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<document id="848886701E57BBEE0320A37E2D73ACD7" ID-DOI="10.3897/zookeys.1223.138220" ID-ZooBank="3557A991-63DF-42D8-B8B1-F310CD52FD00" ID-publisher-id="138220" URI-arpha="7352E8C9-65CA-546F-BE46-EF55E52C51A6" URI-zoobank="http://zoobank.org/3557A991-63DF-42D8-B8B1-F310CD52FD00" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1736187739607" checkinUser="pensoft" docAuthor="Hendrich, Lars &amp; Brancucci, Michel" docDate="2025" docId="B0D7A09D232B508A842FE1359BE49671" docLanguage="en" docName="ZooKeys 1223: 87-147" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/138220/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sandracottus maculatus Hendrich &amp; Brancucci" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="147" masterDocId="7352E8C965CA546FBE46EF55E52C51A6" masterDocTitle="Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)" masterLastPageNumber="147" masterPageNumber="87" pageNumber="87" updateTime="1736200074934" updateUser="admin">
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<mods:title id="077A901936DDCD952FA2A4A53EA0A154">Revision of the Oriental and Australasian diving beetle genus Sandracottus Sharp, 1882 (Coleoptera, Dytiscidae, Dytiscinae)</mods:title>
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<mods:namePart id="25AD2695560D5B542ED3746C5FD1B911">Hendrich, Lars</mods:namePart>
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<mods:affiliation id="647B48031EED1D7D97CF0BE70087AA27">SNSB - Zoologische Staatssammlung München, Münchhausenstraße 21, D 81247 München, Germany</mods:affiliation>
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<mods:namePart id="ABC02AFA474A4FB972CEEEA60BA6C4B8">Brancucci, Michel</mods:namePart>
<mods:affiliation id="080395367E1CE54B39E9B8967D290B73">Naturhistorisches Museum Basel, Basel, Switzerland</mods:affiliation>
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<mods:date id="4282FE3B5FC0B22A70634EEA954DAFE8">2025</mods:date>
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<mods:number id="08046C8BEA3A1277F80B0D8118AF59A2">2025-01-06</mods:number>
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<mods:identifier id="532EF3023229D7AB1F1CB3C92D21AA36" type="DOI">10.3897/zookeys.1223.138220</mods:identifier>
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<paragraph id="90111BEE4B87D2520BE91E710CAE9B19">
<taxonomicName id="DEA5827CB5FC848F88320FF80B51331E" ID-CoL="6XHNK" authority="(Wehncke, 1876)" authorityName="Hendrich &amp; Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="8AFF7D8328E6B49AE2FE8079B6F1D88B" italics="true">Sandracottus maculatus</emphasis>
(
<bibRefCitation id="07FE553040F8F6DB6AD5D43003570273" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="B10D6966B6BF5C9E8126F380D10BF43C" type="description">
<paragraph id="219E461EB0C3B4068160C153485624AD">
<figureCitation id="761CFB9C9CA36DF54AFDD62BE54BC55B" captionStart="Figures 811" captionStartId="F3" captionText="Figures 811. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">Figs 10</figureCitation>
,
<figureCitation id="BA8DDFF464C37DDE3A6820B1B5E25C39" captionStart="Figures 2022" captionStartId="F6" captionText="Figures 2022. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 20 Sandracottus jaechi 21 S. maculatus and 22 S. rotundus." figureDoi="10.3897/zookeys.1223.138220.figures20-22" httpUri="https://binary.pensoft.net/fig/1214413">21</figureCitation>
,
<figureCitation id="ADAF5710EDCCB22060AA54DED56885DE" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Distribution of Sandracottus bizonatus [red squares] and S. maculatus [black dots] in Southeast Asia." figureDoi="10.3897/zookeys.1223.138220.figure24" httpUri="https://binary.pensoft.net/fig/1214415">24</figureCitation>
,
<figureCitation id="A200BC58BC4801569CA24A6BCC1CAE96" captionStart="Figures 3742" captionStartId="F16" captionText="Figures 3742. Habitats of Sandracottus 37, 38 Laos, Attapeu prov., Annam Highlands Mts., Dong Amphan NBCA, Nong Fa [puddles near lake shore] and lake, habitat of S. hunteri 39 Laos, Katamtok, rest pools along a rocky river bed, habitat of S. maculatus (photos: J. Hájek) 40 Thailand, near Salawin NP, open and muddy puddle made by elephants (photo: J. Šťastný), and 41 Sakaerat Biosphere Reserve, rocky pool in an otherwise dry stream bed, habitats of S. hunteri and S. maculatus (photo: W. Atthakor) 42 Philippines, Luzon, Pampanga Prov., Bano San Juan, Mt. Arayat NP, spring creek, habitat of S. insignis (photo: H. Freitag)." figureDoi="10.3897/zookeys.1223.138220.figures37-42" httpUri="https://binary.pensoft.net/fig/1214423">39</figureCitation>
,
<figureCitation id="00A87F5185DA64AD9FD5A37A34A62710" captionStart="Figures 3742" captionStartId="F16" captionText="Figures 3742. Habitats of Sandracottus 37, 38 Laos, Attapeu prov., Annam Highlands Mts., Dong Amphan NBCA, Nong Fa [puddles near lake shore] and lake, habitat of S. hunteri 39 Laos, Katamtok, rest pools along a rocky river bed, habitat of S. maculatus (photos: J. Hájek) 40 Thailand, near Salawin NP, open and muddy puddle made by elephants (photo: J. Šťastný), and 41 Sakaerat Biosphere Reserve, rocky pool in an otherwise dry stream bed, habitats of S. hunteri and S. maculatus (photo: W. Atthakor) 42 Philippines, Luzon, Pampanga Prov., Bano San Juan, Mt. Arayat NP, spring creek, habitat of S. insignis (photo: H. Freitag)." figureDoi="10.3897/zookeys.1223.138220.figures37-42" httpUri="https://binary.pensoft.net/fig/1214423">41</figureCitation>
,
<figureCitation id="F7255ED58FA3644F09A713575DF18C4E" captionStart="Figures 4345" captionStartId="F17" captionText="Figures 4345. Habitats of Sandracottus 43, 44 Malaysia, Sabah, Kinabatangan riv. 8. - 15. VI. 2003, Uncle Tan´s camp, habitat of S. maculatus (photos: J. Šťastný) 45 Indonesia, Sulawesi, Togian Islands, Kaldidiri Island near Paradise Island Resort, Babirusa puddles (on the left), habitat of S. rotundus (photo: J. Haft)." figureDoi="10.3897/zookeys.1223.138220.figures43-45" httpUri="https://binary.pensoft.net/fig/1214424">43</figureCitation>
,
<figureCitation id="29857C896255B555031AFA19C570B703" captionStart="Figures 4345" captionStartId="F17" captionText="Figures 4345. Habitats of Sandracottus 43, 44 Malaysia, Sabah, Kinabatangan riv. 8. - 15. VI. 2003, Uncle Tan´s camp, habitat of S. maculatus (photos: J. Šťastný) 45 Indonesia, Sulawesi, Togian Islands, Kaldidiri Island near Paradise Island Resort, Babirusa puddles (on the left), habitat of S. rotundus (photo: J. Haft)." figureDoi="10.3897/zookeys.1223.138220.figures43-45" httpUri="https://binary.pensoft.net/fig/1214424">44</figureCitation>
,
<figureCitation id="9A59515428DF8389DD033FFC619CF33B" captionStart="Figures 5457" captionStartId="F20" captionText="Figures 5457. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">56</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="FFA31F45AD88CC42D7FB1066C0A08B17">
<treatmentCitationGroup id="17B9EDC0C5E8834EAB9BA262C3C5A8B4">
<treatmentCitation id="CEB27FC81F8B0668F534678900AB8B03" author="Wehncke E" page="196" year="1876">
<taxonomicName id="3613F480B60AC4DA51A8703E72D31E80" authority="Wehncke, 1876: 196" authorityName="Wehncke" authorityPageNumber="196" authorityYear="1876" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="C5171EDDD0AACBB8FB1A222C3A8DD960" italics="true">Hydaticus maculatus</emphasis>
<comment id="BE6859B7FF6AFE98AE9D6BAF1B8B84AA">
<bibRefCitation id="57A37B1E91AC0BD1E8C7E3EDF4CF1668" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876: 196</bibRefCitation>
(type locality “
<collectingCountry id="C6FAF1F07B90527D33B950F090A0C208" name="Thailand">Siam</collectingCountry>
” [
<collectingCountry id="C9ABABD1993E30E4493A3564A6DD3443" name="Thailand">Thailand</collectingCountry>
]).
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="45159866EDC9CF5B6B8439CD13BD050C">
<treatmentCitationGroup id="1E1D93982A37D63CCE4438DA55C6AC24">
<treatmentCitation id="854798F503754353AFA35BA5CE083CF8">
<taxonomicName id="623A52D1691B9FB06FE595DEEC021B1B" authority="J. Balfour-Browne, 1944: 355" authorityName="J. Balfour-Browne" authorityPageNumber="355" authorityYear="1944" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="wehnckei">
<emphasis id="098A10486960E161C47B5B322989A8F1" italics="true">Sandracottus wehnckei</emphasis>
<comment id="85666E0A02C606C9ED5EE891AA0553D9">
J. Balfour-Browne, 1944: 355, replacement name for
<taxonomicName id="72D2C34BDB99CA9E6324627ED36E632F" authority="Wehncke, 1876" authorityName="Wehncke" authorityYear="1876" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="22969248A38E8268D9F2BC7A50FAAB1F" italics="true">Hydaticus maculatus</emphasis>
<bibRefCitation id="D0B5ED2D60CC2B4C0AC013B77B3B5427" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
</taxonomicName>
(objective synonym of
<taxonomicName id="8E7C78095F18D6E00A0DDB7DEDB4FEAD" authorityName="Wehncke" authorityYear="1876" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="50FFDD00BFF4BB23CA6FB322B52EF021" italics="true">Hydaticus maculatus</emphasis>
</taxonomicName>
);
<treatmentCitation id="976F9E5FA8A79F9696DE0EB069E29EC0" author="Vazirani TG" page="277" year="1969">
<bibRefCitation id="BFC822CD2D1D9F0440F37ADEE5AEC8D7" DOI="10.1080/00305316.1968.10433885" author="Vazirani TG" firstAuthor="Vazirani" issue="1968" journalOrPublisher="Oriental Insects" page="277" pagination="221-341" refId="B52" refString="Vazirani TG (1969) A review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India. Oriental Insects 2 (1968): 221341. https://doi.org/10.1080/00305316.1968.10433885" title="A review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India." volume="2" year="1969">Vazirani 1969: 277</bibRefCitation>
</treatmentCitation>
(descr., syst.); 1977: 86 (cat.); Nilsson 2001: 84 (cat.).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2C35314A0F65E8F5E3417361987D1A80">
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<treatmentCitation id="796F33D1BFFB829BC715AA22FD6FD6C5">
<taxonomicName id="68B414397A3D8F540992CEB70D825CCB" ID-CoL="6XHNK" authority="(Wehncke, 1876)" authorityName="Hendrich &amp; Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="3AED368B019D8B48C90FB85C5E3AD1B2" italics="true">Sandracottus maculatus</emphasis>
<comment id="93233A0B3332385DB57BC4C47700F2A4">
(
<bibRefCitation id="BA7C09EC8D248FC5BE86EDCAD702F5F1" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
):
<treatmentCitation id="E8066B51E3F312118E2C838588419578" author="Sharp D" page="686" year="1882">
<bibRefCitation id="F764A451638C4F470FE4ED4705B430C8" DOI="10.5962/bhl.title.9530" author="Sharp D" firstAuthor="Sharp" page="686" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 1791003. [+ pls. 718] https://doi.org/10.5962/bhl.title.9530" year="1882">Sharp 1882: 686</bibRefCitation>
</treatmentCitation>
(comb. nov.);
<treatmentCitation id="E4487360790DC40AE8D5318646C7C6F8" author="Régimbart M" page="338" year="1899">
<bibRefCitation id="F34A50D65F8AF393D1C1CC08EEC4DA62" DOI="10.1080/21686351.1899.12279737" author="Régimbart M" firstAuthor="Regimbart" journalOrPublisher="Annales de la Société Entomologique de France" page="338" pagination="186-367" refId="B42" refString="Régimbart M (1899) Révision des Dytiscidae de la région Indo-Sino-Malaise. Annales de la Société Entomologique de France 68: 186367. https://doi.org/10.1080/21686351.1899.12279737" title="Révision des Dytiscidae de la région Indo-Sino-Malaise." volume="68" year="1899">Régimbart 1899: 338</bibRefCitation>
</treatmentCitation>
(descr.);
<treatmentCitation id="9482EC74BF8247FD4D9CCF68E7908D63" author="Zimmermann A" page="235" year="1920">
<bibRefCitation id="6C198B5926964AD06483BF749E27AA9F" author="Zimmermann A" firstAuthor="Zimmermann" journalOrPublisher="In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71" page="235" refId="B60" refString="Zimmermann A (1920) Dytiscidae. In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71, 326 pp." title="Dytiscidae." year="1920">Zimmermann 1920: 235</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="DCFCA140441C0C57944FEE27FA4EE44F" author="Ghosh SK &amp; Nilsson AN" page="18" year="2012">
<bibRefCitation id="5B261180059A42CA2CA054CA943E43B0" author="Ghosh SK &amp; Nilsson AN" firstAuthor="Ghosh" journalOrPublisher="Skörvnöpparn, Umea, Supplement" page="18" pagination="1-77" refId="B18" refString="Ghosh SK, Nilsson AN (2012) Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae). Skörvnöpparn, Umea, Supplement 3: 177." title="Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae)." volume="3" year="2012">Ghosh and Nilsson 2012: 18</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="D25B7BC9DDA75B39B14824636A63783C" author="Balke M &amp; Bergsten J &amp; Wang L-J &amp; Hendrich L" page="146" year="2017">
<bibRefCitation id="A8198887E9C45DB8DEC7565196DA2F40" DOI="10.3897/zookeys.647.11231" author="Balke M &amp; Bergsten J &amp; Wang L-J &amp; Hendrich L" etAl="et al." firstAuthor="Balke" journalOrPublisher="ZooKeys" page="146" pagination="137-151" refId="B6" refString="Balke M, Bergsten J, Wang L-J, Hendrich L (2017) A new genus and two new species of Southeast Asian Bidessini as well as new synonyms for Oceanian species (Coleoptera, Dytiscidae). ZooKeys 647: 137151. https://doi.org/10.3897/zookeys.647.11231" title="A new genus and two new species of Southeast Asian Bidessini as well as new synonyms for Oceanian species (Coleoptera, Dytiscidae)." volume="647" year="2017">Balke et al. 2017: 146</bibRefCitation>
</treatmentCitation>
(faun., habitat);
<treatmentCitation id="11DD441ADAC3F8268B9D84AFCD98071F" author="Atthakor W &amp; Hendrich L &amp; Sangpradub N &amp; Balke M" page="93" year="2018">
<bibRefCitation id="7DC250E0B12E8E7C06DAC0B6B55C5F7A" author="Atthakor W &amp; Hendrich L &amp; Sangpradub N &amp; Balke M" etAl="et al." firstAuthor="Atthakor" issue="1" journalOrPublisher="Spixiana" page="93" pagination="91-98" refId="B3" refString="Atthakor W, Hendrich L, Sangpradub N, Balke M (2018) Diving beetles of the Sakaerat Biosphere Reserve, Nakhon Ratchasima Province, with four new records for Thailand (Coleoptera, Dytiscidae). Spixiana 41 (1): 9198." title="Diving beetles of the Sakaerat Biosphere Reserve, Nakhon Ratchasima Province, with four new records for Thailand (Coleoptera, Dytiscidae)." volume="41" year="2018">Atthakor et al. 2018: 93</bibRefCitation>
</treatmentCitation>
(faunistics, habitat); Hájek and
<treatmentCitation id="2073040BC91C752C41D57842E2E5CB19" author="Nilsson A &amp; Hájek J" page="91" year="2024">
<bibRefCitation id="40FD06D92E4E5A7614863DEF37AA001F" author="Nilsson A &amp; Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
</treatmentCitation>
(cat.).
</comment>
</taxonomicName>
</treatmentCitation>
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</paragraph>
<paragraph id="9AF762DACC1D83994F75AEEE4845D87F">
<treatmentCitationGroup id="9585691F18E0E75648CF4F87B38C4215">
<treatmentCitation id="3AD16023F11AAFB4A2082752C72753B0">
<taxonomicName id="BA078332DC1FE4B20DBF92D3928741F3" ID-CoL="6XHKZ" authority="Heller, 1934 a: 280" authorityName="Heller" authorityPageNumber="280" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="angulifer">
<emphasis id="3E0CE0494BEA1D28C8D968E3393E890D" italics="true">Sandracottus angulifer</emphasis>
<comment id="5E30BF8F465D6603DD0D328226A84421">
Heller, 1934 a: 280 (type locality
<collectingRegion id="4890B15DD655AA4B1A873AC9657C3865" country="Philippines" name="Davao">Davao</collectingRegion>
, Mindanao,
<collectingCountry id="EA802A874FD341CE6AC7A8F0A088114A" name="Philippines">Philippines</collectingCountry>
); Hájek and
<treatmentCitation id="236AE4BF7253E076A7784C187F15289D" author="Nilsson A &amp; Hájek J" page="91" year="2024">
<bibRefCitation id="52ACB5341011863430F78BABC585C700" author="Nilsson A &amp; Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
</treatmentCitation>
(cat.) (syn. nov.).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="722D9028F49856B26DA92EF8A0AD7A70">
<treatmentCitationGroup id="068EB3B2E4A3C8A83790B1EA95CB6225">
<treatmentCitation id="B99568CD0302601750123E9E4934B439" author="Satô M" page="41" year="1978">
<taxonomicName id="10C539FD11C3B7E3DBFAA9335DF6EA04" ID-CoL="6XHNN" authority="Sato, 1978: 41" authorityName="M. Sato DET. M. SATO" authorityPageNumber="41" authorityYear="1978" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="0EE7FDBC32943FDBE36C8AA22A2FA07E" italics="true">Sandracottus palawanensis</emphasis>
<comment id="DDC8504ED33DBBD2DE62CF9FDAE5D4B9">
<bibRefCitation id="43993F0C5985709720997F69F893C270" author="Satô M" journalOrPublisher="Bulletin of the National Science Museum (A, Zoology)" pagination="39-42" refId="B43" refString="Sato M (1978) Two new dytiscid beetles from Palawan, the Philippines. Bulletin of the National Science Museum (A, Zoology) 5: 39 - 42." title="Two new dytiscid beetles from Palawan, the Philippines." volume="5" year="1978">Satô, 1978: 41</bibRefCitation>
(type locality Sabang,
<collectingRegion id="790969F3DC47CEB81448D33F7182B00C" country="Philippines" name="Palawan">Palawan</collectingRegion>
,
<collectingCountry id="CC7B93EB706F43FBD83249F5355CA2BA" name="Philippines">Philippines</collectingCountry>
); Hájek and
<treatmentCitation id="BA080A5140C486008DCE2BC5A318EB8E" author="Nilsson A &amp; Hájek J" page="91" year="2024">
<bibRefCitation id="E618089B0F4E8D477743533B88CF6836" author="Nilsson A &amp; Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
</treatmentCitation>
(cat.) (syn. nov.).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="807257887963A6752A0D23F7D78433DE">
<treatmentCitationGroup id="6AC158F6B4136CA93BAB4C9AFD105CBF">
<treatmentCitation id="7E7D7FCAB8FB454839AECB8BF96C0064" author="Sharp D" page="690" year="1882">
<taxonomicName id="01DE5BC76BC55FE56048D745AA2B36AB" ID-CoL="79HNH" authority="Sharp, 1882: 690" authorityName="Sharp. In" authorityPageNumber="690" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nauticus">
<emphasis id="3F9B45446F13E9F58B01F2363A86FBD6" italics="true">Sandracottus nauticus</emphasis>
<comment id="EF63F55E09FEF3DB1F13B0F94ADFEE3C">
<bibRefCitation id="52A4D46C9BA0426AF446D810FE31A172" DOI="10.5962/bhl.title.9530" author="Sharp D" refId="B45" refString="Sharp D (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2: 179 - 1003. [+ pls. 7 - 18] https: // doi. org / 10.5962 / bhl. title. 9530" year="1882">Sharp, 1882: 690</bibRefCitation>
(type locality Borneo);
<treatmentCitation id="81D0D0020DE65BD5A3609A0343EE66C6" author="Régimbart M" page="339" year="1899">
<bibRefCitation id="1014C5747999BB6D1138DA830C81A752" DOI="10.1080/21686351.1899.12279737" author="Régimbart M" firstAuthor="Regimbart" journalOrPublisher="Annales de la Société Entomologique de France" page="339" pagination="186-367" refId="B42" refString="Régimbart M (1899) Révision des Dytiscidae de la région Indo-Sino-Malaise. Annales de la Société Entomologique de France 68: 186367. https://doi.org/10.1080/21686351.1899.12279737" title="Révision des Dytiscidae de la région Indo-Sino-Malaise." volume="68" year="1899">Régimbart 1899: 339</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="C045BA1EB26F45557AB71C35B9338AB9" author="Zimmermann A" page="235" year="1920">
<bibRefCitation id="4FD7508C179054BDA8A7F252FAB89BB7" author="Zimmermann A" firstAuthor="Zimmermann" journalOrPublisher="In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71" page="235" refId="B60" refString="Zimmermann A (1920) Dytiscidae. In: Junk W, Schenkling S (Eds) Berlin, Coleopterorum Catalogus 4, pars 71, 326 pp." title="Dytiscidae." year="1920">Zimmermann 1920: 235</bibRefCitation>
</treatmentCitation>
(cat.); Hájek and
<treatmentCitation id="CC810FEB03177DA8F83E0A30959A53B4" author="Nilsson A &amp; Hájek J" page="91" year="2024">
<bibRefCitation id="391A5DB172CE7D5166757C15F9A07E6A" author="Nilsson A &amp; Hájek J" refId="B38" refString="Nilsson A, Hajek J (2024) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2024. Distributed as a PDF file via Internet, 323 pp. http: // www. waterbeetles. eu / documents / W _ CAT _ Dytiscidae _ 2024. pdf" url="http://www.waterbeetles.eu/documents/W_CAT_Dytiscidae_2024.pdf" year="2024">Nilsson 2024: 91</bibRefCitation>
</treatmentCitation>
(cat.) (syn. nov.).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWKCK" type="type material">
<paragraph id="6F806E062810928A97AD673513F882B1">
<heading id="D2B7E29E4F02BBDA1CC5C233EAB03CDB" reason="title">
Type material of
<taxonomicName id="7BA15B0B2379A287DCAA60E5377F2813" authorityName="Hendrich &amp; Brancucci" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="DDD6539E73EA9859C936F7FF1823D47B" italics="true">Sandracottus maculatus</emphasis>
</taxonomicName>
.
</heading>
</paragraph>
<paragraph id="2E34ECE62718E3CA6BDA3C781BD66CDD">
<materialsCitation id="43F81273293940400ADBFE4058F2C921" collectionCode="MNHN" collectorName="Schoenich" country="Cambodia" location="Cambodscha" specimenCount="1" specimenCount-male="1" typeStatus="neotype">
<emphasis id="D0CD8262DE38C8166A9ADB60AA45C49B" bold="true" italics="true">
<typeStatus id="DC317C49A452B76E953AEEC71BB8C8C7" type="neotype">Neotype</typeStatus>
</emphasis>
(herein designated):
<specimenCount id="0F61054AD8621337004B477F0E5D30A1" type="male">Male</specimenCount>
, “
<collectorName id="EAE2F3203E7C77B7E3ED7369D6EE8242">Schoenich</collectorName>
” [?], “
<collectingCountry id="FC48BCE2EE17E7CF5C6BA09BA405D203">Cambodscha</collectingCountry>
” [yellowish handwritten label], “ Ex. Coll. E. WEHNCKE Acq 1884 ” [white printed label], “ NEOTYPUS
<taxonomicName id="B15D121A40A2D27DA69ABEB4FA101B36" authority="L. Hendrich &amp; M. Brancucci" authorityName="L. Hendrich &amp; M. Brancucci" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="6561886B934AB56E4097CF8B27D9F57D" italics="true">
<originalDetermination id="79C0FB1CC1AE2E2AC8A3F03E2B9263FE">Hydaticus maculatus</originalDetermination>
</emphasis>
L. Hendrich &amp; M. Brancucci
</taxonomicName>
des. 2010 ” [red printed label] (
<abbrev id="ABBRID0EXLCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="41DD713FA815757594028CFDE47E1640" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
)
</materialsCitation>
. Examined.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2LCK" type="comments">
<paragraph id="1005BC5F9221374D9724B75F9F3F637F">
<heading id="8DF8B4CF8033D585329DE96CC2283575" reason="title">Comments on classification.</heading>
</paragraph>
<paragraph id="9D97F0A820CF964CBF91784637178FDE">
It was impossible to find any type material labelled “
<collectingCountry id="B72E2B8361007E64F618538B00477069" name="Thailand">Siam</collectingCountry>
” among the historical specimens in the
<abbrev id="ABBRID0EBMCK" xlink_title="Natural History Museum, London">NHMUK</abbrev>
,
<abbrev id="ABBRID0EFMCK" xlink_title="Muséum national d´Histoire naturelle, Paris">MNHN</abbrev>
, and
<abbrev id="ABBRID0EJMCK" xlink_title="Museum für Naturkunde, Berlin">ZHMB</abbrev>
, so we decided to designate a
<typeStatus id="E8A680DA1FF5CB5F7DFD7246D993A82A" type="neotype">neotype</typeStatus>
from
<collectingCountry id="23149190254AE78F94F9D021E7F38BFF" name="Cambodia">Cambodia</collectingCountry>
found in Wehncke´s collection and deposited in
<abbrev id="ABBRID0ENMCK" xlink_title="Muséum national d´Histoire naturelle, Paris">MNHN</abbrev>
. The
<typeStatus id="7B102E6BF061EA54E2591C8FEB5C999D" type="neotype">neotype</typeStatus>
is designated to support taxonomic stability, as there are several morphological similar
<taxonomicName id="6EB7C8C64CF0E7476E9326CCA8596540">
<emphasis id="692B163A22290A4FD4A721EBC2B9DD2E" italics="true">Sandracottus</emphasis>
species
</taxonomicName>
described from the Indomalayan region, and it is paramount for present and future investigations to possess unambiguously characterised name-bearing specimens for all Indomalayan species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYMCK" type="type material">
<paragraph id="8038C308C5B4DFA17CB63B57B278C248">
<heading id="D217877FACF44E116981FCC0FBD98AD1" reason="title">
Type material of
<taxonomicName id="6C2BB7D9FBC3082FC0E2FBBF5891EA09" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="angulifer">
<emphasis id="EB866187821A5CA934B76B789AA64A9F" italics="true">Sandracottus angulifer</emphasis>
</taxonomicName>
.
</heading>
</paragraph>
<paragraph id="48E12589C18EAA32BA986084CD4FFAE0">
<materialsCitation id="4AECA811BF37C518827ADF16E6A17FA4" collectionCode="SMTD" collectorName="Baker" determinerName="Hendrich &amp; Brancucci" location="Mindanao" specimenCount="1" stateProvince="Davao" typeStatus="holotype">
<emphasis id="698F5A654927EB1D4461BB6C1E8A5943" bold="true" italics="true">
<typeStatus id="C81150F7B4CB3A9500BA29F3CE5AACF9" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="D5DA60BC348A6AFAD2DAAC0ED6720B05" type="male">Male</specimenCount>
, “
<collectingRegion id="4CD4304EC1A708C6606F46C2C16A8213" country="Philippines" name="Davao">Davao</collectingRegion>
<location id="F50F2C35A503241965236F1EAF67B272" LSID="urn:lsid:plazi:treatment:B0D7A09D232B508A842FE1359BE49671:F50F2C35A503241965236F1EAF67B272" name="Mindanao" stateProvince="Davao">Mindanao</location>
<collectorName id="E9D4D008C2169A648EF1830C4E2FDFFF">Baker</collectorName>
”, “ 7251 ”, “
<taxonomicName id="59AE941E6FADC01FED65CFF72CF5232B" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="angulifer">
<emphasis id="2BD0DD2DEDFAD9CF6B9DBA952A1E9713" italics="true">
<originalDetermination id="7068803C6B33A7805EA67C9B65B9E285">S. angulifer</originalDetermination>
</emphasis>
</taxonomicName>
Typus ” [red label], “ 1932 12 ”, “ Staatl. Museum für Tierkunde, Dresden ”, “
<taxonomicName id="ED3880298B44C7DD8B7AAD4989694C2F" authority="(Wehncke, 1876)" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<originalDetermination id="A067C6C397DFA9415CAE49A4DCFEAFF0">
<emphasis id="89ED145F8F6C4B888B8EE3025B121845" italics="true">Sandracottus maculatus</emphasis>
(
<bibRefCitation id="DA7BF2FD09B7EA1176D7096926E6559E" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
)
</originalDetermination>
</taxonomicName>
<determinerName id="526D55CD6B3B2F71E43210110A589941">Hendrich &amp; Brancucci</determinerName>
det. 2006 ” (
<abbrev id="ABBRID0EEOCK" xlink_title="Staatliches Museum für Tierkunde">
<collectionCode id="C09D98D705FDA336ED766A59746A1B7E">SMTD</collectionCode>
</abbrev>
)
</materialsCitation>
. Examined.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIOCK" type="type material">
<paragraph id="59FC93A73FC8EA967618256FF53C3BC2">
<heading id="B5C89305BABD9E443CCEFA4F54EB4A58" reason="title">
Type material of
<taxonomicName id="BEDF6190BBA7DF9C4AE8204657957ABE" authorityName="M. Sato DET. M. SATO" authorityYear="1978" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="448A90025EEABFFA1A068F81F0C810A7" italics="true">Sandracottus palawanensis</emphasis>
</taxonomicName>
.
</heading>
</paragraph>
<paragraph id="F414C9C34BA44AE04369044D92AB1CB9">
<materialsCitation id="570F1A395DA3251BA5B98BDD1E942FCB" collectingDate="1977-07-13" collectionCode="NMST" collectorName="M. Sato" country="Philippines" determinerName="Hendrich &amp; Brancucci" location="Sabang" specimenCount="1" specimenCount-male="1" stateProvince="Palawan" typeStatus="holotype">
<emphasis id="A5E2B55393EC3A32F3914E56C4A4E100" bold="true" italics="true">
<typeStatus id="099AA4FD81871BD2DE02759950E36D4A" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="D7C3AABCF352A890368203912B7093E0" type="male">Male</specimenCount>
, “
<location id="E9FEAF98C5044FA5C27BA9095F65F2B4" LSID="urn:lsid:plazi:treatment:B0D7A09D232B508A842FE1359BE49671:E9FEAF98C5044FA5C27BA9095F65F2B4" country="Philippines" name="Sabang" stateProvince="Palawan">Sabang</location>
,
<collectingRegion id="F9B89AF0F172774FCC7D81495E2BFA9F" country="Philippines" name="Palawan">Palawan</collectingRegion>
<collectingCountry id="E68C8A3BC609B658553CDE4740AD39CC" name="Philippines">Philippines</collectingCountry>
<date id="F23A5316B56C816ACB871C8E88CDEF79" value="1977-07-13">
<collectingDate id="4C5698333F354B8CE125D41EA16A2AE2" value="1977-07-13">July 13, 1977</collectingDate>
</date>
<collectorName id="BCABF7667280642D5BC5D7BCF19E1391">M. Sâto</collectorName>
leg. ” [handwritten], “ HOLOTYPE
<taxonomicName id="E904D18D327241541CCF0B6C77E380D9" authority="M. Sato" authorityName="M. Sato" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="palawanensis">
<originalDetermination id="871A6102AA406659914BD52FDEC714CC">
<emphasis id="812A9AA600365BCB4AD5AA984CF34739" italics="true">Sandracottus palawanensis</emphasis>
M. Satô
</originalDetermination>
</taxonomicName>
DET. M.
<bibRefCitation id="2DDB426BEAD8C321021EC61D6ECEA566" author="SATO" firstAuthor="SATO" journalOrPublisher="Bulletin of the National Science Museum (A, Zoology)" pagination="39-42" refId="B43" refString="Satô M (1978) Two new dytiscid beetles from Palawan, the Philippines. Bulletin of the National Science Museum (A, Zoology) 5: 3942." title="Two new dytiscid beetles from Palawan, the Philippines." volume="5" year="1978">SATÔ 1978</bibRefCitation>
” [red label], “
<taxonomicName id="D5CCFEC047BE205C675A380426119BB1" authority="(Wehncke, 1876)" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<originalDetermination id="F4DAABE202AFE07E55C7F4B0ED04D877">
<emphasis id="D6C5A63E08B04FB3BC665F4D0C86CD9B" italics="true">Sandracottus maculatus</emphasis>
(
<bibRefCitation id="B16B703B12F38CA1E062031E7F256A54" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
)
</originalDetermination>
</taxonomicName>
<determinerName id="41859F311D46C441BAD2B646067E4CB5">Hendrich &amp; Brancucci</determinerName>
det. 2010 ” (
<abbrev id="ABBRID0EYPCK" xlink_title="National Museum of Nature and Science">
<collectionCode id="DFB7C20A9B15885BCE0646D9B746E61D">NMST</collectionCode>
</abbrev>
)
</materialsCitation>
. Examined.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3PCK" type="type material">
<paragraph id="1271E4D8FF2A686E7E0C69BEA022BA26">
<heading id="496717BEA41C26D311C275E40BCF1879" reason="title">
Type material of
<taxonomicName id="CE9B97B65359754E74B086F50CBF7347" authorityName="Sharp. In" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nauticus">
<emphasis id="AF1B12CC47F0E133B609F87674FAAEAC" italics="true">Sandracottus nauticus</emphasis>
</taxonomicName>
.
</heading>
</paragraph>
<paragraph id="3D782FA003B54B14B0B1BBE70944FF27">
<materialsCitation id="688270A6B4E7D06B667F33860869260D" collectionCode="NHMUK" collectorName="Male" specimenCount="1" typeStatus="holotype">
<emphasis id="B78E08BEDDC602393EA94A78ABE78702" bold="true" italics="true">
<typeStatus id="59DF2F80E28E4CA4DEF382C32EEEB1FD" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectorName id="09FF4C06D8B5A92F5E893BF41E60E0B1">Male</collectorName>
, “ Borneo 987
<taxonomicName id="5BB618DBBC17AF401B25C4FF0D52325F" authorityName="Sharp. In" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nauticus">
<emphasis id="F8BEE85F5F880BF4711ED53505C8A45C" italics="true">nauticus</emphasis>
</taxonomicName>
”, “ Type H. T. ”, “ Sharp Coll. 1905-313 ” (
<abbrev id="ABBRID0E3QCK" xlink_title="Natural History Museum, London">
<collectionCode id="34720EAA3AA1758696F81D3830F232F2" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
). “
<taxonomicName id="617339D3E049F5ED1CF966FC9EC2DFB7" authority="(Wehncke, 1876)" baseAuthorityName="Wehncke" baseAuthorityYear="1876" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="013495583847AF4C8FF94D56EF9E9727" italics="true">Sandracottus maculatus</emphasis>
(
<bibRefCitation id="C42B4CF36B9A5C41903709DD47991AD0" author="Wehncke E" refId="B57" refString="Wehncke E (1876) Neue Dytisciden. Stettiner Entomologische Zeitung 37: 194 - 197, 356 - 360." year="1876">Wehncke, 1876</bibRefCitation>
)
</taxonomicName>
Hendrich &amp; Brancucci det. 2006 ”. Examined.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELRCK" type="material">
<paragraph id="30806A0A82402D1C1DCACAD0E65A19AF">
<heading id="3CF6D452F6B9F9A0D201A9965BFA5D56" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="76F2378A702B6148476137A2F19503B6">
<emphasis id="8D468964224BAC44BA0E7FBD9EFE84BE" bold="true">
(
<specimenCount id="C4AC97D51479745C722FC00AB1B5953D" count="150" type="generic">150 specimens</specimenCount>
)
</emphasis>
: •
<emphasis id="BF58FCD64F9022C57AD3657B4915A0A3" bold="true">
<collectingCountry id="73B059D03E1ED13BC5C1BC07F4B8D8AF" name="Cambodia">Cambodia</collectingCountry>
</emphasis>
:
<specimenCount id="0928198D0B8DD5307B68FB5AD4774F30" count="1" type="generic">1 ex.</specimenCount>
, “ Cambodia
<collectingRegion id="CD925794E236DD9A93C9A118B5115734" country="Cambodia" name="Pailin">Pailin</collectingRegion>
<quantity id="4BB555C741F9C5697185F9862DA5CAE4" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</quantity>
<date id="53FFBB206F53B1EE42D7028C9EA4BAF4" value="2008-06-11" valueMax="2008-06-16" valueMin="2008-06-11">11. - 16. VI. 2008</date>
S. Murzin leg. Coll. Hendrich &quot; (
<abbrev id="ABBRID0EVRCK" xlink_title="Collection Lars Hendrich">
<collectionCode id="E9FB730560B624139D870D98E8AA9AF6">CLH</collectionCode>
</abbrev>
). •
<emphasis id="9A127EA5AEAAF308ABCD7D9231695989" bold="true">
<collectingCountry id="F7F162FE4AC102CD2E1387841A73D42A" name="Laos">Laos</collectingCountry>
</emphasis>
:
<specimenCount id="5F12979DABFE42DBC1FCE93C9879CE0E" count="7" type="generic">7 exs.</specimenCount>
, “ LAOS centr., Kham Mouan Prov. Nakai vill. env. ca
<quantity id="9010D12F8632D0F2E39F882196F47D0F" metricMagnitude="4" metricUnit="m" metricValue="7.0" unit="km" value="70.0">70 km</quantity>
NNE Muang
<collectingRegion id="D777BFB477C11909AD64927FE7ACA52E" country="Laos" name="Khammouan">Khammouan</collectingRegion>
,
<quantity id="382C1BAE5949C4C252621AE0C086349F" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500600 m</quantity>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5RCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.150000,17.716667]}">
<geoCoordinate id="C0ADEB321F48160D0D9FAF47B2B50F0C" degrees="17" direction="north" minutes="43" orientation="latitude" precision="925" value="17.716667">17 ° 43 ' N</geoCoordinate>
,
<geoCoordinate id="B0ED00A437494422B7726071C683F787" degrees="105" direction="east" minutes="09" orientation="longitude" precision="925" value="105.15">105 ° 09 ' E</geoCoordinate>
</named-content>
,
<date id="3AFA8F533CE0ADE0AB216BFB7C2A3AAB" value="2002-05-07" valueMax="2002-05-25" valueMin="2002-05-07">7. - 25. V. 2002</date>
M. Strba leg. / Coll. HENDRICH ” (
<abbrev id="ABBRID0EDSCK" xlink_title="Collection Lars Hendrich">
<collectionCode id="11AA4376BD1ADE39FD6DBBBDD8065C3A">CLH</collectionCode>
</abbrev>
);
<specimenCount id="51DD228C469AD0C77F4010D875A9F84D" count="1" type="generic">1 ex.</specimenCount>
, “ LAOS south,
<collectingRegion id="4FEA70724C22A4A90F9B8F49234515A7" country="Laos" name="Attapu">Attapeu prov.</collectingRegion>
Bolaven Plateau
<date id="A972AA5CE3A4A8FB04337757388EA58B" value="1999-04-18" valueMax="1999-04-30" valueMin="1999-04-18">18. - 30. IV. 1999</date>
,
<quantity id="5758B45D4D5A22DBA5E29FEB387EFE35" metricMagnitude="4" metricUnit="m" metricValue="1.5" unit="km" value="15.0">15 km</quantity>
SE of Ban Houaykong, NONG LOM (lake) env., N
<geoCoordinate id="D969028EC933ED73ACE52B99E230B27E" degrees="15" direction="east" minutes="02" orientation="longitude" precision="925" value="15.033334">15 ° 02´E</geoCoordinate>
106 ° 35´, alt.
<quantity id="E6131288383B8BB18C0B876A82A5CFC3" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
, E. Jendek &amp; O. Sausa leg. ” (
<abbrev id="ABBRID0EHSCK" xlink_title="Collection Lars Hendrich">
<collectionCode id="70DA8D72CFF45CCB69F04130CD340DF1">CLH</collectionCode>
</abbrev>
);
<specimenCount id="EA7ED9459BB6FBFD82741B201C266524" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="DC9C3508093623CCC427C6EB51F49486" country="Laos" name="Champasak">Champasak Prov.</collectingRegion>
, Ban Nam Touad env. (near Xe Katamtok), village, agricultural lands, old secundary and degraded primary forest, light trapping,
<quantity id="093BC930377058ED867EA1F300C46918" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOSCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.583333,15.100000]}">
<geoCoordinate id="E5CDD54C2CF7849AF3F7AB4F69CCD0F4" degrees="15" direction="north" minutes="06" orientation="latitude" precision="925" value="15.1">15 ° 06 ' N</geoCoordinate>
,
<geoCoordinate id="C3F1BACDFF1182B3891F8A9355A6F0FE" degrees="106" direction="east" minutes="35" orientation="longitude" precision="925" value="106.583336">106 ° 35 ' E</geoCoordinate>
</named-content>
,
<date id="75198A24A0C4DC60DA45309465126B3E" value="2010-06-08" valueMax="2010-06-10" valueMin="2010-06-08">8. - 10. VI. 2010</date>
, leg. M. Geiser &amp; D. Hauck ” (
<collectionCode id="793CAB20F58D238154CA5C6D4036849F">NHB</collectionCode>
);
<specimenCount id="A664595FD601ABB209BF95A2103DA59A" count="1" type="generic">1 ex.</specimenCount>
, “ Northern
<collectingRegion id="4B10C83C8671D518880BAB1F0C6C60D8" country="Laos" name="Viangchan">Vientiane Prov.</collectingRegion>
Vang-Vieng,
<quantity id="094647C7983F17E6CEB0F8BD3D9339DD" metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">300 m</quantity>
, N
<geoCoordinate id="2E3E389F10DBB62679FAC554E9928FEF" degrees="18" direction="east" minutes="55.23" orientation="longitude" precision="9" value="18.9205">18 ° 55.23 E</geoCoordinate>
102 ° 26.55, 10. - 15. V. &amp;
<date id="FE4206D370DF16FAC95B053A937F881E" value="2001-06-01" valueMax="2001-06-06" valueMin="2001-06-01">1. - 6. VI. 2001</date>
, J. Kolibac leg. ” (
<abbrev id="ABBRID0ESSCK" xlink_title="Naturhistorisches Museum Basel">
<collectionCode id="023A73E1682CEB730C729A1628206C03" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
</abbrev>
);
<specimenCount id="76578AC470A3C6044B30F5FEEB0B0D2A" count="1" type="generic">1 ex.</specimenCount>
, “ Umg.
<collectingRegion id="5DC14B03972574398EE51C874FFC8161" country="Laos" name="Viangchan">Vientiane</collectingRegion>
,
<date id="41FAC2462A0290C9515EB38C29134AC1" value="1963-03" valueMax="1963-04-31" valueMin="1963-03-00">III. - IV. 1963</date>
, coll. M. Brancucci ” (
<abbrev id="ABBRID0EWSCK" xlink_title="Naturhistorisches Museum Basel">
<collectionCode id="B5E5189D763EF303C31D6F02361B7840" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
</abbrev>
);
<specimenCount id="296F5ABA7516750BBA1B14D8C4673D82" count="2" type="generic">2 exs.</specimenCount>
, “
<collectingRegion id="5ACF78DCCE90D23ECD34836B473E7437" country="Laos" name="Khammouan">Khammouan Prov.</collectingRegion>
, Ban Khoun Ngeun,
<quantity id="B883144ED46CD15299C19D8FB34A0D56" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4SCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[104.483333,18.116667]}">
<geoCoordinate id="51E03DFC004499F6D8D6AF6A97100E5C" degrees="18" direction="north" minutes="07" orientation="latitude" precision="925" value="18.116667">18 ° 07 ' N</geoCoordinate>
<geoCoordinate id="AE1B0598CEE628FF08E13EFA14681147" degrees="104" direction="east" minutes="29" orientation="longitude" precision="925" value="104.48333">104 ° 29 ' E</geoCoordinate>
</named-content>
,
<date id="115CE18A9073D08550C10316CED709F2" value="2001-04-24" valueMax="2001-04-29" valueMin="2001-04-24">24. - 29. IV. 2001</date>
, V. Kuban leg. ” (
<abbrev id="ABBRID0ECTCK" xlink_title="Naturhistorisches Museum Basel">
<collectionCode id="DA2E5F137367EB11427F959322493A3F" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
</abbrev>
);
<specimenCount id="9C5755A94E9733A683981C4464FF7DDE" count="8" type="generic">8 exs.</specimenCount>
, “
<collectingRegion id="F870B11B427FE9FD16B111A133E67EC5" country="Laos" name="Khammouan">Khammouan Prov.</collectingRegion>
, Nakai env.,
<quantity id="47831550876381FF28520D7F2A8D138B" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500600 m</quantity>
,
<date id="096B98F8EAED94482F2FCC59667B1F18" value="2001-05-22" valueMax="2001-06-08" valueMin="2001-05-22">22. V. - 8. VI. 2001</date>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJTCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.150000,17.716667]}">
<geoCoordinate id="FFC6FE09C0B69BF5879D9E2634093FE9" degrees="17" direction="north" minutes="43" orientation="latitude" precision="925" value="17.716667">17 ° 43 ' N</geoCoordinate>
<geoCoordinate id="E8AB2D691011CF1335003241CEEDCCD6" degrees="105" direction="east" minutes="09" orientation="longitude" precision="925" value="105.15">105 ° 09 ' E</geoCoordinate>
</named-content>
, E. Jendek &amp; O. Sausa leg. ” (
<abbrev id="ABBRID0EOTCK" xlink_title="Collection Hans Hebauer">
<collectionCode id="2998B150F44D818B0A44DF6B3265F4D7">CHH</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0ESTCK" xlink_title="Národní muzeum, Praha">
<collectionCode id="062EF3170BFF8BF28C31997C34162468" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
</abbrev>
);
<specimenCount id="9C4CB5CADD53C748AC09754BC7915A33" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="3B994A420C06A2C0384DFBC3B16182DD" country="Laos" name="Champasak">CHAMPASAK prov.</collectingRegion>
, Bolavens Plateau, waterfall ca.,
<quantity id="0BF105127AA3C8B5A9F83034EE050641" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
E TAD KATAMTOK,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZTCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.646667,15.135000]}">
<geoCoordinate id="ED1065929894D692617B9C00306E470A" degrees="15" direction="north" minutes="08.1" orientation="latitude" precision="92" value="15.135">15 ° 08.1 ' N</geoCoordinate>
,
<geoCoordinate id="44FA6B4617F32AA08F97FDD72305DC54" degrees="106" direction="east" minutes="38.8" orientation="longitude" precision="92" value="106.64667">106 ° 38.8 ' E</geoCoordinate>
</named-content>
,
<quantity id="C135F8532FDB974A3489B7BD246E9A1A" metricMagnitude="2" metricUnit="m" metricValue="4.15" unit="m" value="415.0">415 m</quantity>
,
<date id="FA1070DAF652DCDF609B52F7469F4879" value="2010-05-10" valueMax="2010-05-12" valueMin="2010-05-10">1012. V. 2010</date>
, J. Hájek leg. ” (
<abbrev id="ABBRID0E5TCK" xlink_title="Národní muzeum, Praha">
<collectionCode id="8D68F541F84DAC2008BA57FB28012A64" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
</abbrev>
). •
<emphasis id="BE19DF1BE2EEA3FA6D338A1081E78507" bold="true">
<collectingCountry id="155C4D8E45A92F327D3661019B33849F" name="Thailand">Thailand</collectingCountry>
</emphasis>
:
<specimenCount id="8D81871F22416B8FAB1450A69D93F1BE" count="1" type="generic">1 ex.</specimenCount>
, “ N-Thailand,
<collectingRegion id="E65FCD3DF8818AFC9B68F37D1CC0BD81" country="Thailand" name="Khon Kaen">Khon Kaen Prov.</collectingRegion>
, Si Chom Phu,
<quantity id="B41B8CE70FE4E0E7583E629AF96E3052" metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">220 m</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHUCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[144.100000,158.183333]}">16 ° 8531 N, 102 ° 2526 E</named-content>
R. Ohnesorge,
<date id="418E76EBD0EB9188ED75DD2D88DF7EEE" value="2012-09">IX. 2012</date>
”, “ coll. A. Skale Hof / Germany ” (
<abbrev id="ABBRID0EMUCK" xlink_title="Collection André Skale">
<collectionCode id="A67A366EE42F5D2DE963ED1D6D9D5B22" country="USA" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
</abbrev>
);
<specimenCount id="E17E58F3728520DE6EC3EE67B05DF2D3" count="1" type="generic">1 ex.</specimenCount>
, “ Thailand 25 V 52 REElbel ”, “
<collectingRegion id="9085CA761BF462C9FD10D4790D2742A0" country="Thailand" name="Kanchanaburi">Kanchanaburi</collectingRegion>
” (
<abbrev id="ABBRID0EQUCK" xlink_title="United States National Museum of Natural History">
<collectionCode id="23CFC4B4C70681098CF5D2281DC25AAB" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</abbrev>
),
<specimenCount id="E3AFA98F20DC5BDCED179BC0618F2047" count="14" type="generic">14 exs.</specimenCount>
, “
<collectingRegion id="F689730D9F9F9CB868A668347F194686" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima Province</collectingRegion>
, Sakaerat Biosphere Reserve, King Cobra Cave, Thailand Coords:
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXUCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.932017,14.508933]}">
<geoCoordinate id="9599803E13ED0E1FEC20A3BBDA4805C7" degrees="14" direction="north" minutes="30.536" orientation="latitude" precision="1" value="14.508933">14 ° 30.536 N</geoCoordinate>
,
<geoCoordinate id="4C704BCE147C82AC77F7EA0AE77F0A93" degrees="101" direction="east" minutes="55.921" orientation="longitude" precision="1" value="101.932014">101 ° 55.921 E</geoCoordinate>
</named-content>
,
<quantity id="406CB2F6D58D0264BD12C8FFC0E2AFD2" metricMagnitude="2" metricUnit="m" metricValue="3.62" unit="m" value="362.0">362 m</quantity>
a. s. l.,
<date id="F747C422D2F2BBB9C6C8385532330D25" value="2013-10-27">27 Oct 2013</date>
, W. Atthakor leg., Sakaerat Expedition, led by Professor Somsak Panha of Chulalongkorn University ” (
<abbrev id="ABBRID0E3UCK" xlink_title="Collection Srinakharinwirot University">
<collectionCode id="9D024A2F388E107A1BEA0FC551F139E0">CSUT</collectionCode>
</abbrev>
);
<specimenCount id="E8C92339E4550D67502F69D812DF6739" count="1" type="generic">1 ex.</specimenCount>
, “ N Thailand, Doi Pho Ka N. P. Road from Pua
<quantity id="340EC1812332B42BD9228A4CAF4561BC" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
after Park Hq. m 1400, small pools on rock (24)
<date id="E02F5F0B523B6316FEDC31FFAD80BC7A" value="1999-01-02">2. I. 1999</date>
P. Mazzoldi leg. ” (
<abbrev id="ABBRID0EAVCK" xlink_title="Collection Paolo Mazzoldi">
<collectionCode id="95349AE842FBF6EEA2034983540BB56D" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" type="">CPM</collectionCode>
</abbrev>
);
<specimenCount id="C26083A2D3B84590B77C834F164C8810" count="2" type="generic">2 exs.</specimenCount>
, “ Siam: MeSong Forest
<date id="19F180B6791BE25FDB968E59CB9882E3" value="1919-04">IV. 1919</date>
E. J. Godfrey 1920-244 ” (
<abbrev id="ABBRID0EEVCK" xlink_title="Natural History Museum, London">
<collectionCode id="C8F4045B88A1772EB2981C8281E07242" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
);
<specimenCount id="9190F342D5B8BBD815CA91530DE1EC0F" count="2" type="generic">2 exs.</specimenCount>
, “ E Thailand, Ko Chang White Sands Beach,
<date id="9FFB39E25855F46BC9B81883ECACE359" value="1990-12-10">10. XII. 1990</date>
, M. Jäch leg. (12) ” (
<abbrev id="ABBRID0EIVCK" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="E90ACABB0EF4B949ED1506A71CF978B4" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
);
<specimenCount id="D6DA65FC9BFC076AEE57B3E3B468CE1D" count="1" type="generic">1 ex.</specimenCount>
, “ Prov.
<collectingRegion id="F6FA7310CC91E53CB2C207BD192F2009" country="Thailand" name="Rayong">Rayong</collectingRegion>
, Khao Chamao NP, 12. &amp;
<date id="D41D4F6FE95DE580811DBF309DCD736A" value="1990-12-13">13. XII. 1990</date>
, M. Jäch leg. (14) ” (
<abbrev id="ABBRID0EMVCK" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="1835D8CB81E1604B436BA8E9A187D6D7" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
);
<specimenCount id="371744733ACFEFC327B98D2B63B12952" count="1" type="generic">1 ex.</specimenCount>
, “ Tham Sakoen NP, No. 21,
<date id="9A08636EDB444E255ED9F3249529C27F" value="2003-11-29" valueMax="2003-11-30" valueMin="2003-11-29">29. - 30. XI. 2003</date>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETVCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[100.633333,19.383333]}">
<geoCoordinate id="6148ED9717F0381D921A45E4394BC61D" degrees="19" direction="north" minutes="23" orientation="latitude" precision="925" value="19.383333">19 ° 23 ' N</geoCoordinate>
<geoCoordinate id="A056A4F7BFD3783EA3980DBA6F8A0F66" degrees="100" direction="east" minutes="38" orientation="longitude" precision="925" value="100.63333">100 ° 38 ' E</geoCoordinate>
</named-content>
, Peregovits, Foldvari, Korosi, Szappanos &amp; Maklari-Kis leg. ” (
<abbrev id="ABBRID0EYVCK" xlink_title="Természettudományi Múzeum">
<collectionCode id="8CDD494D32E03A720CBECE450E749AF8">TDMB</collectionCode>
</abbrev>
);
<specimenCount id="499A1F85DE43CE516F260FA6694370D8" count="2" type="generic">2 exs.</specimenCount>
, “ Pak Lag,
<date id="31CB87E2C96859F02AC46675ED8FF609" value="1918-08-14">14. VIII. 1918</date>
, Jeanvoine ” (
<abbrev id="ABBRID0E3VCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="CCA619C70A337B126C286E7006C0A33E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="4F0760DBE384BE88F97EA812AEC900BE" count="17" type="generic">17 exs.</specimenCount>
, “ Pak Lag
<date id="345EEA43913434CDD902A90F24E3AEBF" value="1918-08-13">13. VIII. 1918</date>
V. de Salvaza ” (
<abbrev id="ABBRID0EAWCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="CC181FECF6759311032525CA0A2A2EDD" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="8579363A2F6AACFF0AB0DB1ACFDAD076" count="6" type="generic">6 exs.</specimenCount>
, “ Pak Lag,
<date id="E8CEEF45C24D477C7CD0B4C2E6CE2DD7" value="1918-08-13">13. VIII. 1918</date>
V. De S. ”, “ Samml. A. Zimmermann ” (
<abbrev id="ABBRID0EEWCK" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="C9FF8B86B0AF7A30F558CE69F3394698" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
);
<specimenCount id="BBEBCB59423F760FA039202D3EE7E498" count="3" type="generic">3 exs.</specimenCount>
, “ Pak Lag
<date id="B6157E76BA22D88268DD9FB7C26FF14C" value="1918-07-18">18. VII. 1918</date>
no.
<date id="CC28940BE7E478CDF4C320A9A91F1437" value="2119-05">2119 V.</date>
de S. ” (
<abbrev id="ABBRID0EIWCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="B12CA60175116B5D1C6DE67FC4A1F0DD" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0EMWCK" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="C19D2692BA195EC476B3F60BB60775DA" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
).
<specimenCount id="A028E279A6228EA09DEFC8AC0F7989DF" count="2" type="generic">2 exs.</specimenCount>
, “ Thailand, E
<collectingRegion id="8AF61041F28E7DBF0BF53D99878BC95F" country="Thailand" name="Krung Thep Mahanakhon">Bangkok</collectingRegion>
,
<collectingRegion id="413941C938B9386999A703778F18AAC8" country="Thailand" name="Nakhon Nayok">Nakhon Nayok</collectingRegion>
: Khlong Maduea,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETWCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.272778,14.354722]}">
<geoCoordinate id="489359E71036E9D166ABE7AD95203099" degrees="14" direction="north" minutes="21" orientation="latitude" precision="15" seconds="17" value="14.354723">14 ° 21 ' 17 &quot; N</geoCoordinate>
,
<geoCoordinate id="1876E2388F334DDA8FAA80A29033FA15" degrees="101" direction="east" minutes="16" orientation="longitude" precision="15" seconds="22" value="101.27278">101 ° 16 ' 22 &quot; E</geoCoordinate>
</named-content>
,
<date id="F5BA1D9079DDB5D0D082EB168D9A92B9" value="2017-03-16">16. 03. 2017</date>
, leg. H. Shaverdo ” (
<abbrev id="ABBRID0EYWCK" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="00AF149D71FC14565FF0FD55C28B875D" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
);
<specimenCount id="32AE20B844F52C7F7E716CFBEA94FA23" count="3" type="generic">3 exs.</specimenCount>
, “ Thailand,
<collectingRegion id="E889C74CC1B27C4AA56084E21236383A" country="Thailand" name="Phetchabun">Phetchabun Province</collectingRegion>
, near Nam Nao National Park, Route 2216 then small road off, in village Khlong Choen,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6WCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.733611,16.674722]}">
<geoCoordinate id="F3AE8D2E724380177DEBE7C9BDA24EB8" degrees="16" direction="north" minutes="40" orientation="latitude" precision="15" seconds="29" value="16.67472">16 ° 40 ' 29 &quot; N</geoCoordinate>
,
<geoCoordinate id="061FB7C39F0FF001BD2881FD75E11BCA" degrees="101" direction="east" minutes="44" orientation="longitude" precision="15" seconds="01" value="101.733604">101 ° 44 ' 01 &quot; E</geoCoordinate>
</named-content>
<date id="846B4A51E57C00ADF8CB94D253D76701" value="2017-03-21">21. 03. 2017</date>
, leg. H. Shaverdo ” (
<abbrev id="ABBRID0EEXCK" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="2AE782B1946CFC1D24354A187F6F6733" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
). •
<emphasis id="F1710B4A10F054E6DFECBC1C4AF69229" bold="true">
<collectingCountry id="33AA1218974F3337B59A98D3D6597436" name="Vietnam">Vietnam</collectingCountry>
</emphasis>
:
<specimenCount id="A0841B2CFB878C1A878916BDAE5B15CD" count="1" type="generic">1 ex.</specimenCount>
, “ VietnamN,
<collectingRegion id="CF0933C48294A917F68818CC18774F09" country="Vietnam" name="Quang Binh">Quang Binh prov.</collectingRegion>
<quantity id="371F644C776BDB6DCB112241BF5A0C49" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
N of Cha Lo,
<quantity id="52457E189A714CC85563E214BBF92D8E" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</quantity>
Vietnam-Laos border area
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENXCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.762500,17.689444]}">
<geoCoordinate id="B15F694A47D9EA162489608FD4F78B7E" degrees="17" direction="north" minutes="41" orientation="latitude" precision="15" seconds="22" value="17.689444">17 ° 41 ' 22 ´´ N</geoCoordinate>
<geoCoordinate id="88E39515F134FD607A7216928A77D10B" degrees="105" direction="east" minutes="45" orientation="longitude" precision="15" seconds="45" value="105.7625">105 ° 45 ' 45 ´´ E</geoCoordinate>
</named-content>
, L. Dembicky leg.,
<date id="07A908C43F6C8F045E567C6860E629F6" value="2010-04-11" valueMax="2010-04-24" valueMin="2010-04-11">11. - 24. iv. 2010</date>
” (
<abbrev id="ABBRID0ESXCK" xlink_title="Naturhistorisches Museum Basel">
<collectionCode id="F48850E9DC794A52078BA53E42DB7876" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
</abbrev>
);
<specimenCount id="B966029A4003A1CABDA82BFB0F2EFB68" count="1" type="generic">1 ex.</specimenCount>
, “ Tonkin Env. De
<collectingRegion id="221E886392F79CAA7976FC98708807CE" country="Vietnam" name="Hoa Binh">Hoa Binh</collectingRegion>
J. Laisi 1902 “, “
<taxonomicName id="519DA1A4B28D25E0C3A3D4C35C13909B" authority="Wke" authorityName="Wke" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="6C83B59B1C9965B9098FC26E43651994" italics="true">S. maculatus</emphasis>
Wke
</taxonomicName>
”, “ Dr. Régimbart vidit 1905 ” (
<abbrev id="ABBRID0EAYCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="F08F81E09E95BBDD2A6BEE7DBF3D87FB" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="E0547CCD588D686390007CE52479677D" count="1" type="generic">1 ex.</specimenCount>
, “ Museum Paris Cochinchine ” (
<abbrev id="ABBRID0EEYCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="4FF73FF561E8E527A1671A607C0C8FDB" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="2E1BC14A538CB74B50FA042E118D4FAF" count="3" type="generic">3 exs.</specimenCount>
, “
<collectingRegion id="BEE7666ABA3C4A487B778F555AEA9030" country="Vietnam" name="Hoa Binh">Hoa Binh</collectingRegion>
Tonkin ” (
<abbrev id="ABBRID0EIYCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="ACBB9846D6618B3C142FDE0BCCEF0E9B" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="AA4C138C8CE2BCB522B747F0539D2716" count="3" type="generic">3 exs.</specimenCount>
, “ Lactho Tonkin A. de Cooman ” (
<abbrev id="ABBRID0EMYCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="C84140B56377102FA96FC3C50D63D770" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="864F14A73FF497ED2AE39F88690EC27E" count="1" type="generic">1 ex.</specimenCount>
, “ S Vietnam,
<collectingRegion id="DBFE41C6B37A0BA45361724A637F4B23" country="Vietnam" name="Ho Chi Minh City">Saigon</collectingRegion>
, jardin botanique [botanical garden],
<date id="3F49F050B031E756F814C0B1ADB5968B" value="1870-10">octobre 1870</date>
, A. Krempf leg. ” (
<abbrev id="ABBRID0EQYCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="0C0781BA7BD8AB70EF1EA219195DD275" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="6C683E99BEC9EF586B79FAB15E13BF3F" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="3338AAD0BDC89292C1185EB7FE9DCE5A" country="Vietnam" name="Hoa Binh">Hoa Binh</collectingRegion>
(Tonkin) A. de Cooman ” (
<abbrev id="ABBRID0EUYCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="16219173992FA34155DFA86E778498BF" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="D077D133C96308DADFB9641DC7D89C0D" count="3" type="generic">3 exs.</specimenCount>
, &quot;
<collectingRegion id="06869DBD0FA0F4917CDAC1A85666112D" country="Vietnam" name="Hoa Binh">Hoa Binh</collectingRegion>
(Tonkin) A. de Cooman ” (
<abbrev id="ABBRID0EYYCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="74DA378460527D5DD6A8F70845E015F2" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="730629E8B5A45B1046EB7BE8CE405FC4" count="2" type="generic">2 exs.</specimenCount>
, “ Tonkin occ. env. de
<collectingRegion id="08FF01FCB1F29C938F348BF2CDC5FCCB" country="Vietnam" name="Hoa Binh">Hoa Binh</collectingRegion>
1919 A. de Cooman ” (
<abbrev id="ABBRID0E3YCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="8C4BFBD8B72F32C236ED26136AB0EA91" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="5E94B324D8C819405CF35AEC8E0AEF6C" count="4" type="generic">4 exs.</specimenCount>
, “ S-Vietnam, Nam Cat Tien NP,
<date id="59A1C437F00504BF56DBEA2857AFF28E" value="1994-05-01" valueMax="1994-05-15" valueMin="1994-05-01">1. - 15. V. 1994</date>
, Pacholatko &amp; Dembicky leg. ” (
<abbrev id="ABBRID0EAZCK" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="09D85BDFAE04C625C41132ABAF39CE3E" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
);
<specimenCount id="A7B5AC53EB43C3F7289BF7E2257EA960" count="1" type="generic">1 ex.</specimenCount>
, “ Vietnam,
<collectingRegion id="2A28C3CCA4D5C344B72CE5BF720F4F13" country="Vietnam" name="Dong Nai">Dong NAI</collectingRegion>
, Nam Cat Tien NP,
<quantity id="D193F53CA513B76C2A59C40FD6454ECF" metricMagnitude="2" metricUnit="m" metricValue="1.2" unit="m" value="120.0">120 m</quantity>
,
<date id="558E8F86720B108F6F665B9696D4E68F" value="1998-09-18">18 - IX- 1998</date>
, leg. L. J. Wang ” (
<abbrev id="ABBRID0EEZCK" xlink_title="Collection Liang-Jong Wang">
<collectionCode id="28E22F9428BC1475998E3374E4F7161A">CLJW</collectionCode>
</abbrev>
). •
<emphasis id="19CF84D5382C2BB976F3216BDCA9DE67" bold="true">
<collectingCountry id="F16153E93507799CE26B2342B9B380A1" name="Malaysia">Malaysia</collectingCountry>
</emphasis>
: &quot; E-Malaysia,
<collectingRegion id="5CE01AEB8EFC2E9DAE027AA22A02A555" country="Malaysia" name="Sabah">Sabah</collectingRegion>
Borneo, Mt. Trus [= Trusmadi or Trus Madi] March-April 2010 Local leg. ”, “ coll. A. Skale Hof / Germany ” (
<abbrev id="ABBRID0EKZCK" xlink_title="Collection André Skale">
<collectionCode id="409649FDEB633422FE2A0CDE7AE2681A" country="USA" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
</abbrev>
);
<specimenCount id="5190CD841692E355FD9D9ACE91D46127" count="1" type="generic">1 ex.</specimenCount>
, “ NE Borneo Sandakan, Coll. Guignot ” (
<abbrev id="ABBRID0EOZCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="E6A03DF0F7EF039AA8B268F628585E54" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="216640DC8E542B501D44EA5D5E76E19D" count="1" type="generic">1 ex.</specimenCount>
, “
<taxonomicName id="D12824DFD6D9F1D67772B03D2DDEA218" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="04267DD010D2BA8655FC4765AE333A66" italics="true">Hydaticus</emphasis>
sp.
</taxonomicName>
?
<collectingRegion id="D35D6F82EF7B2931DBFE7087C28374E5" country="Malaysia" name="Pulau Pinang">Pinang</collectingRegion>
&quot; [West Malaysia,
<collectingRegion id="850E9A491F97858C8858ED0C775544C6" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
], “ maculatus Wehncke ” [handwritten label by Régimbart] (
<abbrev id="ABBRID0EYZCK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="9BC0F8B951DAAACA92BD5947388CE7F0" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="0CCBEFA5F5C732894314CA20BD0B2AAD" count="1" type="generic">1 ex.</specimenCount>
, “ Borneo Sandakan Windrath ”,
<taxonomicName id="28E9FA04258D29886D1040DCC46277D9" authority="Shp" authorityName="Sharp. In" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nauticus">
<emphasis id="A8A3FBCB8C6364DAC906FBA00D7AD80F" italics="true">nauticus</emphasis>
Shp
</taxonomicName>
” (
<abbrev id="ABBRID0EF1CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="0C533B33E8B03DF89B90B19325EF78E9" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
),
<specimenCount id="D911172C23D5D431665A5772A5868FF7" count="1" type="generic">1 ex.</specimenCount>
, “ Borneo, Kinabalu [
<collectingRegion id="5F47F40067CF3BCE2E7FF9EAEC826869" country="Malaysia" name="Sabah">Sabah</collectingRegion>
], Whitehead, Fry Coll. 1905-100 ” (
<abbrev id="ABBRID0EJ1CK" xlink_title="Natural History Museum, London">
<collectionCode id="0271CD8EF48517594D9E09906A170695" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
);
<specimenCount id="2391430A0285586204CA4B57120D0076" count="1" type="generic">1 ex.</specimenCount>
, “ Borneo 987, Sharp Coll. 1905-313 ” (
<abbrev id="ABBRID0EN1CK" xlink_title="Natural History Museum, London">
<collectionCode id="323182C5E208A76F4158A5B818974A7A" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
);
<specimenCount id="11524C58153FE4898DEB85C5C85345CB" count="3" type="generic">3 exs.</specimenCount>
, “ Borneo,
<collectingRegion id="0CF8235924A632773CF2F0BFC8B3B0AF" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, Kampung Pisang, Pisang env., tributary of Kuamut river,
<date id="F5EB2E4961F527C3599EC81856F23848" value="1998-06-29">29. VI. 1998</date>
, J. Kodada &amp; F. Ciampor leg. ” (
<abbrev id="ABBRID0ER1CK" xlink_title="Naturhistorisches Museum Wien">
<collectionCode id="0E4890466E2FA9B1F21515C2D897EA18" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
</abbrev>
);
<specimenCount id="56C10738C32ACAA28C5D9E59F32A20E3" count="1" type="generic">1 ex.</specimenCount>
, “ Borneo
<collectingRegion id="A35504903299B9221D3FF62A176417BB" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, Hot Springs,
<date id="9596435BEF9CDA6B2316F5936AABE40E" value="1999-05-26">26. V. 1999</date>
, Bacovský leg. ” (
<abbrev id="ABBRID0EV1CK" xlink_title="Národní muzeum, Praha">
<collectionCode id="D7FE8A86EF1F4ED7C4A5937E3E3ACF78" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
</abbrev>
);
<specimenCount id="A81FE27E7279334C300AD910364CC783" count="20" type="generic">20 exs.</specimenCount>
, “ Borneo,
<collectingRegion id="964934CAECE6C2EE2750662734006491" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, Sungai Kinabatangan, stream near Danau Blandung Besar,
<date id="4CC58B2C74569C65937E1A76909357D7" value="1994-04-11">11. IV. 1994</date>
, H. K. Lua leg. ” (
<abbrev id="ABBRID0EZ1CK" xlink_title="Collection Lars Hendrich">
<collectionCode id="83B2142596266AA69E5EC99C99BFFF58">CLH</collectionCode>
</abbrev>
,
<abbrev id="ABBRID0E41CK" xlink_title="Zoological Reference Collection, Lee Kong Chian Natural History Museum National University of Singapore">
<collectionCode id="0E495D7BF8A525D357E6FBC9A5A39425" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" type="">ZRC</collectionCode>
</abbrev>
);
<specimenCount id="6B7D57B3E7E4E774979A775D9F76DC6B" count="2" type="generic">2 exs.</specimenCount>
, “
<collectingRegion id="B8BD1D9422C97BE6EF7E64E926C3E64E" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, Kinabatangan riv.
<date id="5F38F1777DB46D77EFCCAEEA97AE9CE1" value="2003-06-08" valueMax="2003-06-15" valueMin="2003-06-08">8. - 15. VI. 2003</date>
, Uncle Tan´s camp, J. Šťastný leg. ” (
<abbrev id="ABBRID0EB2CK" xlink_title="Collection Jaroslav Šťastný">
<collectionCode id="9B0EB9E5345840A92FF2E9753000C277">CJS</collectionCode>
</abbrev>
);
<specimenCount id="7CC5A03F4EAA7B2FBC67070EC86C25AE" count="1" type="generic">1 ex.</specimenCount>
, “ Dindings 96-85 ” [Malaysia,
<collectingRegion id="A17964B15F54097FB6B4785278FDEB35" country="Malaysia" name="Perak">Perak</collectingRegion>
, Manjung District], “ Determined by Dr Régimbart
<taxonomicName id="283B15B71CE081562F7EA02179594B93" authority="Sharp" authorityName="Sharp" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="variety" species="maculatus" variety="nauticus">
<emphasis id="7CFAC7AF61F584D8DC6F479FD2C04FCC" italics="true">Sandracottus maculatus</emphasis>
v.
<emphasis id="8D19324CCE088C471CB4E1F3405D7EA2" italics="true">nauticus</emphasis>
Sharp
</taxonomicName>
”;
<specimenCount id="43AA2750B5F7AA8E651646E7BE92FCD9" count="1" type="generic">1 ex.</specimenCount>
, “ Ex. Coll. E. Wehncke Acq 1864 ” (
<abbrev id="ABBRID0EY2CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="C2A09F7277FDF8F4C3042E4BA100FEC9" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="D49FAAFFDB8871B6D2A3F9F594ED9999" count="3" type="generic">3 exs.</specimenCount>
, “ West Malaysia,
<collectingRegion id="3CD153DD8D7D129713953D7FB998DFA0" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
988, Sharp Coll. 1905-313 ” (
<abbrev id="ABBRID0E32CK" xlink_title="Natural History Museum, London">
<collectionCode id="19C33E9916C381F9324CEE80C4B1B1D1" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
);
<specimenCount id="6058E1C8FC2976793F45E0F6716EE4FC" count="5" type="generic">5 exs.</specimenCount>
, “ West Malaysia,
<collectingRegion id="15B66A2F4897DEDA2FD990441AFD1DCB" country="Malaysia" name="Pulau Pinang">Penang</collectingRegion>
, Pascoe Coll. 93-60 ” (
<abbrev id="ABBRID0EA3CK" xlink_title="Natural History Museum, London">
<collectionCode id="8730A2967B3314CFD3D490CA2B5AE0C1" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</abbrev>
). •
<emphasis id="749E161C143139FF252A252E1EA10175" bold="true">
<collectingCountry id="3A8ED6E85333B9EC5DA3AD64D3E5EC2B" name="Indonesia">Indonesia</collectingCountry>
</emphasis>
:
<specimenCount id="8EBF9AC64D9E3F5F64776BC77C4061B4" count="1" type="generic">1 ex.</specimenCount>
, “ Ned. Indie Leg. A. Koller ”, “ Coll. F. C. Drescher ” (
<abbrev id="ABBRID0EG3CK" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="09B582B4FE62EDCA14E1BD21C273FC82" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
);
<specimenCount id="8E76EED29224ABCAEBB6118A06FF853F" count="5" type="generic">5 exs.</specimenCount>
, “
<collectingRegion id="BF471C3F5597976731AE5D5FB35E9AA4" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
Palembang M. Knappert ” (
<abbrev id="ABBRID0EK3CK" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="E56DB9858C49EE2807EF0BB45F6E950F" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
);
<specimenCount id="85784706B0AF7E3753B1D0595398610A" count="7" type="generic">7 exs.</specimenCount>
, “
<collectingRegion id="1F09C99B4D18A286109B21B5BA3A8EC3" country="Indonesia" name="Kalimantan Timur">East Kalimantan</collectingRegion>
, Distr. Damai
<date id="DD0B653FE1B86406CA8B04D7D93DAE94" value="1995-07-12">12. VII. 1995</date>
Ngelung Stream Mazzoldi P. leg. ” (
<abbrev id="ABBRID0EO3CK" xlink_title="Collection Paolo Mazzoldi">
<collectionCode id="191C1B372E58240F750BDC70EFA97BE6" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" type="">CPM</collectionCode>
</abbrev>
);
<specimenCount id="A7BF8DA839C48C1FA1F9A3DFE2EEC870" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="3D41F5A7C96DA929C34BA6E9A78A2A68" country="Indonesia" name="Kalimantan Timur">East Kalimantan</collectingRegion>
<date id="64D47BA91E3A5E7C36DA57EDD60F1259" value="1995-07-12">12. VII. 1995</date>
, Distr. Tingai stream near Sembuan, opening in forest leg. P Mazzoldi ” (
<abbrev id="ABBRID0ES3CK" xlink_title="Collection Paolo Mazzoldi">
<collectionCode id="CA1E2BD5D67D856E5DA15D6CC2D3969A" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" type="">CPM</collectionCode>
</abbrev>
);
<specimenCount id="6A99E54DDC8FD9D3ADF32ABBC5BEE831" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="8BB65530ED9292C3FF18F5D7424423DC" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
Occ. Toeroe 1902 ”, “ Museum Paris ex. Coll. Oberthür 1952 ” (
<abbrev id="ABBRID0EW3CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="AE30BC03322EA826E2AEDB5A88992E1D" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="5E9D162590A478A8A91A51A1C236124F" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="4461CEFD69B20C1B0F9B4FBFA0E207C4" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
ex. J. Waterstradt 1904 ” (
<collectionCode id="E75AD1A95DF80B7F7EE76CFB5AD32A85">NMHN</collectionCode>
);
<specimenCount id="822DE7ACB274D15F9E0233D28C1960C4" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="B5704F412FBF254A68CFEC6245D56030" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
”, “ maculatus Wehncke (
<abbrev id="ABBRID0E13CK" xlink_title="Muséum national d´Histoire naturelle, Paris">MNHN</abbrev>
);
<specimenCount id="BF3E58A993BD4E58AAE62A5BCE6CA6F4" count="1" type="generic">1 ex.</specimenCount>
, “ Malang
<collectingRegion id="8CF863EA21A04FE71DA576A8201F6EAD" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
” (
<abbrev id="ABBRID0E53CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="D371894E053E463FF037EAA406B84934" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="7A459299EDEE07ACB88F7F2ACDD7EAA5" count="1" type="generic">1 ex.</specimenCount>
, &quot;
<collectingRegion id="07063BD8D2A123D0AA54D3604734A06C" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
, Prov. Pasuruan Kalipari,
<quantity id="9D018C312C7D6B13E7037355A35E7E7D" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300500 m</quantity>
, 1891, W. Doherty leg. &quot; (
<abbrev id="ABBRID0EC4CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="1E138B07E9C1C450D01963E41E36243E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="EC155FBEA1D7FE615D9AE49CA859D152" count="1" type="generic">1 ex.</specimenCount>
, &quot;
<collectingRegion id="356D6B98AC2AE4390FD35ED004F50C4B" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
, Blume leg. &quot; (
<abbrev id="ABBRID0EG4CK" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="41B1A6564A3B30C3ED5751035AD18F9F" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
);
<specimenCount id="3A43F4404DCA2A059C450884D04203CE" count="1" type="generic">1 ex.</specimenCount>
, &quot;
<collectingRegion id="05BF07FE763F993AA14570E3502942E6" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
, Muller leg. &quot; (
<abbrev id="ABBRID0EK4CK" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="2C61A8A9F491F18E10BEB0EF927F3DB5" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
);
<specimenCount id="6E2D4BDC5E5C27482A990ED85B2D4B15" count="4" type="generic">4 exs.</specimenCount>
, “ Z. -
<collectingRegion id="9BF5C6383EB94CDBF4B1DF38F4B67D47" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
, Ranau,
<quantity id="8C68B2865D508C84449852CC93C0FA0B" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500700 m</quantity>
, Juli 35 ” (
<abbrev id="ABBRID0EO4CK" xlink_title="Nationaal Natuurhistorisch Museum">
<collectionCode id="FDD5EB3AF01C5A9DF493D85ADF783A63" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</abbrev>
);
<specimenCount id="CAC2A5BA5A3E8D95999621625E39CB2A" count="1" type="generic">1 ex.</specimenCount>
, “ SW
<collectingRegion id="20B8665FA40FFB3857EF3621C5E9CFEF" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
, Gautang Bay,
<date id="8D14F8B5A90D1FCCEAF9A2E09FD13DF4" value="1937-03">III. 1937</date>
, M. A. Lieftinck leg. ” (
<abbrev id="ABBRID0ES4CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="CE414D027374B5BEB2E78CFA95C77D76" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="C8F7665B200FD738C1555DF524219A4D" count="3" type="generic">3 exs.</specimenCount>
, “
<collectingRegion id="C4F0478B30F4C4A87EEA6CC6169A8191" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
, Palembang ” (
<abbrev id="ABBRID0EW4CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="2DDFFA434AA946980A0F8D8C4F565931" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="650A4FDB6F0A84F6A893272E21E52F72" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="50889BB7D608DC6BBE037BC6C8F3E346" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
, Dolok Merangir,
<date id="FB31DD6A064FDB15DB27C2B209579BB8" value="1983-05-08" valueMax="1983-06-10" valueMin="1983-05-08">8. V. - 10. VI. 1983</date>
, E. W. Diehl leg. ” (
<abbrev id="ABBRID0E14CK" xlink_title="Naturhistorisches Museum Basel">
<collectionCode id="89713C156039FDF14FACE802D7ED79A7" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" type="">NMB</collectionCode>
</abbrev>
);
<specimenCount id="CB796C04A6989E14B9D93C1884B84B63" count="1" type="generic">1 ex.</specimenCount>
, “ Mentawai Islands, Siberut,
<quantity id="5BBD763299C212A60203621379F68EDD" metricMagnitude="1" metricUnit="m" metricValue="2.75" metricValueMax="5.0" metricValueMin="0.5" unit="m" value="27.5" valueMax="50.0" valueMin="5.0">550 m</quantity>
,
<date id="4DA09EF7DEC173670B8EB2387F16F328" value="2005-04-03">3. IV. 2005</date>
, S. Jákl leg. ” (
<abbrev id="ABBRID0E54CK" xlink_title="Národní muzeum, Praha">
<collectionCode id="B722F369EE7DA8C4D550F2FD657802FD" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
</abbrev>
);
<specimenCount id="AD29966CE3CC904BD5C87F7BED499482" count="2" type="generic">2 exs.</specimenCount>
, “
<collectingRegion id="615891F75CC1847E6B583EA3D8997B4B" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
, Palembang Coll. Peschet ” (
<abbrev id="ABBRID0EC5CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="98DC8A92262C58581F3E2751EE686E02" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="9C09AAA94EAEC503DC2FDFB376990B82" count="2" type="generic">2 exs.</specimenCount>
, “
<collectingRegion id="E6CC145A8BFD8A789FC82C7DE12AE6C5" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
Palembang ”, “ Samml. A. Zimmermann ” (
<abbrev id="ABBRID0EG5CK" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="C9D959C04314AAC2A146B50AC28B79C2" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
);
<specimenCount id="87A5D59CACD73AF7A780A43B6E47F972" count="1" type="generic">1 ex.</specimenCount>
, “
<collectingRegion id="6E2FCB3CF193A6F669A82BA5FD3D0B21" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
Medan Hayek leg. (
<abbrev id="ABBRID0EK5CK" xlink_title="Zoologische Staatssammlung, München">
<collectionCode id="EEED8AE341D168ED75605D37269EE9FD" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" type="Museum">ZSM</collectionCode>
</abbrev>
). •
<emphasis id="CF2B41C3538CBAD86193DA49AF52B67E" bold="true">
<collectingCountry id="3600B19D55DE3698FDECC640386893EA" name="Philippines">Philippines</collectingCountry>
</emphasis>
:
<specimenCount id="85E6904C6A2281C614A46A1B1C9A456A" count="1" type="generic">1 ex.</specimenCount>
, “ Mindanao
<collectingRegion id="B88CD443C38315A33ECCBA25332D811B" country="Philippines" name="Davao">Davao</collectingRegion>
<date id="FC60DF5CEFA5EAC7AAC7D4460F34C16D" value="1932-12">XII. 1932</date>
, Baker leg. ” (
<abbrev id="ABBRID0EQ5CK" xlink_title="Staatliches Museum für Tierkunde">
<collectionCode id="C349668AAA382D655F34B84F4672F8F1">SMTD</collectionCode>
</abbrev>
);
<specimenCount id="D7AE0D9FFB0B99DA570736E43D74CE8B" count="1" type="generic">1 ex.</specimenCount>
, “ Mindanao ” (
<abbrev id="ABBRID0EU5CK" xlink_title="Muséum national d´Histoire naturelle, Paris">
<collectionCode id="E9DAFEE9230B15C0E12EB3313BF930B7" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</abbrev>
);
<specimenCount id="EB73BD15564BD742DD12A68F69A8E4CE" count="1" type="generic">1 ex.</specimenCount>
, “ Philippines: Luzon, Bicol,
<collectingRegion id="BEEA80027FC0812B00251E44153D6DA0" country="Philippines" name="Camarines Sur">Camarines Sur</collectingRegion>
, Lagonoy ca.
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E25CK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[123.521139,13.737694]}">
<geoCoordinate id="35507EEB8B66B17C9AC615E4F533571D" degrees="13" direction="north" minutes="44" orientation="latitude" precision="1" seconds="15.7" value="13.737695">13 ° 44 ' 15.7 &quot; N</geoCoordinate>
,
<geoCoordinate id="E75C5D1790705E45F5F1DF880CE7B298" degrees="123" direction="east" minutes="31" orientation="longitude" precision="1" seconds="16.1" value="123.52114">123 ° 31 ' 16.1 &quot; E</geoCoordinate>
</named-content>
09.2019 leg. local collector ” (
<abbrev id="ABBRID0EA6CK" xlink_title="Collection Leopold Wendlandt">
<collectionCode id="AECEC941DA2F40F318DB8A9D83D6B484">CLW</collectionCode>
</abbrev>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE6CK" type="remarks">
<paragraph id="D9BDF766A905F009A383836563E0B734">
<heading id="87537F3DF564DE3F55F9F09488001753" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="57752F05EB952B3A5B3A65D7D062B49A">
Specimens from Borneo and
<collectingRegion id="F92925C92F7AFE018BD3E044F2E2C4CF" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
with reduced yellow basal markings on elytra (habitus near
<taxonomicName id="32A7589BFE48E87F7C0806A292388A79" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="6BFE919E6038B03BEBFE23424F27D0BE" italics="true">femoralis</emphasis>
</taxonomicName>
) were described as
<taxonomicName id="5E56361407D3A538BC478B7DDABD672E" authority="Sharp. In" authorityName="Sharp. In" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nauticus">
<emphasis id="641521348C7624D7A745FEE1AFE2DD42" italics="true">S. nauticus</emphasis>
Sharp. In
</taxonomicName>
the shape of the median lobe of aedeagus, some specimens from Borneo lies between
<taxonomicName id="AA9ACF4A8A8F57AE447A8F62F1452304" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="1FB7DA28F5A4887345878D3767658882" italics="true">S. femoralis</emphasis>
</taxonomicName>
and
<taxonomicName id="AB020E764698940FD03081ABE36CA367" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="3F2A7216D0A171B4A1425A9EB9B7141A" italics="true">S. maculatus</emphasis>
</taxonomicName>
from mainland Asia. Furthermore, we do not have any molecular data for either form, so we here list
<taxonomicName id="132D8F12C8391905DC6707A8D45E46C4" authorityName="Sharp. In" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nauticus">
<emphasis id="5D87B3D7019EAC903EBB5382AA803E1F" italics="true">S. nauticus</emphasis>
</taxonomicName>
as an objective junior synonym of
<taxonomicName id="0163D7ABC1A00E01FF0D898E620AC8C3" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="A0DCDE8AAA02CA6A37B7A9DF949F9037" italics="true">S. maculatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENBDK" type="redescription">
<paragraph id="3CB26B2750FE370B1CC52133FA7BC2D3">
<heading id="0724FA411ED185901AE1B906872FC1CC" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="C5E4FBBB57D984702845CA842450F887">Body large, broad oval, submatt, only slightly shiny, black with ferrugineus brown markings. Ventral side completely dark brown to black, legs testaceous, hind legs somewhat darker.</paragraph>
<paragraph id="8460C4E061F9BA0A66EE72E41E8F0054">
Head testaceous with posterior part broadly black leaving free two small testaceous spots on vertex (Figs
<figureCitation id="8CBC269E1C9C1F1D6437B5ACA421FEE1" captionStart="Figures 811" captionStartId="F3" captionText="Figures 811. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">10</figureCitation>
,
<figureCitation id="03C268F3633F7138E5F8F303E8286D64" captionStart="Figures 5457" captionStartId="F20" captionText="Figures 5457. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">56</figureCitation>
). Surface submatt, shagreened, consisting of dense and minute punctures but not very uniform in size and of larger and much sparser ones, more numerous on frons. Clypeal grooves and punctures alongside eyes marked, punctures medium-sized but coalescent. Antennae testaceous; antennomeres slender, antennomere V 4 × as long as broad.
</paragraph>
<paragraph id="48650B68697016884C4EC9E99FFCD126">
Pronotum black with margins broadly testaceous, submatt (Figs
<figureCitation id="FF7558FAC39315C6A6D3184D7EAEA024" captionStart="Figures 811" captionStartId="F3" captionText="Figures 811. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">10</figureCitation>
,
<figureCitation id="EE932048E4C70BDF40DFAD66A2D5B55C" captionStart="Figures 5457" captionStartId="F20" captionText="Figures 5457. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">56</figureCitation>
). Surface distinctly shagreened, with very dense punctation; punctures medium-sized and distant of only 12 × their own diameter. Anterior puncture line interrupted in middle; punctures relatively small but coalescent, building wrinkles and becoming sparse towards middle and lacking in very middle of anterior margin. Posterior puncture line with coarse and coalescent punctures in middle of each sides building distinct wrinkles, distinctly smaller and spaced on disc.
</paragraph>
<paragraph id="C75C8214D6F123912CC59BBF2E153093">
Elytra black with testaceous markings consisting of one basal band sometimes reduced to different spots at base, one postmedian band, one preapical short and reduced band, and an apical spot (Figs
<figureCitation id="F17A706405DEA637AFB6EAD74B5EFB88" captionStart="Figures 811" captionStartId="F3" captionText="Figures 811. Habitus of 8 Sandracottus insignis (Luzon, Philippines) 9 S. jaechi (paratype from Nuwara Eliya, Sri Lanka) 10 S. maculatus (Thailand) 11 S. rotundus (Sulawesi, Togian Islands)." figureDoi="10.3897/zookeys.1223.138220.figures8-11" httpUri="https://binary.pensoft.net/fig/1214410">10</figureCitation>
,
<figureCitation id="73DE5C21EB3DA08219ED68B457D951DD" captionStart="Figures 5457" captionStartId="F20" captionText="Figures 5457. Habitus of 54 Sandracottus jaechi (holotype, Sri Lanka, Nuwara Eliya) 55 Sandracottus insignis (Philippines; Luzon, Los Banos) 56 S. maculatus (Borneo, Sabah, Sungai Kinabatangan) 57 S. rotundus (Indonesia, Sulawesi, Togian Islands, Kaldidiri Island)." figureDoi="10.3897/zookeys.1223.138220.figures54-57" httpUri="https://binary.pensoft.net/fig/1214427">56</figureCitation>
), submatt. Epipleura ferrugineus brown. Surface distinctly shagreened and covered with small and dense punctures as well as with larger and much more sparser ones. Puncture lines with groups of medium-sized punctures mostly grouped by five or six punctures; groups closer together on discal row. Sutural puncture line marked only by few punctures on apical part.
</paragraph>
<paragraph id="901F4E0301B603B8BD4E272081BAB9B2">Ventral side dark brown. Legs particularly fore and mid legs testaceous, hind legs ferrugineus brown to dark brown. Prosternal process short and broad, 1.4 × longer than broad, flattened finely but distinctly sculptured; posterior margin broadly rounded. Metatibia with sparse medium-sized punctures on whole surface. Ventrites II VI very superficially shagreened, slightly and longitudinally wrinkled on lateral parts, densely covered with very small punctures and larger sparser ones. Posterior margins rounded, deeply bordered with some large coalescent punctures on middle of each side along margin.</paragraph>
<paragraph id="B3E7631640EFD2827F46D49DE040788C">
Measurements:
<abbrev id="ABBRID0ETCDK" xlink_title="total length">TL</abbrev>
=
<quantity id="4FD3CBCDA5EE975F5FDD44BFBC5E3632" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.66" metricValueMin="1.34" unit="mm" value="15.0" valueMax="16.6" valueMin="13.4">13.416.6 mm</quantity>
,
<abbrev id="ABBRID0EXCDK" xlink_title="total length minus head length">TL-h</abbrev>
=
<quantity id="C711099DDDB8B4EFE2DDC02B424A7A71" metricMagnitude="-2" metricUnit="m" metricValue="1.4149999999999998" metricValueMax="1.52" metricValueMin="1.31" unit="mm" value="14.149999999999999" valueMax="15.2" valueMin="13.1">13.115.2 mm</quantity>
,
<abbrev id="ABBRID0E2CDK" xlink_title="maximum width">TW</abbrev>
=
<quantity id="B379C2D0526E32F9CA1776DD4E0923FB" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" unit="mm" value="9.25" valueMax="10.0" valueMin="8.5">8.510 mm</quantity>
.
</paragraph>
<paragraph id="E9557D1FA7E661B50D60A5A3B91446A4">
<specimenCount id="E74ABF1357BE3C0B4087D6E743EC6900" count="1" type="male"></specimenCount>
. Protarsomeres I III strongly enlarged with three larger suckers and numerous smaller one. Mesotarsomeres I III with two rows of small suckers. Median lobe of aedeagus, in ventral view, elongate, parallel-sided on complete length, lobes narrowly rounded at apex (Fig.
<figureCitation id="BF60F6E04DDFCEE6E01CC6AA1D4A5E07" captionStart="Figures 2022" captionStartId="F6" captionText="Figures 2022. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 20 Sandracottus jaechi 21 S. maculatus and 22 S. rotundus." figureDoi="10.3897/zookeys.1223.138220.figures20-22" httpUri="https://binary.pensoft.net/fig/1214413">21 a</figureCitation>
). Parameres narrow and elongate, and slightly longer than median lobe (Fig.
<figureCitation id="CE4E350C197540F7BDDAC90DC1D679CF" captionStart="Figures 2022" captionStartId="F6" captionText="Figures 2022. Median lobe of aedeagus in ventral view (a), and right paramere in lateral view (b): 20 Sandracottus jaechi 21 S. maculatus and 22 S. rotundus." figureDoi="10.3897/zookeys.1223.138220.figures20-22" httpUri="https://binary.pensoft.net/fig/1214413">21 b</figureCitation>
).
</paragraph>
<paragraph id="7CDC4BC6C397A6BD1EFBC89CCB9694E0">
<specimenCount id="622F62F248A00D3CA84B322BD79D4665" count="1" type="female"></specimenCount>
. Similar to male. Pro- and mesotarsi not enlarged. Microsculpture on ventrite VI as in male.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKDDK" type="Differential diagnosis">
<paragraph id="1A8A72ACB01491DE0347063E5D56878D">
<heading id="B3006EBE17378F0F27D071696B13220E" reason="title">Differential diagnosis.</heading>
</paragraph>
<paragraph id="DD0A4AB38EF627AE0CBD656FB65539DD">
The species is clearly distinguished from all other Indomalayan species of the genus by its colouration and larger size. It is the largest species of the genus. From the dorsal colouration
<taxonomicName id="7A835589703D5EA26A4A060D3F7870FD" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="F37EA1E535A2E7B37BE1C06F80ED0D5E" italics="true">S. maculatus</emphasis>
</taxonomicName>
is near to
<taxonomicName id="FD6D48F662ABA47A4EDECBC92FE6D716" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="AB783FA2997A4CCD1721E817A7F6B763" italics="true">S. femoralis</emphasis>
</taxonomicName>
which is generally smaller in size (see
<taxonomicName id="4DC79CE4E22CF4EC75DA93E4E686B6EA" authorityName="Heller" authorityYear="1934" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="3855864AB2F189E3331281D772278AFB" italics="true">S. femoralis</emphasis>
</taxonomicName>
). Furthermore, both species can be separated by the shapes of their median lobes.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EREDK" type="distribution">
<paragraph id="0D0764700BC0CA13D7E02F2749A92CB3">
<heading id="527D99CC1C0F9CF01362BB99249CD7C4" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="70BBA98EB0FCDE93F41040CB7A5A5A01">
<collectingCountry id="F2FFE49ED235DA5BB2E440661454C8FC" name="Cambodia">Cambodia</collectingCountry>
,
<collectingCountry id="C2CB1273C53DEE542B7A2F020827492F" name="Laos">Laos</collectingCountry>
,
<collectingCountry id="25B2FFF98F717A2FC6FEC6053A20889A" name="Thailand">Thailand</collectingCountry>
,
<collectingCountry id="CDEA8AE521F15CA6AAC9787B1D81DA01" name="Vietnam">Vietnam</collectingCountry>
,
<collectingCountry id="1C907371EF75A66A0F2F785325ECE291" name="Malaysia">Malaysia</collectingCountry>
(
<collectingRegion id="E8975EE1AEBE241AA5BFBB8FB8F2DC7D" country="Malaysia" name="Sabah">Sabah</collectingRegion>
),
<collectingCountry id="0A14F49EFB6AB7769F9CE1EB281123AE" name="Indonesia">Indonesia</collectingCountry>
(
<collectingRegion id="AD3C72FEF0CB2BB8353B4669D446A892" country="Indonesia" name="Kalimantan Utara">Kalimantan</collectingRegion>
,
<collectingRegion id="48B7F26175D9D112F53DB4548F9D6E3C" country="Indonesia" name="Sumatera Utara">Sumatra</collectingRegion>
,
<collectingRegion id="A82981CD5B68F5FA81CE9088EA429440" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
, Siberut) and
<collectingCountry id="2D98ADCB25A75968DA81275EEC094ABA" name="Philippines">Philippines</collectingCountry>
(Luzon, Mindanao,
<collectingRegion id="9E041365AD2059E17D7C2525C69F31FE" country="Philippines" name="Palawan">Palawan</collectingRegion>
) (Fig.
<figureCitation id="B012477ECD3C6C413D11B52CEEB9D62E" captionStart="Figure 24" captionStartId="F8" captionText="Figure 24. Distribution of Sandracottus bizonatus [red squares] and S. maculatus [black dots] in Southeast Asia." figureDoi="10.3897/zookeys.1223.138220.figure24" httpUri="https://binary.pensoft.net/fig/1214415">24</figureCitation>
). Specimens were collected from approximately sea level to
<quantity id="48B1B5A0FABDAC1FF77096C4E5227C5A" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</quantity>
a. s. l.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2EDK" type="habitat">
<paragraph id="DBEA9B2ED618BA6644B541206BC7A455">
<heading id="5288AB2951F20606F65F0B082CFABDF8" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="5C13F00A7372FADDBEDAEE157C139B28">
A widespread but rare species. According to various collectors all recent records are from well vegetated pools, forest pools, or rest pools of intermittent streams and rivers, mainly located in primary rainforest (Fig.
<figureCitation id="1AFEF27DA1F5775674D8B7C9FF958F1D" captionStart="Figures 4345" captionStartId="F17" captionText="Figures 4345. Habitats of Sandracottus 43, 44 Malaysia, Sabah, Kinabatangan riv. 8. - 15. VI. 2003, Uncle Tan´s camp, habitat of S. maculatus (photos: J. Šťastný) 45 Indonesia, Sulawesi, Togian Islands, Kaldidiri Island near Paradise Island Resort, Babirusa puddles (on the left), habitat of S. rotundus (photo: J. Haft)." figureDoi="10.3897/zookeys.1223.138220.figures43-45" httpUri="https://binary.pensoft.net/fig/1214424">43</figureCitation>
). At least most habitats are partly shaded and enriched with decaying leaves and twigs (Fig.
<figureCitation id="3F67E81AD19F57670E0C2A83DD6E766A" captionStart="Figures 4345" captionStartId="F17" captionText="Figures 4345. Habitats of Sandracottus 43, 44 Malaysia, Sabah, Kinabatangan riv. 8. - 15. VI. 2003, Uncle Tan´s camp, habitat of S. maculatus (photos: J. Šťastný) 45 Indonesia, Sulawesi, Togian Islands, Kaldidiri Island near Paradise Island Resort, Babirusa puddles (on the left), habitat of S. rotundus (photo: J. Haft)." figureDoi="10.3897/zookeys.1223.138220.figures43-45" httpUri="https://binary.pensoft.net/fig/1214424">44</figureCitation>
). In
<collectingCountry id="AC36619F8BF17641E9EDBA995ADE49C8" name="Thailand">Thailand</collectingCountry>
(Sakaerat Biosphere Reserve) the species was collected at the end of the dry season in a rest pond of a rocky stream, with very dark water and accumulated leaf litter. The pond was more or less permanent but with a high fluctuation of water level (Fig.
<figureCitation id="8AC8C8E0150E8F0FEB24E3E7C3936308" captionStart="Figures 3742" captionStartId="F16" captionText="Figures 3742. Habitats of Sandracottus 37, 38 Laos, Attapeu prov., Annam Highlands Mts., Dong Amphan NBCA, Nong Fa [puddles near lake shore] and lake, habitat of S. hunteri 39 Laos, Katamtok, rest pools along a rocky river bed, habitat of S. maculatus (photos: J. Hájek) 40 Thailand, near Salawin NP, open and muddy puddle made by elephants (photo: J. Šťastný), and 41 Sakaerat Biosphere Reserve, rocky pool in an otherwise dry stream bed, habitats of S. hunteri and S. maculatus (photo: W. Atthakor) 42 Philippines, Luzon, Pampanga Prov., Bano San Juan, Mt. Arayat NP, spring creek, habitat of S. insignis (photo: H. Freitag)." figureDoi="10.3897/zookeys.1223.138220.figures37-42" httpUri="https://binary.pensoft.net/fig/1214423">41</figureCitation>
). The species was syntopic with
<taxonomicName id="D315467FAA3679287BD112E2DD23B43C" authority="(Atthakor et al. 2018)" baseAuthorityName="Atthakor" baseAuthorityYear="2018" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="hunteri">
<emphasis id="29C522296032D39B85E7AFAA31604F07" italics="true">S. hunteri</emphasis>
(
<bibRefCitation id="E38B66DE4349FD1135F4D5B66B492741" author="Atthakor" etAl="et al." firstAuthor="Atthakor" issue="1" journalOrPublisher="Spixiana" pagination="91-98" refId="B3" refString="Atthakor W, Hendrich L, Sangpradub N, Balke M (2018) Diving beetles of the Sakaerat Biosphere Reserve, Nakhon Ratchasima Province, with four new records for Thailand (Coleoptera, Dytiscidae). Spixiana 41 (1): 9198." title="Diving beetles of the Sakaerat Biosphere Reserve, Nakhon Ratchasima Province, with four new records for Thailand (Coleoptera, Dytiscidae)." volume="41" year="2018">Atthakor et al. 2018</bibRefCitation>
)
</taxonomicName>
. In
<collectingCountry id="6D243ED535636D5FE1DABCC3E154E9E0" name="Vietnam">Vietnam</collectingCountry>
, the species was collected in shallow water made by a passing jeep on a path. Syntopically occurring dytiscid species include
<taxonomicName id="71E7BCD33B54BEAE8F47698062D13261" authority="Balke, Wang, Bergsten &amp; Hendrich, 2017" authorityName="Balke, Wang, Bergsten &amp; Hendrich" authorityYear="2017" class="Insecta" family="Dytiscidae" genus="Leiodytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kualalipis">
<emphasis id="22D762ED6BEA67A3945AB52F1479B4F0" italics="true">Leiodytes kualalipis</emphasis>
<bibRefCitation id="D9E9932E1A882E791355459C6182EE0E" DOI="10.3897/zookeys.647.11231" author="Balke M &amp; Bergsten J &amp; Wang L-J &amp; Hendrich L" journalOrPublisher="ZooKeys" pagination="137-151" refId="B6" refString="Balke M, Bergsten J, Wang L-J, Hendrich L (2017) A new genus and two new species of Southeast Asian Bidessini as well as new synonyms for Oceanian species (Coleoptera, Dytiscidae). ZooKeys 647: 137 - 151. https: // doi. org / 10.3897 / zookeys. 647.11231" title="A new genus and two new species of Southeast Asian Bidessini as well as new synonyms for Oceanian species (Coleoptera, Dytiscidae)." volume="647" year="2017">Balke, Wang, Bergsten &amp; Hendrich, 2017</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="07222C72688BDCBC365333B5E1A6C4B1" authority="(Redtenbacher, 1867)" baseAuthorityName="Redtenbacher" baseAuthorityYear="1867" class="Insecta" family="Dytiscidae" genus="Leiodytes" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nicobaricus">
<emphasis id="82CABA529FE991C8E47DF9A310F76D39" italics="true">L. nicobaricus</emphasis>
(Redtenbacher, 1867)
</taxonomicName>
, and
<taxonomicName id="67250D2BA6B61BDB272217D41D860C7D" authority="(Clark, 1863) (Balke et al. 2017)" authorityName="Balke et al." authorityYear="2017" baseAuthorityName="Clark" baseAuthorityYear="1863" class="Insecta" family="Dytiscidae" genus="Hydroglyphus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="EB3B550731C38A8E898B3B3919014554" italics="true">Hydroglyphus orientalis</emphasis>
(Clark, 1863) (
<bibRefCitation id="925DDE685BB02E67FFFC4D8D08FF70A8" DOI="10.3897/zookeys.647.11231" author="Balke" etAl="et al." firstAuthor="Balke" journalOrPublisher="ZooKeys" pagination="137-151" refId="B6" refString="Balke M, Bergsten J, Wang L-J, Hendrich L (2017) A new genus and two new species of Southeast Asian Bidessini as well as new synonyms for Oceanian species (Coleoptera, Dytiscidae). ZooKeys 647: 137151. https://doi.org/10.3897/zookeys.647.11231" title="A new genus and two new species of Southeast Asian Bidessini as well as new synonyms for Oceanian species (Coleoptera, Dytiscidae)." volume="647" year="2017">Balke et al. 2017</bibRefCitation>
)
</taxonomicName>
. In
<collectingCountry id="D82980A3B77ADDFB43B9C5A29213D305" name="Laos">Laos</collectingCountry>
,
<taxonomicName id="83A63C871FB0F02D82C2FE93141A8587" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="21FE6FC730A38E98454C69662E514D01" italics="true">S. maculatus</emphasis>
</taxonomicName>
was collected in a residual pool near the river (Fig.
<figureCitation id="3F6F9C6B3B7B29B4055B4C67A0D5514B" captionStart="Figures 3742" captionStartId="F16" captionText="Figures 3742. Habitats of Sandracottus 37, 38 Laos, Attapeu prov., Annam Highlands Mts., Dong Amphan NBCA, Nong Fa [puddles near lake shore] and lake, habitat of S. hunteri 39 Laos, Katamtok, rest pools along a rocky river bed, habitat of S. maculatus (photos: J. Hájek) 40 Thailand, near Salawin NP, open and muddy puddle made by elephants (photo: J. Šťastný), and 41 Sakaerat Biosphere Reserve, rocky pool in an otherwise dry stream bed, habitats of S. hunteri and S. maculatus (photo: W. Atthakor) 42 Philippines, Luzon, Pampanga Prov., Bano San Juan, Mt. Arayat NP, spring creek, habitat of S. insignis (photo: H. Freitag)." figureDoi="10.3897/zookeys.1223.138220.figures37-42" httpUri="https://binary.pensoft.net/fig/1214423">39</figureCitation>
). The pool had a diameter of nearly
<quantity id="802300DA3272A9AAF677C8A7CAEFDC86" metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">1 m</quantity>
, depth was ~
<quantity id="DD59B5958FD1FECC36E8728F7D6C0A46" metricMagnitude="-1" metricUnit="m" metricValue="4.0" unit="cm" value="40.0">40 cm</quantity>
, bottom with a thick layer of decaying leaves (Hájek pers. comm. 2023).
</paragraph>
<caption id="E8B74CAE91BA0744D9E255A36601580B" ID-DOI="10.3897/zookeys.1223.138220.figures43-45" ID-arpha="A50EB81C-D38E-56F6-8FDE-AEFC0360BFD1" httpUri="https://binary.pensoft.net/fig/1214424" startId="F17">
<paragraph id="BBD2DA90E849395493E6F6229F2C708D">
<label id="8ED2145257E46ADCD4816C2AEE919C7F">Figures 4345.</label>
</paragraph>
<paragraph id="3F093FDE12591738CC1C59862B498841">
Habitats of
<taxonomicName id="34D604A5CB992E75A2C0A47566FBF611" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="095FDF22C93C1CADE087D9AAB804F7C7" italics="true">Sandracottus</emphasis>
</taxonomicName>
<emphasis id="262DC1B01235D03CD51BFB5C46A1CA1A" bold="true">43, 44</emphasis>
Malaysia, Sabah, Kinabatangan riv. 8. - 15. VI. 2003, Uncle Tan´s camp, habitat of
<taxonomicName id="2049B6F8D1F4E6F0528ADB0C62133816" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="46528CFA3FBEE8DCC68DBD177FB7FF00" italics="true">S. maculatus</emphasis>
</taxonomicName>
(photos: J. Šťastný)
<emphasis id="4E79ADC12BE00733EB6D06A1BB88DF65" bold="true">45</emphasis>
Indonesia, Sulawesi, Togian Islands, Kaldidiri Island near Paradise Island Resort, Babirusa puddles (on the left), habitat of
<taxonomicName id="019D52A1833D05E6F082E2F087A513E7" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Sandracottus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="rotundus">
<emphasis id="E5EA36ABB7FDE4EB183E220B7583842F" italics="true">S. rotundus</emphasis>
</taxonomicName>
(photo: J. Haft).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>