168 lines
18 KiB
XML
168 lines
18 KiB
XML
<document id="94FEEB844E4760CB61EA507BAF457063" ID-DOI="10.11646/zootaxa.5028.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5453967" ID-ZooBank="C678F0FE-1B62-4F34-8A66-449CF9806B50" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630655426570" checkinUser="plazi" docAuthor="Santos, Bruna S., Marques, Mariana P., Bauer, Aaron M. & Ceríaco, Luis M. P." docDate="2021" docId="B007F528FFF9FFF416D7FA4FFBB45E81" docLanguage="en" docName="zootaxa.5028.1.1.pdf" docOrigin="Zootaxa 5028 (1)" docSource="http://zoobank.org/c678f0fe-1b62-4f34-8a66-449cf9806b50" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Philothamnus angolensis Bocage 1882" docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="4C3E8D50FFD8FFD51640FFA1FFD2597B" masterDocTitle="Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903 – 1906): a taxonomic revision and new records of a forgotten collection" masterLastPageNumber="80" masterPageNumber="1" pageNumber="34" updateTime="1699281765633" updateUser="plazi">
|
||
<mods:mods id="67E3029D848F7756F9C561BA765374F4" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="47FE40ACAB0127A986E81F3665E2D2E9">
|
||
<mods:title id="96C44526EB0D9F8584088A83653C79A8">Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903 – 1906): a taxonomic revision and new records of a forgotten collection</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="3A6FECBD6F1B71215BB8F64B1F8FA58F" type="personal">
|
||
<mods:role id="D145B6335737F378892EA4E5198E20FB">
|
||
<mods:roleTerm id="E15D84C905B59B80B88E8C205E4BD2DC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="839704D4553A75794FBAF2729BD5CAAE">Santos, Bruna S.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="5ED930512A1CE0B142AB271F2CAE68DD" type="personal">
|
||
<mods:role id="A6CD929C7915D37198D08BF4773A59D4">
|
||
<mods:roleTerm id="37A0F39B4DA537FC930084A84097D94D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D68303FD92B366C0F3ECA50E0C991AAD">Marques, Mariana P.</mods:namePart>
|
||
<mods:nameIdentifier id="14F87D5F60415D348733A49A68DDC264" type="ORCID">0000-0002-1712-2632</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="5320EA7D59421401A57632ED0D1413B7" type="email">mptlmarques@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="E39FF4C134E3425B1E0AC66C0F0ED561" type="personal">
|
||
<mods:role id="61C0A5C3E6DCEFCBA13EE30D052BF6EF">
|
||
<mods:roleTerm id="E0EF84AAFA9C70B7803524E2D624AFC9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BAC4D86F9D61CD6559DB8F14215E2847">Bauer, Aaron M.</mods:namePart>
|
||
<mods:nameIdentifier id="671AA16C83861E5F167E3D4B9F45BC50" type="ORCID">0000-0001-6839-8025</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="F8030DCD4DA91C6365F19ABD954A8E50" type="email">aaron.bauer@villanova.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="898C45BA526CFD3CCE64AF5D7D8C7088" type="personal">
|
||
<mods:role id="339F6D59B9ECBA483C51E8DF4E512219">
|
||
<mods:roleTerm id="D27AA3FCA83A0878EE0F029E61C97C30">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4884991BCC729D0E880C547EDAC37342">Ceríaco, Luis M. P.</mods:namePart>
|
||
<mods:nameIdentifier id="B28608EF0A30D56A18AEDE668753BF44" type="ORCID">0000-0002-3798-2664</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="657D13CF1B8B7BAAD55CDAF6AE8D953F" type="email">brunascesantos@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="7394EC9CF9D62CEE64388174EAD51BBD">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E400CBE793C4D97F987CE43A2E682AD9" type="host">
|
||
<mods:titleInfo id="BFC06EA0D7B9F236A193860F541E7807">
|
||
<mods:title id="938CA865616419CBCF5A56272A8E84D3">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="0D6DBAD329E99C0B381B3FCBBC018726">
|
||
<mods:date id="EABA347A72317FFFBF375FE4B1F908D5">2021</mods:date>
|
||
<mods:detail id="818D1332BF3BC58A35903FD217282B94" type="pubDate">
|
||
<mods:number id="CEE5C70E286E42168F222A63671A557E">2021-09-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="18F87C2B684F9B477366CCCCCEF989D4" type="volume">
|
||
<mods:number id="E37FC308182711D4C05EDAECBF20D8AE">5028</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="29F348343F5E29F3F27B5B5271761FB0" type="issue">
|
||
<mods:number id="9207CD76E7BEC206C5A794BBD4B8CD06">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="8ED9FFBEB7C4D36ECE30540161287537" unit="page">
|
||
<mods:start id="3BA31688ACBCECAC3A5E66BDB75BD399">1</mods:start>
|
||
<mods:end id="0AE76276EE566F4A67E7B4128E093C68">80</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="455F12585DE2BF197A985254601C9ED4">
|
||
<mods:url id="BA2B7EDBF374DBCCABE155938071F48F">http://zoobank.org/c678f0fe-1b62-4f34-8a66-449cf9806b50</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="F00A3E91338551099D13D836B4F56397">journal article</mods:classification>
|
||
<mods:identifier id="9D961D2807EA045A9087EC2D3F10665D" type="DOI">10.11646/zootaxa.5028.1.1</mods:identifier>
|
||
<mods:identifier id="2CC5D1DE0F2B2BAF1007B863439F0582" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="943BFA97058CA25A31100381D8DD8DB2" type="Zenodo-Dep">5453967</mods:identifier>
|
||
<mods:identifier id="DF2472344366E5AA59E8D2249CEF337B" type="ZooBank">C678F0FE-1B62-4F34-8A66-449CF9806B50</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="B007F528FFF9FFF416D7FA4FFBB45E81" ID-DOI="http://doi.org/10.5281/zenodo.5495381" ID-Zenodo-Dep="5495381" LSID="urn:lsid:plazi:treatment:B007F528FFF9FFF416D7FA4FFBB45E81" httpUri="http://treatment.plazi.org/id/B007F528FFF9FFF416D7FA4FFBB45E81" lastPageNumber="34" pageId="33" pageNumber="34">
|
||
<subSubSection id="70B417B5FFF9FFF416D7FA4FFDA75F73" box="[151,629,1518,1545]" pageId="33" pageNumber="34" type="nomenclature">
|
||
<paragraph id="3811443EFFF9FFF416D7FA4FFDA75F73" blockId="33.[151,629,1518,1545]" box="[151,629,1518,1545]" pageId="33" pageNumber="34">
|
||
<heading id="6359F352FFF9FFF416D7FA4FFDA75F73" bold="true" box="[151,629,1518,1545]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="4">
|
||
<taxonomicName id="FFAE3FBDFFF9FFF416D7FA4FFDA75F73" authority="Bocage, 1882" authorityName="Bocage" authorityYear="1882" box="[151,629,1518,1545]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="0ADA982CFFF9FFF416D7FA4FFDA75F73" bold="true" box="[151,629,1518,1545]" pageId="33" pageNumber="34">
|
||
<emphasis id="0ADA982CFFF9FFF416D7FA4FFE195F72" bold="true" box="[151,459,1518,1545]" italics="true" pageId="33" pageNumber="34">Philothamnus angolensis</emphasis>
|
||
Bocage, 1882
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70B417B5FFF9FFF416D7F991FD645F15" pageId="33" pageNumber="34" type="materials_examined">
|
||
<paragraph id="3811443EFFF9FFF416D7F991FD645F15" blockId="33.[151,1437,1584,2043]" pageId="33" pageNumber="34">
|
||
<emphasis id="0ADA982CFFF9FFF416D7F991FE175F31" bold="true" box="[151,453,1584,1610]" pageId="33" pageNumber="34">
|
||
Material (
|
||
<specimenCount id="2EA88FB7FFF9FFF41750F991FE655F31" box="[272,439,1584,1610]" pageId="33" pageNumber="34" type="generic">one specimen</specimenCount>
|
||
):
|
||
</emphasis>
|
||
One adult specimen (MHNCUP/REP 182, formerly UP-MHNFCP-017392;
|
||
<figureCitation id="A09558BBFFF9FFF4136CF991FA5D5F31" box="[1324,1423,1584,1610]" captionStart="FIGURE S" captionStartId="78.[151,248,2006,2030]" captionTargetBox="[170,1416,1100,1980]" captionTargetId="figure-51@78.[170,1416,1100,1980]" captionTargetPageId="78" captionText="FIGURE S74. Specimen of Philotamnus angolensis (MHNCUP/REP 182), from unknown locality. Photo by Bruna Santos." figureDoi="http://doi.org/10.5281/zenodo.5454131" httpUri="https://zenodo.org/record/5454131/files/figure.png" pageId="33" pageNumber="34">Fig. S74</figureCitation>
|
||
), collected from unknown locality, date unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="70B417B5FFF9FFF41687F9D9FBB45E81" pageId="33" pageNumber="34" type="discussion">
|
||
<paragraph id="3811443EFFF9FFF41687F9D9FBB45E81" blockId="33.[151,1437,1584,2043]" pageId="33" pageNumber="34">
|
||
<emphasis id="0ADA982CFFF9FFF41687F9D9FE805FE9" bold="true" box="[199,338,1656,1682]" pageId="33" pageNumber="34">Comments:</emphasis>
|
||
<taxonomicName id="FFAE3FBDFFF9FFF4171AF9D9FDA75FE9" authorityName="Bocage" authorityYear="1882" box="[346,629,1656,1682]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="0ADA982CFFF9FFF4171AF9D9FDA75FE9" box="[346,629,1656,1682]" italics="true" pageId="33" pageNumber="34">Philothamnus angolensis</emphasis>
|
||
</taxonomicName>
|
||
is a generalist species that inhabits both subtropical forest and savanna biomes and has a widespread distribution in sub-Saharan Africa, occurring in most of
|
||
<collectingCountry id="40B904AEFFF9FFF4122FF93DFB135FCD" box="[1135,1217,1692,1718]" name="Angola" pageId="33" pageNumber="34">Angola</collectingCountry>
|
||
with the exception of the southwestern desert (
|
||
<bibRefCitation id="5C3F39CFFFF9FFF41790F961FD6B5FA1" author="Marques, M. P. & Ceriaco, L. M. P. & Blackburn, D. C. & Bauer, A. M." box="[464,697,1728,1754]" pageId="33" pageNumber="34" pagination="1 - 501" refId="ref38562" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
|
||
Marques
|
||
<emphasis id="0ADA982CFFF9FFF4147CF960FDA45FA1" box="[572,630,1728,1754]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5C3F39CFFFF9FFF41486F961FC075FA1" author="Engelbrecht, H. M. & Branch, W. R. & Greenbaum, E. & Alexander, G. J. & Jackson, K. & Burger, M. & Conradie, W. & Kusamba, C. & Boulou, A-G. & Tolley, K. A." box="[710,981,1728,1754]" pageId="33" pageNumber="34" pagination="357 - 365" refId="ref36844" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., Zassi- Boulou, A-G. & Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
|
||
Engelbrecht
|
||
<emphasis id="0ADA982CFFF9FFF41516F960FC5D5FA1" box="[854,911,1728,1754]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="5C3F39CFFFF9FFF415A8F960FB0D5FA1" author="Marques, M. P. & Ceriaco, L. M. P. & Blackburn, D. C. & Bauer, A. M." box="[1000,1247,1728,1755]" pageId="33" pageNumber="34" pagination="1 - 501" refId="ref38562" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. & Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
|
||
Marques
|
||
<emphasis id="0ADA982CFFF9FFF41213F960FB5E5FA1" box="[1107,1164,1728,1754]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
(2018)
|
||
</bibRefCitation>
|
||
commented that Angolan records of
|
||
<taxonomicName id="FFAE3FBDFFF9FFF41738F945FCEA5F85" authority="(Leach, 1819)" baseAuthorityName="Leach" baseAuthorityYear="1819" box="[376,824,1764,1790]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="irregularis">
|
||
<emphasis id="0ADA982CFFF9FFF41738F945FD415F85" box="[376,659,1764,1790]" italics="true" pageId="33" pageNumber="34">Philothamnus irregularis</emphasis>
|
||
(Leach, 1819)
|
||
</taxonomicName>
|
||
correspond mainly to
|
||
<taxonomicName id="FFAE3FBDFFF9FFF41276F944FB185F85" authorityName="Bocage" authorityYear="1882" box="[1078,1226,1764,1790]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="0ADA982CFFF9FFF41276F944FB185F85" box="[1078,1226,1764,1790]" italics="true" pageId="33" pageNumber="34">P. angolensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FFAE3FBDFFF9FFF41341F945FE085E58" authority="(Gunther, 1863)" baseAuthorityName="Gunther" baseAuthorityYear="1863" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="hoplogaster">
|
||
<emphasis id="0ADA982CFFF9FFF41341F945FECF5E59" italics="true" pageId="33" pageNumber="34">Philothamnus hoplogaster</emphasis>
|
||
(Günther, 1863)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="5C3F39CFFFF9FFF417A6F8A9FD425E59" author="Ferreira, J. B." box="[486,656,1800,1826]" pageId="33" pageNumber="34" pagination="111 - 117" refId="ref37152" refString="Ferreira, J. B. (1904) Reptis e amphibios de Angola da regiao ao norte do Quanza (Colleccao Newton 1903). Jornal de Sciencias Mathematicas Physicas e Naturaes, Segunda Serie, 7 (25), 111 - 117." type="journal article" year="1904">Ferreira (1904)</bibRefCitation>
|
||
cited an adult specimen from “Chingo” as “
|
||
<taxonomicName id="FFAE3FBDFFF9FFF412C1F8A9FEEE5E3D" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="variety" species="irregularis" variety="angolensis">
|
||
<emphasis id="0ADA982CFFF9FFF412C1F8A9FA4E5E59" box="[1153,1436,1800,1826]" italics="true" pageId="33" pageNumber="34">Philothamnus irregularis</emphasis>
|
||
var
|
||
<emphasis id="0ADA982CFFF9FFF41684F88DFEEE5E3D" box="[196,316,1836,1862]" italics="true" pageId="33" pageNumber="34">angolensis</emphasis>
|
||
</taxonomicName>
|
||
” that exhibited an exceptional number of temporal scales, 2+2 (MHNCUP/REP 197, see account below) and a “blueish green” specimen of “
|
||
<taxonomicName id="FFAE3FBDFFF9FFF4143FF8F1FC4B5E11" baseAuthorityName="Leach" baseAuthorityYear="1819" box="[639,921,1872,1898]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="irregularis">
|
||
<emphasis id="0ADA982CFFF9FFF4143FF8F1FC4B5E11" box="[639,921,1872,1898]" italics="true" pageId="33" pageNumber="34">Philothamnus irregularis</emphasis>
|
||
</taxonomicName>
|
||
” from “Cambondo” with 1+1 temporal scales (
|
||
<bibRefCitation id="5C3F39CFFFF9FFF416DEF8D5FEEF5EF4" author="Ferreira, J. B." box="[158,317,1908,1935]" pageId="33" pageNumber="34" pagination="159 - 171" refId="ref37198" refString="Ferreira, J. B. (1906) Algumas especies novas ou pouco conhecidas de amphibios e reptis de Angola (Coleccao Newton - 1903 - 1904). Jornal de Sciencias Mathematicas Physicas e Naturaes, Segunda Serie, 7 (26), 159 - 171." type="journal article" year="1906">Ferreira 1906</bibRefCitation>
|
||
). An additional specimen, without locality, was also cited as “
|
||
<taxonomicName id="FFAE3FBDFFF9FFF415BBF8D5FAD15EF5" authorityName="Pfeffer" authorityYear="1893" box="[1019,1283,1908,1934]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="guntheri">
|
||
<emphasis id="0ADA982CFFF9FFF415BBF8D5FAD15EF5" box="[1019,1283,1908,1934]" italics="true" pageId="33" pageNumber="34">Philothamnus Guntheri</emphasis>
|
||
</taxonomicName>
|
||
” (a synonym of
|
||
<taxonomicName id="FFAE3FBDFFF9FFF416F6F838FE995EC9" authorityName="Bocage" authorityYear="1882" box="[182,331,1944,1970]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="0ADA982CFFF9FFF416F6F838FE995EC9" box="[182,331,1944,1970]" italics="true" pageId="33" pageNumber="34">P. angolensis</emphasis>
|
||
</taxonomicName>
|
||
) with 1+1 temporal scales. Upon analysis, we believe the extant specimen does not represent any of the published records, as it has 1+2 temporal scales.
|
||
<bibRefCitation id="5C3F39CFFFF9FFF41545F81DFBCF5EAD" author="Engelbrecht, H. M. & Branch, W. R. & Greenbaum, E. & Alexander, G. J. & Jackson, K. & Burger, M. & Conradie, W. & Kusamba, C. & Boulou, A-G. & Tolley, K. A." box="[773,1053,1980,2006]" pageId="33" pageNumber="34" pagination="357 - 365" refId="ref36844" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., Zassi- Boulou, A-G. & Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
|
||
Engelbrecht
|
||
<emphasis id="0ADA982CFFF9FFF415D3F81CFC1E5EAD" box="[915,972,1980,2006]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
recently reported
|
||
<taxonomicName id="FFAE3FBDFFF9FFF412AAF81CFAAC5EAD" authorityName="Bocage" authorityYear="1882" box="[1258,1406,1980,2006]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="0ADA982CFFF9FFF412AAF81CFAAC5EAD" box="[1258,1406,1980,2006]" italics="true" pageId="33" pageNumber="34">P. angolensis</emphasis>
|
||
</taxonomicName>
|
||
as a species differentiated by morphological characteristics supported by genetic analysis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |