,273,711,736]" captionStart-0="FIGURES 511" captionStart-1="FIGURES 1234" captionStartId-0="3.[151,264,1940,1963]" captionStartId-1="4.[151,264,1507,1530]" captionTargetBox-0="[228,1355,203,1921]" captionTargetBox-1="[314,1269,197,1483]" captionTargetId-0="figure@3.[218,1371,193,1921]" captionTargetId-1="figure@4.[313,1273,193,1486]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 511. Virgilia cocovora n. sp., holotype. 5 Habitus, dorsal view. 6 Habitus, lateral view. 7 Frons and clypeus, anterior view. 8 Labels. 9 Anal tube, pygopher and gonostylus, lateral view. 10 Aedeagus s.l., lateral view. 11 Aedeagus s.s., lateral view." captionText-1="FIGURES 1234. Virgilia cocovora n. sp., paratypes. 1216 female MNHN(EH)23545 12 Habitus, dorsal view. 13 Habitus, ventral view 14 Habitus, lateral view 15 Frons and clypeus, anterior view 16 Labels. 1721 male 17 Habitus, dorsal view. 18 Habitus, ventral view 19 Habitus, lateral view 20 Frons and clypeus, anterior view 21 Labels. 2225 female 22 Habitus, dorsal view 23 Habitus, lateral view 24 Frons and clypeus, anterior view 25 Labels. 2629 female MNHN(EH)235457 26 Habitus, dorsal view 27 Habitus, lateral view 28 Frons and clypeus, anterior view 29 Labels. 3034 female MNHN(EH)23548 30 Habitus, dorsal view. 31Habitus, ventral view 32Habitus, lateral view 33 Labels34 Frons and clypeus, anterior view." httpUri-0="https://zenodo.org/record/1241862/files/figure.png" httpUri-1="https://zenodo.org/record/1241864/files/figure.png" pageId="2" pageNumber="137">Figs 534</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B96571FFCDFFD1D785842FFE05FC20" pageId="2" pageNumber="137" type="materials_examined">
<paragraph id="3A1C36FAFFCDFFD1D785842FFB28FCD4" blockId="2.[151,1436,783,2032]" pageId="2" pageNumber="137">
<emphasis id="08D7EAE8FFCDFFD1D785842FFEDEFCB0" bold="true" box="[151,333,783,808]" pageId="2" pageNumber="137">
<typeStatus id="E5188858FFCDFFD1D785842FFF41FCB0" box="[151,210,783,808]" pageId="2" pageNumber="137">Type</typeStatus>
material.
</emphasis>
Male
<typeStatus id="E5188858FFCDFFD1D6B1842FFD99FCB0" box="[419,522,783,808]" pageId="2" pageNumber="137" type="holotype">holotype</typeStatus>
:
<collectingCountry id="42B4766AFFCDFFD1D50E842FFD0CFCB0" box="[540,671,783,808]" name="Philippines" pageId="2" pageNumber="137">Philippines</collectingCountry>
: Mindanao,
<collectingRegion id="F867F818FFCDFFD1D42A842FFB85FCB0" box="[824,1046,783,808]" country="Philippines" name="Compostela" pageId="2" pageNumber="137">Compostela Valley</collectingRegion>
, New
<collectingRegion id="F867F818FFCDFFD1D37E8430FB2EFCBF" box="[1132,1213,784,807]" country="Philippines" name="Bataan" pageId="2" pageNumber="137">Bataan</collectingRegion>
, Camanlangan, on coconut,
<date id="4E1D103AFFCDFFD1D7EF8413FEF1FCD4" box="[253,354,819,844]" pageId="2" pageNumber="137" value="2016-02-03">3.2.2016</date>
, M.V. Navasero &amp; G.A.S. Burgonio rec &amp; leg,
<collectionCode id="5CB2AE3FFFCDFFD1D4628413FC5BFCD4" box="[880,968,819,844]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="137">MNHN</collectionCode>
(EH)23546 (
<collectionCode id="5CB2AE3FFFCDFFD1D3418414FB3CFCD4" box="[1107,1199,820,844]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="137">MNHN</collectionCode>
).
</paragraph>
<paragraph id="3A1C36FAFFCDFFD1D7D58478FE05FC20" blockId="2.[151,1436,783,2032]" pageId="2" pageNumber="137">
Paratypes: 1♂?, 4♀?.
<materialsCitation id="8ACB3CA7FFCDFFD1D6D98477FE05FC20" ID-GBIF-Occurrence="1841140032" collectingDate="2016-02-03" collectionCode="MNHN" collectorName="M. V. Navasero &amp; G. A. S. Burgonio" country="Philippines" location="Camanlangan" municipality="Mindanao" pageId="2" pageNumber="137" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Compostela Valley">
<collectingCountry id="42B4766AFFCDFFD1D6D98477FDDFFCE8" box="[459,588,855,880]" name="Philippines" pageId="2" pageNumber="137">Philippines</collectingCountry>
,
<collectingMunicipality id="DA78AC80FFCDFFD1D5498477FD43FCE8" box="[603,720,855,880]" pageId="2" pageNumber="137">Mindanao</collectingMunicipality>
,
<collectingRegion id="F867F818FFCDFFD1D5CD8477FC2AFCE8" box="[735,953,855,880]" country="Philippines" name="Compostela" pageId="2" pageNumber="137">Compostela Valley</collectingRegion>
, New
<collectingRegion id="F867F818FFCDFFD1D31B8478FBC9FCF7" box="[1033,1114,856,879]" country="Philippines" name="Bataan" pageId="2" pageNumber="137">Bataan</collectingRegion>
,
<location id="3F7C6021FFCDFFD1D37B8477FA95FCE8" LSID="urn:lsid:plazi:treatment:B20A87ECFFCDFFD6D7858584FB38FE85:3F7C6021FFCDFFD1D37B8477FA95FCE8" box="[1129,1286,855,880]" country="Philippines" municipality="Mindanao" name="Camanlangan" pageId="2" pageNumber="137" stateProvince="Compostela Valley">Camanlangan</location>
, on coconut,
<collectingDate id="5E59E9D2FFCDFFD1D785845BFF68FC0C" box="[151,251,891,916]" pageId="2" pageNumber="137" value="2016-02-03">
<date id="4E1D103AFFCDFFD1D785845BFF68FC0C" box="[151,251,891,916]" pageId="2" pageNumber="137" value="2016-02-03">3.2.2016</date>
</collectingDate>
,
<collectorName id="9756532CFFCDFFD1D615845CFE3CFC0B" box="[263,431,892,915]" pageId="2" pageNumber="137">M.V. Navasero</collectorName>
&amp;
<collectorName id="9756532CFFCDFFD1D6C0845CFD1FFC0C" box="[466,652,891,916]" pageId="2" pageNumber="137">G.A.S. Burgonio</collectorName>
rec. &amp; leg.,
<specimenCount id="2CA5FD73FFCDFFD1D404845CFCABFC0C" box="[790,824,892,916]" pageId="2" pageNumber="137" type="female">3♀</specimenCount>
?
<collectionCode id="5CB2AE3FFFCDFFD1D45F845CFC36FC0C" box="[845,933,892,916]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="137">MNHN</collectionCode>
(EH) 23545, 23547 and 23548 (
<collectionCode id="5CB2AE3FFFCDFFD1D217845CFACDFC0C" box="[1285,1374,892,916]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="137">MNHN</collectionCode>
)
<specimenCount id="2CA5FD73FFCDFFD1D27F845CFA03FC0C" box="[1389,1424,892,916]" pageId="2" pageNumber="137" type="male">1♂</specimenCount>
? &amp;
<specimenCount id="2CA5FD73FFCDFFD1D7A68480FF40FC20" box="[180,211,928,952]" pageId="2" pageNumber="137" type="female">1♀</specimenCount>
? (MNH-UPLB).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="72B96571FFCDFFD1D7D584E3FC8AFC44" box="[199,793,963,988]" pageId="2" pageNumber="137" type="distribution">
<paragraph id="3A1C36FAFFCDFFD1D7D584E3FC8AFC44" blockId="2.[151,1436,783,2032]" box="[199,793,963,988]" pageId="2" pageNumber="137">
<emphasis id="08D7EAE8FFCDFFD1D7D584E3FECAFC44" bold="true" box="[199,345,963,988]" pageId="2" pageNumber="137">Distribution</emphasis>
.
<collectingCountry id="42B4766AFFCDFFD1D67584E3FE7BFC44" box="[359,488,963,988]" name="Philippines" pageId="2" pageNumber="137">Philippines</collectingCountry>
, Mindanao Island (
<figureCitation id="A2982A7FFFCDFFD1D5AD84E3FC82FC44" box="[703,785,963,988]" captionStart="FIGURE 65" captionStartId="12.[151,250,1765,1787]" captionTargetBox="[151,1436,183,1756]" captionTargetId="figure@12.[151,1436,183,1756]" captionTargetPageId="12" captionText="FIGURE 65.Distribution map." httpUri="https://zenodo.org/record/1241880/files/figure.png" pageId="2" pageNumber="137">Fig. 65</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B96571FFCDFFD1D7D584C7FE7EFBF4" pageId="2" pageNumber="137" type="diagnosis">
<paragraph id="3A1C36FAFFCDFFD1D7D584C7FE7EFBF4" blockId="2.[151,1436,783,2032]" pageId="2" pageNumber="137">
<emphasis id="08D7EAE8FFCDFFD1D7D584C7FED2FB98" bold="true" box="[199,321,999,1024]" pageId="2" pageNumber="137">Diagnosis.</emphasis>
Easy to recognise from the other three
<taxonomicName id="FDA34D79FFCDFFD1D40384C7FCF6FC67" authorityName="Stal" authorityYear="1870" box="[785,869,999,1023]" class="Insecta" family="Lophopidae" genus="Virgilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="08D7EAE8FFCDFFD1D40384C7FCF6FC67" box="[785,869,999,1023]" italics="true" pageId="2" pageNumber="137">Virgilia</emphasis>
</taxonomicName>
by the coloration of its frons, it bears six brown spots (
<figureCitation id="A2982A7FFFCDFFD1D7F3832BFEB6FBBC" box="[225,293,1035,1060]" captionStart="FIGURES 13" captionStartId="2.[151,264,584,607]" captionTargetBox="[151,1436,273,562]" captionTargetId="figure@2.[151,1436,273,570]" captionTargetPageId="2" captionText="FIGURES 13. Schematic patterns on Virgilia frons. 1 V. luzonensis. 2 V.cocovora.3 V.nigropicta.4 V.imuganensis." httpUri="https://zenodo.org/record/1241860/files/figure.png" pageId="2" pageNumber="137">Fig. 2</figureCitation>
) where the
<taxonomicName id="FDA34D79FFCDFFD1D6BE832CFDAEFBBC" authorityName="Baker" authorityYear="1925" box="[428,573,1036,1060]" class="Insecta" family="Lophopidae" genus="Virgilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="08D7EAE8FFCDFFD1D6BE832CFDAEFBBC" box="[428,573,1036,1060]" italics="true" pageId="2" pageNumber="137">V. luzonensis</emphasis>
</taxonomicName>
presents two patches and a stripe (
<figureCitation id="A2982A7FFFCDFFD1D4D8832BFB9CFBBC" box="[970,1039,1035,1060]" captionStart="FIGURES 13" captionStartId="2.[151,264,584,607]" captionTargetBox="[151,1436,273,562]" captionTargetId="figure@2.[151,1436,273,570]" captionTargetPageId="2" captionText="FIGURES 13. Schematic patterns on Virgilia frons. 1 V. luzonensis. 2 V.cocovora.3 V.nigropicta.4 V.imuganensis." httpUri="https://zenodo.org/record/1241860/files/figure.png" pageId="2" pageNumber="137">Fig. 2</figureCitation>
),
<taxonomicName id="FDA34D79FFCDFFD1D334832CFB5DFBBB" authorityName="Soulier-Perkins &amp; Navasero &amp; Yap &amp; Wilson" authorityYear="2018" box="[1062,1230,1036,1059]" class="Insecta" family="Lophopidae" genus="Virgilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="imuganensis">
<emphasis id="08D7EAE8FFCDFFD1D334832CFBA0FBBB" box="[1062,1075,1036,1059]" italics="true" pageId="2" pageNumber="137">V</emphasis>
.
<emphasis id="08D7EAE8FFCDFFD1D350832CFB5DFBBB" box="[1090,1230,1036,1059]" italics="true" pageId="2" pageNumber="137">imuganensis</emphasis>
</taxonomicName>
colouration along the media and sub-lateral carinae (Fig. 4) and
<taxonomicName id="FDA34D79FFCDFFD1D5BA8310FCAAFBDF" authorityName="Stal" authorityYear="1870" box="[680,825,1072,1095]" class="Insecta" family="Lophopidae" genus="Virgilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="nigropicta">
<emphasis id="08D7EAE8FFCDFFD1D5BA8310FCAAFBDF" box="[680,825,1072,1095]" italics="true" pageId="2" pageNumber="137">V. nigropicta</emphasis>
</taxonomicName>
a single large patch, halfmoon shaped, touching the fronto-clypeal margin (
<figureCitation id="A2982A7FFFCDFFD1D6898373FE4CFBF4" box="[411,479,1107,1132]" captionStart="FIGURES 13" captionStartId="2.[151,264,584,607]" captionTargetBox="[151,1436,273,562]" captionTargetId="figure@2.[151,1436,273,570]" captionTargetPageId="2" captionText="FIGURES 13. Schematic patterns on Virgilia frons. 1 V. luzonensis. 2 V.cocovora.3 V.nigropicta.4 V.imuganensis." httpUri="https://zenodo.org/record/1241860/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72B96571FFCDFFD6D7D58357FC24FF4D" lastPageId="5" lastPageNumber="140" pageId="2" pageNumber="137" type="description">
<paragraph id="3A1C36FAFFCDFFD1D7D58357FDA4FB2B" blockId="2.[151,1436,783,2032]" pageId="2" pageNumber="137">
<emphasis id="08D7EAE8FFCDFFD1D7D58357FEC2FB08" bold="true" box="[199,337,1143,1168]" pageId="2" pageNumber="137">Description</emphasis>
. Total length of male
<typeStatus id="E5188858FFCDFFD1D54C8357FD56FB08" box="[606,709,1143,1168]" pageId="2" pageNumber="137" type="holotype">holotype</typeStatus>
:
<quantity id="FD5B9B1FFFCDFFD1D5C68358FCA1FB17" box="[724,818,1144,1168]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="137" unit="cm" value="1.17">1.17 cm</quantity>
(tegmina included), male
<typeStatus id="E5188858FFCDFFD1D3798358FB43FB08" box="[1131,1232,1144,1168]" pageId="2" pageNumber="137" type="paratype">paratype</typeStatus>
:
<quantity id="FD5B9B1FFFCDFFD1D3CD8358FAD3FB17" box="[1247,1344,1144,1168]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="137" unit="cm" value="1.17">1.17 cm</quantity>
, female
<typeStatus id="E5188858FFCDFFD1D78583BCFE91FB2C" box="[151,258,1180,1204]" pageId="2" pageNumber="137" type="paratype">paratypes</typeStatus>
between 1.31 and
<quantity id="FD5B9B1FFFCDFFD1D6C783BCFDA0FB2B" box="[469,563,1180,1204]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="2" pageNumber="137" unit="cm" value="1.49">1.49 cm</quantity>
.
</paragraph>
<paragraph id="3A1C36FAFFCDFFD1D7D5839FFE0EF910" blockId="2.[151,1436,783,2032]" pageId="2" pageNumber="137">Head. In dorsal view, head 1.78 times wider across eyes than long in midline. Vertex 1.74 longer than wide, posterior margin almost straight, anterior margin V shaped, lateral margins posterior two-third parallel and last anterior third diverging slightly. In frontal view, frons 1.04 wider across the widest part at gena level than long in midline, fronto-vertex and fronto-clypeal margins slightly rounded and respectively convex and concave, lateral margins regular and underlining the pear-shaped frons, the widest area is sharply curved, sub-lateral and median carinae present but not reaching the fronto-clypeal suture. In lateral view, ocellus present located below compound eye and anteriorly to antennae, ocellar carinae absent, genal carinae present. Prothorax larger than head and 4.15 times wider than long in midline, longest length being in midline. Mesothorax 1.45 times wider than long in midline. Tegmina 3.12 longer than wide at midline, ScP+R(+MA) forking first, then CuA and finally MP, all forking before the end of clavus and respectively around 1/5, 1/3 and 1/2 of the total tegminas length. Metatibiae bearing 3 lateral and 7 stout apical spines, first metatarsal segment bearing a series of apical spines organised in a triangular area, longer than cumulative length of second and third metatarsal segment, second segment reduced to a lobe without any spine.</paragraph>
<paragraph id="3A1C36FAFFCDFFD1D7D581B3FE58F868" blockId="2.[151,1436,783,2032]" pageId="2" pageNumber="137">Male terminalia. In lateral view, anal tube elongated posteriorly, 5.25 times longer than its biggest thickness measured just before the epiproct insertion, ventral margin developed just before the apex into an irregular tooth shaped extension pointing postero-ventrally. Pygofer with dorso-posterior margin largely and regularly rounded, posterior margin clearly inverted S shaped, posterior and ventral margin meeting in a sharp acute angle. Gonostylus with dorsal margin developing on its 2/3 posterior length a regular dorso-anterior oriented hook shaped extension, just posterior to the hook, the dorsal margin is curved ventrally, ventral margin roughly straight and posterior margin generally convex and irregularly curved. Periandrium with dorsal margin starting horizontally and bending up to its half at 90° and finishing in a rounded lobe, ventral margin curving as well and finishing dorsally in a hillock. Aedeagus bearing a ventral posterior process sickle-shaped and a short ventral anterior process strongly bent and pointing ventrally.</paragraph>
<caption id="6EDC6672FFCCFFD0D78580B4FE80F870" httpUri="https://zenodo.org/record/1241862/files/figure.png" pageId="3" pageNumber="138" startId="3.[151,264,1940,1963]" targetBox="[228,1355,203,1921]" targetPageId="3">
<paragraph id="3A1C36FAFFCCFFD0D78580B4FE80F870" blockId="3.[151,1436,1940,2024]" pageId="3" pageNumber="138">
<emphasis id="08D7EAE8FFCCFFD0D78580B4FEDAF832" bold="true" box="[151,329,1940,1963]" pageId="3" pageNumber="138">FIGURES 511.</emphasis>
<taxonomicName id="FDA34D79FFCCFFD0D64080B5FD95F833" authority="Soulier-Perkins &amp; Navasero &amp; Yap &amp; Wilson, 2018" authorityName="Soulier-Perkins &amp; Navasero &amp; Yap &amp; Wilson" authorityYear="2018" box="[338,518,1941,1963]" class="Insecta" family="Lophopidae" genus="Virgilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="cocovora" status="sp. nov.">
<emphasis id="08D7EAE8FFCCFFD0D64080B5FD95F833" box="[338,518,1941,1963]" italics="true" pageId="3" pageNumber="138">Virgilia cocovora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13E45793FFCCFFD0D50280B5FDDFF832" box="[528,588,1941,1962]" pageId="3" pageNumber="138" rank="species">
<emphasis id="08D7EAE8FFCCFFD0D50280B5FDDFF832" bold="true" box="[528,588,1941,1962]" pageId="3" pageNumber="138">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype.
<emphasis id="08D7EAE8FFCCFFD0D5D780B5FD41F832" bold="true" box="[709,722,1941,1962]" pageId="3" pageNumber="138">5</emphasis>
Habitus, dorsal view.
<emphasis id="08D7EAE8FFCCFFD0D4D080B5FC5CF832" bold="true" box="[962,975,1941,1962]" pageId="3" pageNumber="138">6</emphasis>
Habitus, lateral view.
<emphasis id="08D7EAE8FFCCFFD0D3D280B5FB5EF832" bold="true" box="[1216,1229,1941,1962]" pageId="3" pageNumber="138">7</emphasis>
Frons and clypeus, anterior view.
<emphasis id="08D7EAE8FFCCFFD0D6398094FEAAF851" bold="true" box="[299,313,1972,1993]" pageId="3" pageNumber="138">8</emphasis>
Labels.
<emphasis id="08D7EAE8FFCCFFD0D6838094FE0CF851" bold="true" box="[401,415,1972,1993]" pageId="3" pageNumber="138">9</emphasis>
Anal tube, pygopher and gonostylus, lateral view.
<emphasis id="08D7EAE8FFCCFFD0D4BE8094FC56F851" bold="true" box="[940,965,1972,1993]" pageId="3" pageNumber="138">10</emphasis>
Aedeagus s.l., lateral view.
<emphasis id="08D7EAE8FFCCFFD0D3F88094FA91F851" bold="true" box="[1258,1282,1972,1993]" pageId="3" pageNumber="138">11</emphasis>
Aedeagus s.s., lateral view.
</paragraph>
</caption>
<caption id="6EDC6672FFCBFFD7D78582C3FAD0F90B" httpUri="https://zenodo.org/record/1241864/files/figure.png" pageId="4" pageNumber="139" startId="4.[151,264,1507,1530]" targetBox="[314,1269,197,1483]" targetPageId="4">
<paragraph id="3A1C36FAFFCBFFD7D78582C3FAD0F90B" blockId="4.[151,1437,1507,1683]" pageId="4" pageNumber="139">
<emphasis id="08D7EAE8FFCBFFD7D78582C3FEC6FA61" bold="true" box="[151,341,1507,1530]" pageId="4" pageNumber="139">FIGURES 1234.</emphasis>
<taxonomicName id="FDA34D79FFCBFFD7D64E82C4FD9EFA62" authority="Soulier-Perkins &amp; Navasero &amp; Yap &amp; Wilson, 2018" authorityName="Soulier-Perkins &amp; Navasero &amp; Yap &amp; Wilson" authorityYear="2018" box="[348,525,1508,1530]" class="Insecta" family="Lophopidae" genus="Virgilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="cocovora" status="sp. nov.">
<emphasis id="08D7EAE8FFCBFFD7D64E82C4FD9EFA62" box="[348,525,1508,1530]" italics="true" pageId="4" pageNumber="139">Virgilia cocovora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13E45793FFCBFFD7D50782C4FDDCFA61" box="[533,591,1508,1529]" pageId="4" pageNumber="139" rank="species">
<emphasis id="08D7EAE8FFCBFFD7D50782C4FDDCFA61" bold="true" box="[533,591,1508,1529]" pageId="4" pageNumber="139">n. sp.</emphasis>
</taxonomicNameLabel>
, paratypes.
<emphasis id="08D7EAE8FFCBFFD7D5DE82C4FC98FA61" bold="true" box="[716,779,1508,1529]" pageId="4" pageNumber="139">1216</emphasis>
female MNHN(EH)23545
<emphasis id="08D7EAE8FFCBFFD7D33482C4FBD3FA61" bold="true" box="[1062,1088,1508,1529]" pageId="4" pageNumber="139">12</emphasis>
Habitus, dorsal view.
<emphasis id="08D7EAE8FFCBFFD7D23482C4FAACFA61" bold="true" box="[1318,1343,1508,1529]" pageId="4" pageNumber="139">13</emphasis>
Habitus, ventral view
<emphasis id="08D7EAE8FFCBFFD7D60F8123FEA4F980" bold="true" box="[285,311,1539,1560]" pageId="4" pageNumber="139">14</emphasis>
Habitus, lateral view
<emphasis id="08D7EAE8FFCBFFD7D5088123FDA7F980" bold="true" box="[538,564,1539,1560]" pageId="4" pageNumber="139">15</emphasis>
Frons and clypeus, anterior view
<emphasis id="08D7EAE8FFCBFFD7D4838123FC38F980" bold="true" box="[913,939,1539,1560]" pageId="4" pageNumber="139">16</emphasis>
Labels.
<emphasis id="08D7EAE8FFCBFFD7D3118123FBD1F980" bold="true" box="[1027,1090,1539,1560]" pageId="4" pageNumber="139">1721</emphasis>
male
<emphasis id="08D7EAE8FFCBFFD7D3918123FB0EF980" bold="true" box="[1155,1181,1539,1560]" pageId="4" pageNumber="139">17</emphasis>
Habitus, dorsal view.
<emphasis id="08D7EAE8FFCBFFD7D2918123FA0EF980" bold="true" box="[1411,1437,1539,1560]" pageId="4" pageNumber="139">18</emphasis>
Habitus, ventral view
<emphasis id="08D7EAE8FFCBFFD7D6658101FE02F9AE" bold="true" box="[375,401,1569,1590]" pageId="4" pageNumber="139">19</emphasis>
Habitus, lateral view
<emphasis id="08D7EAE8FFCBFFD7D57D8101FD1BF9AE" bold="true" box="[623,648,1569,1590]" pageId="4" pageNumber="139">20</emphasis>
Frons and clypeus, anterior view
<emphasis id="08D7EAE8FFCBFFD7D4F08101FC68F9AE" bold="true" box="[994,1019,1569,1590]" pageId="4" pageNumber="139">21</emphasis>
Labels.
<emphasis id="08D7EAE8FFCBFFD7D3408101FB02F9AE" bold="true" box="[1106,1169,1569,1590]" pageId="4" pageNumber="139">2225</emphasis>
female
<emphasis id="08D7EAE8FFCBFFD7D3F18101FB6EF9AE" bold="true" box="[1251,1277,1569,1590]" pageId="4" pageNumber="139">22</emphasis>
Habitus, dorsal view
<emphasis id="08D7EAE8FFCBFFD7D7C28160FF79F9CD" bold="true" box="[208,234,1600,1621]" pageId="4" pageNumber="139">23</emphasis>
Habitus, lateral view
<emphasis id="08D7EAE8FFCBFFD7D6DD8160FE7BF9CD" bold="true" box="[463,488,1600,1621]" pageId="4" pageNumber="139">24</emphasis>
Frons and clypeus, anterior view
<emphasis id="08D7EAE8FFCBFFD7D4588160FCF0F9CD" bold="true" box="[842,867,1600,1621]" pageId="4" pageNumber="139">25</emphasis>
Labels.
<emphasis id="08D7EAE8FFCBFFD7D4AF8160FC6EF9CD" bold="true" box="[957,1021,1600,1621]" pageId="4" pageNumber="139">2629</emphasis>
female MNHN(EH)235457
<emphasis id="08D7EAE8FFCBFFD7D2378160FAACF9CD" bold="true" box="[1317,1343,1600,1621]" pageId="4" pageNumber="139">26</emphasis>
Habitus, dorsal view
<emphasis id="08D7EAE8FFCBFFD7D608817FFEA0F9EC" bold="true" box="[282,307,1631,1652]" pageId="4" pageNumber="139">27</emphasis>
Habitus, lateral view
<emphasis id="08D7EAE8FFCBFFD7D530817FFDA8F9EC" bold="true" box="[546,571,1631,1652]" pageId="4" pageNumber="139">28</emphasis>
Frons and clypeus, anterior view
<emphasis id="08D7EAE8FFCBFFD7D4BB817FFC50F9EC" bold="true" box="[937,963,1631,1652]" pageId="4" pageNumber="139">29</emphasis>
Labels.
<emphasis id="08D7EAE8FFCBFFD7D333817FFBF3F9EC" bold="true" box="[1057,1120,1631,1652]" pageId="4" pageNumber="139">3034</emphasis>
female MNHN(EH)23548
<emphasis id="08D7EAE8FFCBFFD7D291817FFA0EF9EC" bold="true" box="[1411,1437,1631,1652]" pageId="4" pageNumber="139">30</emphasis>
Habitus, dorsal view.
<emphasis id="08D7EAE8FFCBFFD7D661815DFE1EF90A" bold="true" box="[371,397,1661,1682]" pageId="4" pageNumber="139">31</emphasis>
Habitus, ventral view
<emphasis id="08D7EAE8FFCBFFD7D560815DFD1FF90A" bold="true" box="[626,652,1661,1682]" pageId="4" pageNumber="139">32</emphasis>
Habitus, lateral view
<emphasis id="08D7EAE8FFCBFFD7D478815DFC10F90A" bold="true" box="[874,899,1661,1682]" pageId="4" pageNumber="139">33</emphasis>
Labels
<emphasis id="08D7EAE8FFCBFFD7D4C1815DFC7EF90A" bold="true" box="[979,1005,1661,1682]" pageId="4" pageNumber="139">34</emphasis>
Frons and clypeus, anterior view.
</paragraph>
</caption>
<paragraph id="3A1C36FAFFCBFFD6D7D5819AFC24FF4D" blockId="4.[151,1437,1722,2035]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="140" pageId="4" pageNumber="139">Colour. Generally brownish and cream with ochre powdery colour on tegmina that tend to disappear on specimens preserved in the collections. Head cream coloured with: on the vertex, a median discrete carina underlined in brown, two brown lines comma shaped close to the anterior margin and two yellowish-brown patches closes to the posterior margin; on the frons, 3 brown-black dots on each side along the sub-lateral carinae; on the clypeus that tend to get darker toward its apex, a brown patch in the middle, just below the fronto-clypeal suture; on each gena, a circular brown patch around the ocellus. Prothorax mixed with cream and brown with the cream more abundant along the anterior margin and the brown along the posterior margin, mesothorax brown and apex of metathorax cream. Tegmina dominantly dark brown with a series of 7-8 cream lunula shaped areas along the costal and radial margin that can be very small or larger and containing up to two dark stripes starting from the margin of the tegmina, one lunula on the apical margin in the medial area and one ultimate lunula on the margin on the anterior cubital area. Profemur with two dark brown transversal stripes.</paragraph>
</subSubSection>
<subSubSection id="72B96571FFCAFFD6D7D587FFFB38FE85" pageId="5" pageNumber="140" type="etymology">
<paragraph id="3A1C36FAFFCAFFD6D7D587FFFB38FE85" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="140">
<emphasis id="08D7EAE8FFCAFFD6D7D587FFFED5FF60" bold="true" box="[199,326,223,248]" pageId="5" pageNumber="140">Etymology</emphasis>
. The species name is built using the first four letters of coconut “
<emphasis id="08D7EAE8FFCAFFD6D37A87C1FB0EFF60" box="[1128,1181,225,248]" italics="true" pageId="5" pageNumber="140">coco</emphasis>
”, plant on which the specimens were collected and found feeding and the Latin word “
<emphasis id="08D7EAE8FFCAFFD6D4648625FC2CFE84" box="[886,959,261,284]" italics="true" pageId="5" pageNumber="140">vorare</emphasis>
” meaning to devour.
</paragraph>
</subSubSection>
</treatment>
</document>