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<document id="DA57F45BB715B6AF7E8EFD7136FA924F" ID-CLB-Dataset="39303" ID-DOI="10.11646/zootaxa.4057.1.9" ID-GBIF-Dataset="a3d9451b-2e0c-40e6-bafb-2d8a958fd8de" ID-ISSN="1175-5326" ID-Zenodo-Dep="233231" ID-ZooBank="7AC91C1D-5EAB-40A6-B177-3A154AF2F9EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461354776227" checkinUser="plazi" docAuthor="Jacobs, Carmen T., Scholtz, Clarke H. &amp; Strümpher, Werner P." docDate="2015" docId="B01287C7FFB68815FF283D2DCC43E01F" docLanguage="en" docName="zt04057p142.pdf" docOrigin="Zootaxa 4057 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colophon deschodti Jacobs &amp; Scholtz, new species" docType="treatment" docVersion="9" lastPageNumber="141" masterDocId="4C2BFFBFFFB38813FFBF397ACD4FE223" masterDocTitle="Taxonomy of Colophon Gray (Coleoptera: Lucanidae): new species and a status change" masterLastPageNumber="142" masterPageNumber="135" pageNumber="140" updateTime="1698661467203" updateUser="ExternalLinkService">
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<mods:title id="6951D48EFC88B40181B719B764D69A3E">Taxonomy of Colophon Gray (Coleoptera: Lucanidae): new species and a status change</mods:title>
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<mods:namePart id="50D3F8B246B6222C6E48F62DD6E5C057">Jacobs, Carmen T.</mods:namePart>
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<mods:namePart id="E2C1690D7EEEAA673CE524E54581DA09">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="BE6067A7AB2587A5F43C483C200E2394">Strümpher, Werner P.</mods:namePart>
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<mods:date id="1F47487F78E0EA9663521D14B335D2A2">2015</mods:date>
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<treatment id="B01287C7FFB68815FF283D2DCC43E01F" ID-DOI="http://doi.org/10.5281/zenodo.6093239" ID-GBIF-Taxon="119647049" ID-Zenodo-Dep="6093239" LSID="urn:lsid:plazi:treatment:B01287C7FFB68815FF283D2DCC43E01F" httpUri="http://treatment.plazi.org/id/B01287C7FFB68815FF283D2DCC43E01F" lastPageId="6" lastPageNumber="141" pageId="5" pageNumber="140">
<subSubSection id="70A1655AFFB68816FF283D2DCC23E6B0" pageId="5" pageNumber="140" type="nomenclature">
<paragraph id="380436D1FFB68816FF283D2DCE4CE652" blockId="5.[151,771,1111,1171]" box="[151,771,1111,1137]" pageId="5" pageNumber="140">
<heading id="634C81BDFFB68816FF283D2DCE4CE652" bold="true" box="[151,771,1111,1137]" fontSize="11" level="1" pageId="5" pageNumber="140" reason="1">
<emphasis id="0ACFEAC3FFB68816FF283D2DCE4CE652" bold="true" box="[151,771,1111,1137]" pageId="5" pageNumber="140">
<taxonomicName id="FFBB4D52FFB68816FF283D2DCF28E652" ID-CoL="X83R" authority="Jacobs &amp; Scholtz" authorityName="Jacobs &amp; Scholtz" box="[151,615,1111,1137]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="deschodti" status="sp. nov.">
<emphasis id="0ACFEAC3FFB68816FF283D2DCCC9E652" bold="true" box="[151,390,1111,1137]" italics="true" pageId="5" pageNumber="140">Colophon deschodti</emphasis>
Jacobs &amp; Scholtz
</taxonomicName>
,
<taxonomicNameLabel id="11FC57B8FFB68816FDCC3D2DCE4CE652" box="[627,771,1111,1137]" pageId="5" pageNumber="140" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="380436D1FFB68816FF283D00CC23E6B0" blockId="5.[151,771,1111,1171]" box="[151,364,1146,1171]" pageId="5" pageNumber="140">
(
<figureCitation id="A0802A54FFB68816FF203D00CD94E6B0" box="[159,219,1146,1171]" captionStart="FIGURE 4" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[232,1352,1303,1918]" captionTargetId="figure@5.[226,1360,1298,1932]" captionTargetPageId="5" captionText="FIGURE 4. Adult habitus of Colophon deschodti new species: (a) male dorsal view; (b) male ventral view; (c) male protibia." httpUri="https://zenodo.org/record/233235/files/figure.png" pageId="5" pageNumber="140">Fig 4</figureCitation>
ac;
<tableCitation id="7539036AFFB68816FEAF3D00CC2EE6B0" box="[272,353,1146,1171]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="5" pageNumber="140">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB68816FF283DB8CF02E6F8" box="[151,589,1218,1243]" pageId="5" pageNumber="140" type="materials_examined">
<paragraph id="380436D1FFB68816FF283DB8CF02E6F8" blockId="5.[151,589,1218,1243]" box="[151,589,1218,1243]" pageId="5" pageNumber="140">
<emphasis id="0ACFEAC3FFB68816FF283DB8CC72E6F8" bold="true" box="[151,317,1218,1243]" pageId="5" pageNumber="140">
<typeStatus id="E7008873FFB68816FF283DB8CD9DE6F8" box="[151,210,1218,1243]" pageId="5" pageNumber="140">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="E7008873FFB68816FEF33DB8CCFBE6F8" box="[332,436,1218,1243]" pageId="5" pageNumber="140" type="holotype">Holotype</typeStatus>
(
<specimenCount id="2EBDFD58FFB68816FE7A3DB9CCA4E6F8" box="[453,491,1219,1243]" pageId="5" pageNumber="140" type="generic">1♂</specimenCount>
:
<collectionCode id="5EAAAE14FFB68816FE493DB9CF0EE6F8" box="[502,577,1219,1243]" pageId="5" pageNumber="140">ISAM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="6CC46659FFB68816FF283EDBC8C0E594" box="[151,1423,1953,1975]" httpUri="https://zenodo.org/record/233235/files/figure.png" pageId="5" pageNumber="140" targetBox="[232,1352,1303,1918]" targetPageId="5">
<paragraph id="380436D1FFB68816FF283EDBC8C0E594" blockId="5.[151,1423,1953,1975]" box="[151,1423,1953,1975]" pageId="5" pageNumber="140">
<emphasis id="0ACFEAC3FFB68816FF283EDBCC5DE595" bold="true" box="[151,274,1953,1975]" pageId="5" pageNumber="140">FIGURE 4.</emphasis>
Adult habitus of
<taxonomicName id="FFBB4D52FFB68816FE7B3EDBCFC1E594" box="[452,654,1953,1975]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="deschodti" status="sp. nov.">
<emphasis id="0ACFEAC3FFB68816FE7B3EDBCFC1E594" box="[452,654,1953,1975]" italics="true" pageId="5" pageNumber="140">Colophon deschodti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11FC57B8FFB68816FD2A3ED8CE5DE594" box="[661,786,1953,1975]" pageId="5" pageNumber="140" rank="species">new species</taxonomicNameLabel>
: (a) male dorsal view; (b) male ventral view; (c) male protibia.
</paragraph>
</caption>
<subSubSection id="70A1655AFFB58815FF7839EDCE00E2DB" pageId="6" pageNumber="141" type="diagnosis">
<paragraph id="380436D1FFB58815FF7839EDCE00E2DB" blockId="6.[151,1437,151,572]" pageId="6" pageNumber="141">
<emphasis id="0ACFEAC3FFB58815FF7839EDCC75E293" bold="true" box="[199,314,151,176]" pageId="6" pageNumber="141">Diagnosis</emphasis>
.
<taxonomicName id="FFBB4D52FFB58815FEF739E2CF67E293" box="[328,552,152,176]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="deschodti">
<emphasis id="0ACFEAC3FFB58815FEF739E2CF67E293" box="[328,552,152,176]" italics="true" pageId="6" pageNumber="141">Colophon deschodti</emphasis>
</taxonomicName>
is distinguished from other members in the group by the shape of the mentum, which is clearly divided into two projecting tubercles or ridges.
<taxonomicName id="FFBB4D52FFB58815FC3E39C6C92BE2F7" box="[897,1124,188,212]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="deschodti">
<emphasis id="0ACFEAC3FFB58815FC3E39C6C92BE2F7" box="[897,1124,188,212]" italics="true" pageId="6" pageNumber="141">Colophon deschodti</emphasis>
</taxonomicName>
lacks any tubercles on the head, but bears a median point on anterior margin of the head.
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB58815FF78387ECE66E3F3" pageId="6" pageNumber="141" type="materials_examined">
<paragraph id="380436D1FFB58815FF78387ECE66E3F3" blockId="6.[151,1437,151,572]" pageId="6" pageNumber="141">
<emphasis id="0ACFEAC3FFB58815FF78387ECC8EE33E" bold="true" box="[199,449,260,285]" pageId="6" pageNumber="141">
<typeStatus id="E7008873FFB58815FF78387ECC7BE33E" box="[199,308,260,285]" pageId="6" pageNumber="141" type="holotype">Holotype</typeStatus>
description
</emphasis>
.
<emphasis id="0ACFEAC3FFB58815FE6E387ECF6FE33F" box="[465,544,260,284]" italics="true" pageId="6" pageNumber="141">Colour</emphasis>
: Black.
<emphasis id="0ACFEAC3FFB58815FD3D387ECFE0E338" box="[642,687,260,283]" italics="true" pageId="6" pageNumber="141">Size</emphasis>
: length
<quantity id="FF439B34FFB58815FCB2387ECE3AE33F" box="[781,885,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="6" pageNumber="141" unit="mm" value="20.5">20.5 mm</quantity>
, pronotal width
<quantity id="FF439B34FFB58815FB8F387EC9D8E33F" box="[1072,1175,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="6" pageNumber="141" unit="mm" value="10.4">10.4 mm</quantity>
, mandible length:
<quantity id="FF439B34FFB58815FAD4387ECD89E31C" metricMagnitude="-3" metricUnit="m" metricValue="3.28" pageId="6" pageNumber="141" unit="mm" value="3.28">3.28 mm</quantity>
.
<emphasis id="0ACFEAC3FFB58815FF6E3852CC42E363" box="[209,269,296,320]" italics="true" pageId="6" pageNumber="141">Head</emphasis>
: With a distinct anterior margin, which is concave with a median point and a strong oblique ridge over the eye, no tubercles present (
<tableCitation id="7539036AFFB58815FE533836CF0DE347" box="[492,578,332,357]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="6" pageNumber="141">Table 1</tableCitation>
).
<emphasis id="0ACFEAC3FFB58815FDE33836CF88E347" box="[604,711,332,356]" italics="true" pageId="6" pageNumber="141">Mandible</emphasis>
: Apex of mandible deeply impressed on upper surface as in
<taxonomicName id="FFBB4D52FFB58815FA3D3836CDA9E3AB" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="stokoei">
<emphasis id="0ACFEAC3FFB58815FA3D3836CDA9E3AB" italics="true" pageId="6" pageNumber="141">C. stokoei</emphasis>
</taxonomicName>
, but with the tooth on the inner basal margin rounded and nearly obsolete (
<tableCitation id="7539036AFFB58815FBE23815C9FBE3AB" box="[1117,1204,367,392]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="6" pageNumber="141">Table 1</tableCitation>
).
<emphasis id="0ACFEAC3FFB58815FB70380BC864E3AB" box="[1231,1323,369,392]" italics="true" pageId="6" pageNumber="141">Mentum</emphasis>
: Anterior surface with two projecting ridges, causing it to appear distinctly bifid in front ventral view (
<tableCitation id="7539036AFFB58815FB1138EEC84DE38F" box="[1198,1282,404,429]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="6" pageNumber="141">Table 1</tableCitation>
).
<emphasis id="0ACFEAC3FFB58815FAA438EFC81FE38F" box="[1307,1360,405,428]" italics="true" pageId="6" pageNumber="141">Legs</emphasis>
: As in
<taxonomicName id="FFBB4D52FFB58815FF2838C2CC71E3F3" box="[151,318,440,464]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="struempheri">
<emphasis id="0ACFEAC3FFB58815FF2838C2CC71E3F3" box="[151,318,440,464]" italics="true" pageId="6" pageNumber="141">C. struempheri</emphasis>
</taxonomicName>
(
<figureCitation id="A0802A54FFB58815FEF138CDCCDCE3F3" box="[334,403,439,464]" captionStart="FIGURE 4" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[232,1352,1303,1918]" captionTargetId="figure@5.[226,1360,1298,1932]" captionTargetPageId="5" captionText="FIGURE 4. Adult habitus of Colophon deschodti new species: (a) male dorsal view; (b) male ventral view; (c) male protibia." httpUri="https://zenodo.org/record/233235/files/figure.png" pageId="6" pageNumber="141">Fig. 4</figureCitation>
c).
<emphasis id="0ACFEAC3FFB58815FE0438C2CF10E3F3" box="[443,607,440,464]" italics="true" pageId="6" pageNumber="141">Male genitalia</emphasis>
: As in
<emphasis id="0ACFEAC3FFB58815FD0E38C2CE66E3F3" box="[689,809,440,464]" italics="true" pageId="6" pageNumber="141">
<taxonomicName id="FFBB4D52FFB58815FD0E38C2CE6AE3F3" box="[689,805,440,464]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="stokoei">C. stokoei</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB58815FF7838A6CE97E3D6" box="[199,984,476,501]" pageId="6" pageNumber="141" type="distribution">
<paragraph id="380436D1FFB58815FF7838A6CE97E3D6" blockId="6.[151,1437,151,572]" box="[199,984,476,501]" pageId="6" pageNumber="141">
<emphasis id="0ACFEAC3FFB58815FF7838A6CC16E3D6" bold="true" box="[199,345,476,501]" pageId="6" pageNumber="141">Distribution</emphasis>
. Hottentots
<collectingCountry id="40AC7641FFB58815FE5C38A6CF72E3D6" box="[483,573,476,501]" name="Netherlands" pageId="6" pageNumber="141">Holland</collectingCountry>
Mountains, Western Cape Province.
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB58815FF783885CC43E01F" pageId="6" pageNumber="141" type="etymology">
<paragraph id="380436D1FFB58815FF783885CC43E01F" blockId="6.[151,1437,151,572]" pageId="6" pageNumber="141">
<emphasis id="0ACFEAC3FFB58815FF783885CC09E03B" bold="true" box="[199,326,511,536]" pageId="6" pageNumber="141">Etymology</emphasis>
. This species is named after Christian Deschodt in recognition of his contribution to the study of
<taxonomicName id="FFBB4D52FFB58815FF283B5ECC4AE01F" box="[151,261,548,572]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="0ACFEAC3FFB58815FF283B5ECC4AE01F" box="[151,261,548,572]" italics="true" pageId="6" pageNumber="141">Colophon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>