279 lines
80 KiB
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279 lines
80 KiB
XML
<document id="776FA349A53D18D9CF57FB414DBEBA7A" ID-CLB-Dataset="8277" ID-DOI="10.11646/zootaxa.4858.3.1" ID-GBIF-Dataset="f8abea37-7992-482c-8a49-982d08cf9f66" ID-ISSN="1175-5326" ID-Zenodo-Dep="4412321" ID-ZooBank="7BA52F1C-4084-4915-A7D9-8DA99379086B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609688599536" checkinUser="plazi" docAuthor="Uusitalo, Matti, Ueckermann, Edward A. & Theron, Pieter D." docDate="2020" docId="AC6C87BAC466EB6EFF1C67F015EED0AA" docLanguage="en" docName="zootaxa.4858.3.1.pdf" docOrigin="Zootaxa 4858 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alycidae G.Canestrini & Fanzago 1877" docType="key" docVersion="7" lastPageNumber="303" masterDocId="5055FFC2C464EB6CFF8B62731073D77B" masterDocTitle="A review of the family Alycidae (Acari, Acariformes) from South Africa" masterLastPageNumber="340" masterPageNumber="301" pageNumber="303" updateTime="1698851566087" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="75DC5272E7EAF359A85F883D2CCC19DE">A review of the family Alycidae (Acari, Acariformes) from South Africa</mods:title>
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<mods:namePart id="B78840C93D381CCF26E5529F38ABDAB1">Uusitalo, Matti</mods:namePart>
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<mods:affiliation id="83744F126CC476DF0A1FF516C4D69F05">Zoological Museum, Centre for Biodiversity, University of Turku, FIN- 20014 Turku, Finland</mods:affiliation>
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<mods:namePart id="917DEFE0EC9AFA2E80E80E81E83DE580">Ueckermann, Edward A.</mods:namePart>
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<mods:affiliation id="58687602626C83340897FA5375F1E151">Unit for Environmental Sciences and Management, Potchefstroom Campus, North-West University, Private Bag X 6001, Potchefstroom, 2520, South Africa & https: // orcid. org / 0000 - 0003 - 4213 - 4309</mods:affiliation>
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<mods:namePart id="CB36419C0BA1747B237990052E887443">Theron, Pieter D.</mods:namePart>
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<mods:affiliation id="976D3DAF5776F37BFC684AAA9BBBC031">Unit for Environmental Sciences and Management, Potchefstroom Campus, North-West University, Private Bag X 6001, Potchefstroom, 2520, South Africa & https: // orcid. org / 0000 - 0003 - 4213 - 4309</mods:affiliation>
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<mods:title id="A479138D16E8BB3C4685EED932800CC9">Zootaxa</mods:title>
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<mods:date id="7F4BE9B687EC5C2E3F1EC304B741D98E">2020</mods:date>
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<treatment id="AC6C87BAC466EB6EFF1C67F015EED0AA" ID-DOI="http://doi.org/10.5281/zenodo.4412325" ID-GBIF-Taxon="177319609" ID-Zenodo-Dep="4412325" LSID="urn:lsid:plazi:treatment:AC6C87BAC466EB6EFF1C67F015EED0AA" httpUri="http://treatment.plazi.org/id/AC6C87BAC466EB6EFF1C67F015EED0AA" lastPageNumber="303" pageId="2" pageNumber="303">
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<subSubSection id="6CDF6527C466EB6EFF1C67F012E4D2E6" box="[151,663,1411,1437]" pageId="2" pageNumber="303" type="nomenclature">
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<heading id="7F3281C0C466EB6EFF1C67F012E4D2E6" bold="true" box="[151,663,1411,1437]" fontSize="11" level="1" pageId="2" pageNumber="303" reason="1">
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<emphasis id="16B1EABEC466EB6EFF1C67F012E4D2E6" bold="true" box="[151,663,1411,1437]" pageId="2" pageNumber="303">
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Key to subfamilies and tribes of
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<taxonomicName id="E3C54D2FC466EB6EFDA167F012E4D2E6" authorityName="G.Canestrini & Fanzago" authorityYear="1877" box="[554,663,1411,1437]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="2" pageNumber="303" phylum="Arthropoda" rank="family">Alycidae</taxonomicName>
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</emphasis>
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<keyLead id="9F349599C466EB6EFF1C67BB15EFD14E" pageId="2" pageNumber="303">
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1. Cuticular pattern comprising striae with many small lamellae (
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<figureCitation id="BCFE2A29C466EB6EFCC867BB13FBD2A6" box="[835,904,1480,1502]" captionStart="FIGURES 1−5" captionStartId="5.[151,264,1893,1917]" captionTargetBox="[171,1410,188,1853]" captionTargetId="figure-18@5.[165,1421,179,1867]" captionTargetPageId="5" captionText="FIGURES 1−5. Alycus augrabiensis sp. nov. 1—Prodorsum, integument mostly soft on crista, i.e. the area from naso to sensilla (from vi to ve), exp subequal in size with sce; 2—Dorsal habitus; 3—Integument and setal microplates; 4—Ventral habitus, 18–19 genital setae per valve, 2 eugenitals (female), 5 anal setae per valve on PA (cf. Fig. 2); 5—Genital valve of female, 19 genital setae, 1 pair of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412323" httpUri="https://zenodo.org/record/4412323/files/figure.png" pageId="2" pageNumber="303">Figs. 3</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFC1F67BB13DDD2A6" box="[916,942,1480,1501]" captionStart="FIGURES 29−35" captionStartId="11.[151,264,1804,1828]" captionTargetBox="[162,1424,191,1773]" captionTargetId="figure-18@11.[151,1436,181,1781]" captionTargetPageId="11" captionText="FIGURES 29−35. Amphialycus (Amphialycus) acaciae sp. nov. 29—Prodorsum, sclerotized crista, setae vi wide apart from each other; 30—Nasal area in lateral view; 31—Dorsal habitus, holotrichy; 32—Ventral habitus, 22 genital setae per valve, 8–9 anal setae per valve on PA (cf. Fig. 31); 33—Genital valve of female, 21 genital setae, 1 pair of eugenital setae; 34—Chelicera, smooth cheliceral seta more abaxially than the figure suggests; 35—Palp tarsus with 2 pairs of pseudacanthoid eupathids." figureDoi="http://doi.org/10.5281/zenodo.4412337" httpUri="https://zenodo.org/record/4412337/files/figure.png" pageId="2" pageNumber="303">29</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFC3267BB13A0D2A6" box="[953,979,1480,1501]" captionStart="FIGURES 48−53" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[172,1413,196,1808]" captionTargetId="figure-18@15.[160,1426,181,1820]" captionTargetPageId="15" captionText="FIGURES 48−53. Amphialycus (Amphialycus) adustus sp. nov. 48—Prodorsum, sclerotized crista, tapering naso, distance between counterparts of setae vi and sensilla ve subequal; 49—Nasal area in lateral view; 50—Dorsal habitus, neotrichy; 51—Ventral habitus, 21–22 genital setae per valve, 9–10 anal setae per valve; 52—Subcapitulum, rutella with broad shaft, ventrodistal lobes and an apical point,1 pair of adorals; 53—Palp tarsus with 5 barbed eupathids." figureDoi="http://doi.org/10.5281/zenodo.4412345" httpUri="https://zenodo.org/record/4412345/files/figure.png" pageId="2" pageNumber="303">48</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFC5567BB138BD2A5" box="[990,1016,1480,1502]" captionStart="FIGURES 63−67" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[186,1400,189,1865]" captionTargetId="figure-29@18.[179,1409,181,1874]" captionTargetPageId="18" captionText="FIGURES 63−67. Amphialycus (Amphialycus) mayteni sp. nov. 63—Prodorsum, sclerotized crista, reduced naso, distance between counterparts of setae vi and sensilla ve subequal; 64—Nasal area in lateral view; 65—Dorsal habitus, holotrichy; 66—Ventral habitus, neotrichy, 17–20 genital setae per valve, 6 anal setae per valve; 67—Genital valve of male, 21 genital setae, 10 (pairs of) eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412353" httpUri="https://zenodo.org/record/4412353/files/figure.png" pageId="2" pageNumber="303">63</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFB8867BB146ED2A5" box="[1027,1053,1480,1502]" captionStart="FIGURES 76−83" captionStartId="21.[151,264,1868,1892]" captionTargetBox="[182,1405,191,1833]" captionTargetId="figure-18@21.[171,1416,181,1844]" captionTargetPageId="21" captionText="FIGURES 76−83. Amphialycus (Orthacarus) oblongus (Halbert). 76—Prodorsum, naso missing, distance between counterparts of setae vi and sensilla ve smaller than between setae in, setae vi, sce and exp reduced; 77—Dorsal habitus, neotrichy; 78—Ventral habitus, 17 genital setae per valve, 8 anal setae per valve (cf. Fig. 77); 79—Genital valve of female, 16–18 genital setae, 1 pair of eugenital setae; 80—Subcapitulum, 1 pair of adorals; 81—Rutellum; 82—Palp, tarsus with 4 barbed and basally fused eupathids; 83—2 slightly bent claws and densely setulated, claw-like empodium." figureDoi="http://doi.org/10.5281/zenodo.4412359" httpUri="https://zenodo.org/record/4412359/files/figure.png" pageId="2" pageNumber="303">76</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFBA367BB143FD2A5" box="[1064,1100,1480,1502]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="2" pageNumber="303">134</figureCitation>
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C–F); chelicerae chelate, with 2−6 teeth each, and with or without seta (
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<figureCitation id="BCFE2A29C466EB6EFDCB679612E3D280" box="[576,656,1509,1531]" captionStart="FIGURES 16−22" captionStartId="8.[151,264,1897,1921]" captionTargetBox="[167,1414,190,1864]" captionTargetId="figure-29@8.[163,1425,181,1873]" captionTargetPageId="8" captionText="FIGURES 16−22. Pachygnathus nasutus sp. nov. 16—Prodorsum, sclerotized crista, filamentous sensilla ve and sci ciliated, setae exp small; 17—Swollen naso; 18—Beak-like chelicerae, typical to genus; 19—Palp tarsus with 4 pseudacanthoid eupathids; 20—Subcapitulum, 1 pair of adoral setae; 21—Genital valve of female, 13 genital setae, 2 eugenital setae; 22—Tarsus I (2 solenidia), famulus I (arrow), Tibia I (2 solenidia)." figureDoi="http://doi.org/10.5281/zenodo.4412331" httpUri="https://zenodo.org/record/4412331/files/figure.png" pageId="2" pageNumber="303">Figs. 18</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFD12679512C0D280" box="[665,691,1510,1531]" captionStart="FIGURES 29−35" captionStartId="11.[151,264,1804,1828]" captionTargetBox="[162,1424,191,1773]" captionTargetId="figure-18@11.[151,1436,181,1781]" captionTargetPageId="11" captionText="FIGURES 29−35. Amphialycus (Amphialycus) acaciae sp. nov. 29—Prodorsum, sclerotized crista, setae vi wide apart from each other; 30—Nasal area in lateral view; 31—Dorsal habitus, holotrichy; 32—Ventral habitus, 22 genital setae per valve, 8–9 anal setae per valve on PA (cf. Fig. 31); 33—Genital valve of female, 21 genital setae, 1 pair of eugenital setae; 34—Chelicera, smooth cheliceral seta more abaxially than the figure suggests; 35—Palp tarsus with 2 pairs of pseudacanthoid eupathids." figureDoi="http://doi.org/10.5281/zenodo.4412337" httpUri="https://zenodo.org/record/4412337/files/figure.png" pageId="2" pageNumber="303">34</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFD3667961293D280" box="[701,736,1509,1531]" captionStart="FIGURE 135" captionStartId="38.[151,250,1690,1714]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure-29@38.[151,1436,181,1666]" captionTargetPageId="38" captionText="FIGURE 135. Details of Alycidae. A—prodorsum of Petralycus longicornis, B—prodorsum of P. brevicornis, C—palpal tarsi with solenidion ω extending partly over a pair of distal mucronate eupathids, Petralycus sp., D—strongly elongated chelicerae with 5 teeth similar in size on each fixed digit and 2 claws and a clawless empodium (arrow), Petralycus sp., E—cheliceral shafts abruptly tapering into chelae with 2 strong teeth on each fixed digit, Alycus sp., F—cheliceral shafts evenly tapering, elongated, Amphialycus sp. Abbreviations: Na—naso, Ch—chelicera, chb—cheliceral seta, P—palp, I—first leg." figureDoi="http://doi.org/10.5281/zenodo.4412391" httpUri="https://zenodo.org/record/4412391/files/figure.png" pageId="2" pageNumber="303">135</figureCitation>
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D–F;
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<bibRefCitation id="40544B5DC466EB6EFC9C679613E5D280" author="Theron, P. D." box="[791,918,1509,1531]" pageId="2" pageNumber="303" pagination="38 - 45" refId="ref18602" refString="Theron, P. D. (1977) New species of the genus Petralycus Grandjean (Acari: Endeostigmata) from South Africa. Acarologia, 19, 38 - 45." type="journal article" year="1977">Theron 1977</bibRefCitation>
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: fig. 8); rutella well developed (
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<figureCitation id="BCFE2A29C466EB6EFB4D6796157AD280" box="[1222,1289,1509,1531]" captionStart="FIGURES 6−13" captionStartId="6.[151,264,1857,1881]" captionTargetBox="[171,1411,197,1822]" captionTargetId="figure-29@6.[165,1422,181,1834]" captionTargetPageId="6" captionText="FIGURES 6−13. Alycus augrabiensis sp. nov. 6—Rutellum, broad shaft, ventrodistal lobes, and apical point; 7—Subcapitulum, 3 pairs of adorals; 8—Palpal tarsi with 3 pairs of pseudacanthoid eupathids; 9—Genital valve of male, 16 genital setae per valve, 7 (pairs of) eugenitals; 10—Tarsus I (2 solenidia), famulus I (arrow); 11—Tibia I (1 solenidion), Genu I (4 solenidia), Femur I (1 solenidion); 12—Tarsus II (1 solenidion), Tibia II (2 solenidia), Genu II (2 solenidia); 13—Two claws with barbed shafts and a claw-like, oligosetulate empodium." figureDoi="http://doi.org/10.5281/zenodo.4412327" httpUri="https://zenodo.org/record/4412327/files/figure.png" pageId="2" pageNumber="303">Figs. 6</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFA986795155ED280" box="[1299,1325,1510,1531]" captionStart="FIGURES 16−22" captionStartId="8.[151,264,1897,1921]" captionTargetBox="[167,1414,190,1864]" captionTargetId="figure-29@8.[163,1425,181,1873]" captionTargetPageId="8" captionText="FIGURES 16−22. Pachygnathus nasutus sp. nov. 16—Prodorsum, sclerotized crista, filamentous sensilla ve and sci ciliated, setae exp small; 17—Swollen naso; 18—Beak-like chelicerae, typical to genus; 19—Palp tarsus with 4 pseudacanthoid eupathids; 20—Subcapitulum, 1 pair of adoral setae; 21—Genital valve of female, 13 genital setae, 2 eugenital setae; 22—Tarsus I (2 solenidia), famulus I (arrow), Tibia I (2 solenidia)." figureDoi="http://doi.org/10.5281/zenodo.4412331" httpUri="https://zenodo.org/record/4412331/files/figure.png" pageId="2" pageNumber="303">20</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFABD67961523D280" box="[1334,1360,1509,1531]" captionStart="FIGURES 36−42" captionStartId="12.[151,264,1860,1884]" captionTargetBox="[184,1399,192,1831]" captionTargetId="figure-29@12.[176,1411,181,1837]" captionTargetPageId="12" captionText="FIGURES 36−42. Amphialycus (Amphialycus) acaciae sp. nov. 36—Subcapitulum, rutellum with broad shaft and ventrodistal lobes, 2 pairs of adorals; 37—2 claws and multisetulate, claw-like empodium (cf. oligosetulate in Fig. 13); 38—Tarsus I (2 solenidia), famulus I (arrow); 39—Tibia I (2 solenidia), Genu I (3 solenidia), Femur I (2 solenidia); 40—Tarsus II (1 solenidion), famulus II (arrow), Tibia II (2 solenidia), Genu II (2 solenidia); 41—Tarsus III (no solenidia), Tibia III (2 solenidia), Genu III (2 solenidia); 42—Leg IV, Genu IV (1 solenidion)." figureDoi="http://doi.org/10.5281/zenodo.4412339" httpUri="https://zenodo.org/record/4412339/files/figure.png" pageId="2" pageNumber="303">36</figureCitation>
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,
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<figureCitation id="BCFE2A29C466EB6EFAD167961507D280" box="[1370,1396,1509,1531]" captionStart="FIGURES 48−53" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[172,1413,196,1808]" captionTargetId="figure-18@15.[160,1426,181,1820]" captionTargetPageId="15" captionText="FIGURES 48−53. Amphialycus (Amphialycus) adustus sp. nov. 48—Prodorsum, sclerotized crista, tapering naso, distance between counterparts of setae vi and sensilla ve subequal; 49—Nasal area in lateral view; 50—Dorsal habitus, neotrichy; 51—Ventral habitus, 21–22 genital setae per valve, 9–10 anal setae per valve; 52—Subcapitulum, rutella with broad shaft, ventrodistal lobes and an apical point,1 pair of adorals; 53—Palp tarsus with 5 barbed eupathids." figureDoi="http://doi.org/10.5281/zenodo.4412345" httpUri="https://zenodo.org/record/4412345/files/figure.png" pageId="2" pageNumber="303">52</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFAF5679515EAD280" box="[1406,1433,1510,1531]" captionStart="FIGURES 68−75" captionStartId="19.[151,264,1783,1807]" captionTargetBox="[162,1422,193,1749]" captionTargetId="figure-18@19.[151,1436,181,1759]" captionTargetPageId="19" captionText="FIGURES 68−75. Amphialycus (Amphialycus) mayteni sp. nov. 68—Chelicera, robust, elongated, without cheliceral setae; 69—Subcapitulum, rutella with broad shaft, ventrodistal lobes,1 pair of adorals; 70—Rutellum, without apical point; 71—Palp, tarsus with 4 barbed eupathids in 2 groups; 72—Genital valve of female, 24 genital setae, 1 (pair of) eugenital setae; 73—2 slightly bent claws and densely setulated, claw-like empodium; 74—Probably a male tritonymph, genital valve, 15 genital setae per valve, 4 eugenitals, see Kethley (1991); 75—Protonymph, venter, 1 genital seta per valve. 63−75 from RSA, near Potchef- stroom." figureDoi="http://doi.org/10.5281/zenodo.4412355" httpUri="https://zenodo.org/record/4412355/files/figure.png" pageId="2" pageNumber="303">70</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFF6964701112D163" author="Theron, P. D." box="[226,353,1539,1561]" pageId="2" pageNumber="303" pagination="38 - 45" refId="ref18602" refString="Theron, P. D. (1977) New species of the genus Petralycus Grandjean (Acari: Endeostigmata) from South Africa. Acarologia, 19, 38 - 45." type="journal article" year="1977">Theron 1977</bibRefCitation>
|
||
: fig. 10;
|
||
<bibRefCitation id="40544B5DC466EB6EFE3E64701233D163" author="Uusitalo, M." box="[437,576,1539,1561]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: fig. 29); area from naso to anterior sensilla (=crista) more or less sclerotized (
|
||
<figureCitation id="BCFE2A29C466EB6EFABB64701501D163" box="[1328,1394,1539,1561]" captionStart="FIGURES 1−5" captionStartId="5.[151,264,1893,1917]" captionTargetBox="[171,1410,188,1853]" captionTargetId="figure-18@5.[165,1421,179,1867]" captionTargetPageId="5" captionText="FIGURES 1−5. Alycus augrabiensis sp. nov. 1—Prodorsum, integument mostly soft on crista, i.e. the area from naso to sensilla (from vi to ve), exp subequal in size with sce; 2—Dorsal habitus; 3—Integument and setal microplates; 4—Ventral habitus, 18–19 genital setae per valve, 2 eugenitals (female), 5 anal setae per valve on PA (cf. Fig. 2); 5—Genital valve of female, 19 genital setae, 1 pair of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412323" httpUri="https://zenodo.org/record/4412323/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFAF5647015EBD162" box="[1406,1432,1539,1561]" captionStart="FIGURES 16−22" captionStartId="8.[151,264,1897,1921]" captionTargetBox="[167,1414,190,1864]" captionTargetId="figure-29@8.[163,1425,181,1873]" captionTargetPageId="8" captionText="FIGURES 16−22. Pachygnathus nasutus sp. nov. 16—Prodorsum, sclerotized crista, filamentous sensilla ve and sci ciliated, setae exp small; 17—Swollen naso; 18—Beak-like chelicerae, typical to genus; 19—Palp tarsus with 4 pseudacanthoid eupathids; 20—Subcapitulum, 1 pair of adoral setae; 21—Genital valve of female, 13 genital setae, 2 eugenital setae; 22—Tarsus I (2 solenidia), famulus I (arrow), Tibia I (2 solenidia)." figureDoi="http://doi.org/10.5281/zenodo.4412331" httpUri="https://zenodo.org/record/4412331/files/figure.png" pageId="2" pageNumber="303">16</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFF696453108FD14E" box="[226,252,1568,1589]" captionStart="FIGURES 29−35" captionStartId="11.[151,264,1804,1828]" captionTargetBox="[162,1424,191,1773]" captionTargetId="figure-18@11.[151,1436,181,1781]" captionTargetPageId="11" captionText="FIGURES 29−35. Amphialycus (Amphialycus) acaciae sp. nov. 29—Prodorsum, sclerotized crista, setae vi wide apart from each other; 30—Nasal area in lateral view; 31—Dorsal habitus, holotrichy; 32—Ventral habitus, 22 genital setae per valve, 8–9 anal setae per valve on PA (cf. Fig. 31); 33—Genital valve of female, 21 genital setae, 1 pair of eugenital setae; 34—Chelicera, smooth cheliceral seta more abaxially than the figure suggests; 35—Palp tarsus with 2 pairs of pseudacanthoid eupathids." figureDoi="http://doi.org/10.5281/zenodo.4412337" httpUri="https://zenodo.org/record/4412337/files/figure.png" pageId="2" pageNumber="303">29</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFE8D64531153D14E" box="[262,288,1568,1589]" captionStart="FIGURES 48−53" captionStartId="15.[151,264,1844,1868]" captionTargetBox="[172,1413,196,1808]" captionTargetId="figure-18@15.[160,1426,181,1820]" captionTargetPageId="15" captionText="FIGURES 48−53. Amphialycus (Amphialycus) adustus sp. nov. 48—Prodorsum, sclerotized crista, tapering naso, distance between counterparts of setae vi and sensilla ve subequal; 49—Nasal area in lateral view; 50—Dorsal habitus, neotrichy; 51—Ventral habitus, 21–22 genital setae per valve, 9–10 anal setae per valve; 52—Subcapitulum, rutella with broad shaft, ventrodistal lobes and an apical point,1 pair of adorals; 53—Palp tarsus with 5 barbed eupathids." figureDoi="http://doi.org/10.5281/zenodo.4412345" httpUri="https://zenodo.org/record/4412345/files/figure.png" pageId="2" pageNumber="303">48</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFEA164531137D14D" box="[298,324,1568,1590]" captionStart="FIGURES 63−67" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[186,1400,189,1865]" captionTargetId="figure-29@18.[179,1409,181,1874]" captionTargetPageId="18" captionText="FIGURES 63−67. Amphialycus (Amphialycus) mayteni sp. nov. 63—Prodorsum, sclerotized crista, reduced naso, distance between counterparts of setae vi and sensilla ve subequal; 64—Nasal area in lateral view; 65—Dorsal habitus, holotrichy; 66—Ventral habitus, neotrichy, 17–20 genital setae per valve, 6 anal setae per valve; 67—Genital valve of male, 21 genital setae, 10 (pairs of) eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412353" httpUri="https://zenodo.org/record/4412353/files/figure.png" pageId="2" pageNumber="303">63</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFEC56453111BD14D" box="[334,360,1568,1590]" captionStart="FIGURES 76−83" captionStartId="21.[151,264,1868,1892]" captionTargetBox="[182,1405,191,1833]" captionTargetId="figure-18@21.[171,1416,181,1844]" captionTargetPageId="21" captionText="FIGURES 76−83. Amphialycus (Orthacarus) oblongus (Halbert). 76—Prodorsum, naso missing, distance between counterparts of setae vi and sensilla ve smaller than between setae in, setae vi, sce and exp reduced; 77—Dorsal habitus, neotrichy; 78—Ventral habitus, 17 genital setae per valve, 8 anal setae per valve (cf. Fig. 77); 79—Genital valve of female, 16–18 genital setae, 1 pair of eugenital setae; 80—Subcapitulum, 1 pair of adorals; 81—Rutellum; 82—Palp, tarsus with 4 barbed and basally fused eupathids; 83—2 slightly bent claws and densely setulated, claw-like empodium." figureDoi="http://doi.org/10.5281/zenodo.4412359" httpUri="https://zenodo.org/record/4412359/files/figure.png" pageId="2" pageNumber="303">76</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFEF9645311B6D14D" box="[370,453,1568,1590]" captionStart="FIGURE 135" captionStartId="38.[151,250,1690,1714]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure-29@38.[151,1436,181,1666]" captionTargetPageId="38" captionText="FIGURE 135. Details of Alycidae. A—prodorsum of Petralycus longicornis, B—prodorsum of P. brevicornis, C—palpal tarsi with solenidion ω extending partly over a pair of distal mucronate eupathids, Petralycus sp., D—strongly elongated chelicerae with 5 teeth similar in size on each fixed digit and 2 claws and a clawless empodium (arrow), Petralycus sp., E—cheliceral shafts abruptly tapering into chelae with 2 strong teeth on each fixed digit, Alycus sp., F—cheliceral shafts evenly tapering, elongated, Amphialycus sp. Abbreviations: Na—naso, Ch—chelicera, chb—cheliceral seta, P—palp, I—first leg." figureDoi="http://doi.org/10.5281/zenodo.4412391" httpUri="https://zenodo.org/record/4412391/files/figure.png" pageId="2" pageNumber="303">135A, B</figureCitation>
|
||
). (
|
||
<taxonomicName id="E3C54D2FC466EB6EFE556453124BD14D" authorityName="G.Canestrini & Fanzago" authorityYear="1877" box="[478,568,1568,1590]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="2" pageNumber="303" phylum="Arthropoda" rank="subFamily" subFamily="Alycinae">Alycinae</taxonomicName>
|
||
)...................................................................... 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="247A36ACC466EB6EFF1C644D15EFD1D0" blockId="2.[151,1437,1480,2001]" pageId="2" pageNumber="303">
|
||
<keyLead id="9F349599C466EB6EFF1C644D15EFD1D0" pageId="2" pageNumber="303">
|
||
- Cuticular patterning more complex, often reticulate or rosetted primary pattern, with many large lamellae or formed only by large lamellae (
|
||
<figureCitation id="BCFE2A29C466EB6EFEF2642811B8D10B" box="[377,459,1627,1649]" captionStart="FIGURES 90−100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90−100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 9–10 genital setae per valve, 4–6 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="2" pageNumber="303">Figs. 92</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFE5C6428118ED10A" box="[471,509,1627,1649]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">105</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFD836428125ED10B" box="[520,557,1627,1648]" captionStart="FIGURES 119−125" captionStartId="32.[151,264,1868,1892]" captionTargetBox="[175,1414,193,1835]" captionTargetId="figure-29@32.[164,1423,181,1844]" captionTargetPageId="32" captionText="FIGURES 119−125. Laminamichaelia furcula sp. nov. 119—Prodorsum, neotrichous soft integument and in-area, sensilla ve branched, button-like naso, reticulated pattern of large lamellae on crista between strips; 120—Dorsal habitus, strong neotrichy; 121—Vental habitus, neotrichy, 9–10 genital setae per valve, 7–8 anal setae per valve; 122—Palp tarsus; 123—Subcapitulum, neotrichy; 124—Genital area, 10 genital setae per valve, 3 pairs of eugenital setae; 125—Tarsus I (3B), famulus I (arrow), Tibia I (3B1P)." figureDoi="http://doi.org/10.5281/zenodo.4412377" httpUri="https://zenodo.org/record/4412377/files/figure.png" pageId="2" pageNumber="303">119</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFDB264281228D10B" box="[569,603,1627,1648]" captionStart="FIGURE 133" captionStartId="36.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1692]" captionTargetId="figure-29@36.[151,1436,181,1692]" captionTargetPageId="36" captionText="FIGURE 133. Primary and secondary patterns of cuticle in the tribe Bimichaeliini (Bimichaeliinae). A, B—stellate primary pattern by large lamellae, secondary pattern by small lamellae, setae prominent in Laminamichaelia sp. of arbusculosa-group, C, D—primary pattern by clumps of large lamellae connected by small lamellae, setae prominent in Laminamichaelia sp. of subnuda-group, E, F—primary pattern by roundish cells of large lamellae, secondary pattern by small granulae, setae inconspicuous in Bimichaelia sp. Abbreviations: Ch—chelicera, P—palp, Gen—genitalia, C, D, E, F, H, PS—segments of opisthosoma." figureDoi="http://doi.org/10.5281/zenodo.4412385" httpUri="https://zenodo.org/record/4412385/files/figure.png" pageId="2" pageNumber="303">133</figureCitation>
|
||
A–F, 134A, B); chelicerae stylet-like (
|
||
<figureCitation id="BCFE2A29C466EB6EFC5E64281454D10B" box="[981,1063,1627,1649]" captionStart="FIGURES 90−100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90−100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 9–10 genital setae per valve, 4–6 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="2" pageNumber="303">Figs. 94</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFBB96428142AD10B" box="[1074,1113,1627,1648]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">108</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFBEE64281481D10B" author="Uusitalo, M." box="[1125,1266,1627,1649]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: fig. 123); rutella absent (
|
||
<figureCitation id="BCFE2A29C466EB6EFEA0640B1108D1F5" box="[299,379,1656,1678]" captionStart="FIGURES 90−100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90−100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 9–10 genital setae per valve, 4–6 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="2" pageNumber="303">Figs. 95</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFE0F640B11D9D1F6" box="[388,426,1656,1677]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">109</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFE38640B11A9D1F6" box="[435,474,1656,1677]" captionStart="FIGURES 119−125" captionStartId="32.[151,264,1868,1892]" captionTargetBox="[175,1414,193,1835]" captionTargetId="figure-29@32.[164,1423,181,1844]" captionTargetPageId="32" captionText="FIGURES 119−125. Laminamichaelia furcula sp. nov. 119—Prodorsum, neotrichous soft integument and in-area, sensilla ve branched, button-like naso, reticulated pattern of large lamellae on crista between strips; 120—Dorsal habitus, strong neotrichy; 121—Vental habitus, neotrichy, 9–10 genital setae per valve, 7–8 anal setae per valve; 122—Palp tarsus; 123—Subcapitulum, neotrichy; 124—Genital area, 10 genital setae per valve, 3 pairs of eugenital setae; 125—Tarsus I (3B), famulus I (arrow), Tibia I (3B1P)." figureDoi="http://doi.org/10.5281/zenodo.4412377" httpUri="https://zenodo.org/record/4412377/files/figure.png" pageId="2" pageNumber="303">123</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFE69640B121ED1F6" author="Uusitalo, M." box="[482,621,1656,1678]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: fig. 92); crista sclerotized or smooth (
|
||
<figureCitation id="BCFE2A29C466EB6EFC53640B1454D1F6" box="[984,1063,1656,1678]" captionStart="FIGURES 90−100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90−100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 9–10 genital setae per valve, 4–6 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="2" pageNumber="303">Figs. 90</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFBBB640B1425D1F5" box="[1072,1110,1656,1678]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">105</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFBD4640B14F6D1F6" box="[1119,1157,1656,1677]" captionStart="FIGURES 119−125" captionStartId="32.[151,264,1868,1892]" captionTargetBox="[175,1414,193,1835]" captionTargetId="figure-29@32.[164,1423,181,1844]" captionTargetPageId="32" captionText="FIGURES 119−125. Laminamichaelia furcula sp. nov. 119—Prodorsum, neotrichous soft integument and in-area, sensilla ve branched, button-like naso, reticulated pattern of large lamellae on crista between strips; 120—Dorsal habitus, strong neotrichy; 121—Vental habitus, neotrichy, 9–10 genital setae per valve, 7–8 anal setae per valve; 122—Palp tarsus; 123—Subcapitulum, neotrichy; 124—Genital area, 10 genital setae per valve, 3 pairs of eugenital setae; 125—Tarsus I (3B), famulus I (arrow), Tibia I (3B1P)." figureDoi="http://doi.org/10.5281/zenodo.4412377" httpUri="https://zenodo.org/record/4412377/files/figure.png" pageId="2" pageNumber="303">119</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFB05640B156BD1F6" author="Uusitalo, M." box="[1166,1304,1656,1678]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: figs. 90, 100, 110, 120, 130); posterior pair of sensilla globular (
|
||
<figureCitation id="BCFE2A29C466EB6EFD4E64E6136AD1D0" box="[709,793,1685,1707]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">Fig. 106</figureCitation>
|
||
); empodia clawless (
|
||
<figureCitation id="BCFE2A29C466EB6EFC6964E6144DD1D0" box="[994,1086,1685,1707]" captionStart="FIGURES 90−100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90−100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 9–10 genital setae per valve, 4–6 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="2" pageNumber="303">Figs. 100</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFBC364E5141ED1D0" box="[1096,1133,1686,1707]" captionStart="FIGURES 112−116" captionStartId="29.[151,264,1852,1876]" captionTargetBox="[160,1426,193,1816]" captionTargetId="figure-18@29.[151,1436,181,1828]" captionTargetPageId="29" captionText="FIGURES 112−116. Laminamichaelia shibai nom. nov. 112—Genital area of male, 5 genital setae per valve, 6 (pairs of) eugenital setae; 113—Tarsus I (2B), famulus I (arrow), Tibia I (2B1P = 2 baculiform solenidia and 1 piliform solenidion), Genu I (4P); 114—Tarsus II (1B), Tibia II (2B1P), Genu II (1B2P); 115—Deutonymph, venter, 3 genital setae per valve, 3 anal setae per valve; 116—Protonymph, venter, 1 genital seta per valve, 3–4 anal setae per valve." figureDoi="http://doi.org/10.5281/zenodo.4412371" httpUri="https://zenodo.org/record/4412371/files/figure.png" pageId="2" pageNumber="303">113</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFBFC64E514EDD1D7" box="[1143,1182,1686,1708]" captionStart="FIGURES 126−130" captionStartId="33.[151,264,1889,1913]" captionTargetBox="[158,1425,193,1855]" captionTargetId="figure-18@33.[151,1436,181,1865]" captionTargetPageId="33" captionText="FIGURES 126−130. Laminamichaelia furcula sp. nov. 126—Tarsus II (2B), Tibia II (2B); 127—2 slightly bent claws and a setulated empodium; 128—Tritonymph, venter, 6−7 genital setae per valve, 5−6 anal setae per valve; 129—Deutonymph, venter, 4 genital setae per valve, 4−6 anal setae per valve; 130—Protonymph, venter, 1 genital seta per valve, 4 anal setae per valve." figureDoi="http://doi.org/10.5281/zenodo.4412379" httpUri="https://zenodo.org/record/4412379/files/figure.png" pageId="2" pageNumber="303">127</figureCitation>
|
||
). (
|
||
<taxonomicName id="E3C54D2FC466EB6EFB3D64E61538D1D0" authorityName="Womersley" authorityYear="1944" box="[1206,1355,1685,1707]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="2" pageNumber="303" phylum="Arthropoda" rank="subFamily" subFamily="Bimichaeliinae">Bimichaeliinae</taxonomicName>
|
||
)..... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="9F312E09C466EB6EFF1C64C015EFD020" pageId="2" pageNumber="303">
|
||
<paragraph id="247A36ACC466EB6EFF1C64C015EFD078" blockId="2.[151,1437,1480,2001]" pageId="2" pageNumber="303">
|
||
<keyLead id="9F349599C466EB6EFF1C64C015EFD078" pageId="2" pageNumber="303">
|
||
2. Both pairs of prodorsal sensilla are either filamentous (
|
||
<figureCitation id="BCFE2A29C466EB6EFD6464C01355D1B2" box="[751,806,1715,1737]" captionStart="FIGURES 1−5" captionStartId="5.[151,264,1893,1917]" captionTargetBox="[171,1410,188,1853]" captionTargetId="figure-18@5.[165,1421,179,1867]" captionTargetPageId="5" captionText="FIGURES 1−5. Alycus augrabiensis sp. nov. 1—Prodorsum, integument mostly soft on crista, i.e. the area from naso to sensilla (from vi to ve), exp subequal in size with sce; 2—Dorsal habitus; 3—Integument and setal microplates; 4—Ventral habitus, 18–19 genital setae per valve, 2 eugenitals (female), 5 anal setae per valve on PA (cf. Fig. 2); 5—Genital valve of female, 19 genital setae, 1 pair of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412323" httpUri="https://zenodo.org/record/4412323/files/figure.png" pageId="2" pageNumber="303">Fig. 1</figureCitation>
|
||
) or slightly swollen (
|
||
<figureCitation id="BCFE2A29C466EB6EFC7864C01448D1B2" box="[1011,1083,1715,1737]" captionStart="FIGURES 23−28" captionStartId="9.[151,264,1908,1932]" captionTargetBox="[185,1396,189,1873]" captionTargetId="figure-18@9.[176,1411,181,1884]" captionTargetPageId="9" captionText="FIGURES 23−28. Pachygnathus nasutus sp. nov. 23—Genital valve of male, 15 genital setae per valve, 10 pairs of eugenitals; 24—Deutonymph, genital area, 5 genital setae per valve; 25—Protonymph, genital area, 1 genital seta per valve; 26—Protonymph, subcapitulum; 27—Dorsal habitus of larva, holotrichy, sensilla ve and sci slightly swollen, 1 pair of palpal eupathids; 28—Ventral habitus of larva, genital area absent, 3–4 anal setae per valve. 16−28 from RSA, near Potchefstroom." figureDoi="http://doi.org/10.5281/zenodo.4412335" httpUri="https://zenodo.org/record/4412335/files/figure.png" pageId="2" pageNumber="303">Fig. 27</figureCitation>
|
||
); chelicerae with 1−2 strong teeth on each fixed digit (
|
||
<figureCitation id="BCFE2A29C466EB6EFE0E64A311A6D19E" box="[389,469,1744,1766]" captionStart="FIGURES 29−35" captionStartId="11.[151,264,1804,1828]" captionTargetBox="[162,1424,191,1773]" captionTargetId="figure-18@11.[151,1436,181,1781]" captionTargetPageId="11" captionText="FIGURES 29−35. Amphialycus (Amphialycus) acaciae sp. nov. 29—Prodorsum, sclerotized crista, setae vi wide apart from each other; 30—Nasal area in lateral view; 31—Dorsal habitus, holotrichy; 32—Ventral habitus, 22 genital setae per valve, 8–9 anal setae per valve on PA (cf. Fig. 31); 33—Genital valve of female, 21 genital setae, 1 pair of eugenital setae; 34—Chelicera, smooth cheliceral seta more abaxially than the figure suggests; 35—Palp tarsus with 2 pairs of pseudacanthoid eupathids." figureDoi="http://doi.org/10.5281/zenodo.4412337" httpUri="https://zenodo.org/record/4412337/files/figure.png" pageId="2" pageNumber="303">
|
||
Figs.
|
||
<geoCoordinate id="41F1506BC466EB6EFE3064A311A6D19E" box="[443,469,1744,1765]" degrees="34, 135" direction="east" orientation="longitude" pageId="2" pageNumber="303" precision="55" value="34.135">34</geoCoordinate>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFE5464A3125DD19D" box="[479,558,1744,1766]" captionStart="FIGURE 135" captionStartId="38.[151,250,1690,1714]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure-29@38.[151,1436,181,1666]" captionTargetPageId="38" captionText="FIGURE 135. Details of Alycidae. A—prodorsum of Petralycus longicornis, B—prodorsum of P. brevicornis, C—palpal tarsi with solenidion ω extending partly over a pair of distal mucronate eupathids, Petralycus sp., D—strongly elongated chelicerae with 5 teeth similar in size on each fixed digit and 2 claws and a clawless empodium (arrow), Petralycus sp., E—cheliceral shafts abruptly tapering into chelae with 2 strong teeth on each fixed digit, Alycus sp., F—cheliceral shafts evenly tapering, elongated, Amphialycus sp. Abbreviations: Na—naso, Ch—chelicera, chb—cheliceral seta, P—palp, I—first leg." figureDoi="http://doi.org/10.5281/zenodo.4412391" httpUri="https://zenodo.org/record/4412391/files/figure.png" pageId="2" pageNumber="303">135E, F</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFDB364A312B7D19E" author="Uusitalo, M." box="[568,708,1744,1766]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: fig. 63); usually with a pair of rudimentary posterolateral eyes (
|
||
<figureCitation id="BCFE2A29C466EB6EFABB64A31501D19E" box="[1328,1394,1744,1766]" captionStart="FIGURES 1−5" captionStartId="5.[151,264,1893,1917]" captionTargetBox="[171,1410,188,1853]" captionTargetId="figure-18@5.[165,1421,179,1867]" captionTargetPageId="5" captionText="FIGURES 1−5. Alycus augrabiensis sp. nov. 1—Prodorsum, integument mostly soft on crista, i.e. the area from naso to sensilla (from vi to ve), exp subequal in size with sce; 2—Dorsal habitus; 3—Integument and setal microplates; 4—Ventral habitus, 18–19 genital setae per valve, 2 eugenitals (female), 5 anal setae per valve on PA (cf. Fig. 2); 5—Genital valve of female, 19 genital setae, 1 pair of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412323" httpUri="https://zenodo.org/record/4412323/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFAF564A315EBD19D" box="[1406,1432,1744,1766]" captionStart="FIGURES 16−22" captionStartId="8.[151,264,1897,1921]" captionTargetBox="[167,1414,190,1864]" captionTargetId="figure-29@8.[163,1425,181,1873]" captionTargetPageId="8" captionText="FIGURES 16−22. Pachygnathus nasutus sp. nov. 16—Prodorsum, sclerotized crista, filamentous sensilla ve and sci ciliated, setae exp small; 17—Swollen naso; 18—Beak-like chelicerae, typical to genus; 19—Palp tarsus with 4 pseudacanthoid eupathids; 20—Subcapitulum, 1 pair of adoral setae; 21—Genital valve of female, 13 genital setae, 2 eugenital setae; 22—Tarsus I (2 solenidia), famulus I (arrow), Tibia I (2 solenidia)." figureDoi="http://doi.org/10.5281/zenodo.4412331" httpUri="https://zenodo.org/record/4412331/files/figure.png" pageId="2" pageNumber="303">16</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFF69649E108ED078" box="[226,253,1773,1795]" captionStart="FIGURES 63−67" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[186,1400,189,1865]" captionTargetId="figure-29@18.[179,1409,181,1874]" captionTargetPageId="18" captionText="FIGURES 63−67. Amphialycus (Amphialycus) mayteni sp. nov. 63—Prodorsum, sclerotized crista, reduced naso, distance between counterparts of setae vi and sensilla ve subequal; 64—Nasal area in lateral view; 65—Dorsal habitus, holotrichy; 66—Ventral habitus, neotrichy, 17–20 genital setae per valve, 6 anal setae per valve; 67—Genital valve of male, 21 genital setae, 10 (pairs of) eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412353" httpUri="https://zenodo.org/record/4412353/files/figure.png" pageId="2" pageNumber="303">63</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFE8C649E11E1D078" author="Uusitalo, M." box="[263,402,1773,1795]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: figs. 12, 75, 82); empodia with an apical hook (
|
||
<figureCitation id="BCFE2A29C466EB6EFCE9649E13C1D078" box="[866,946,1773,1795]" captionStart="FIGURES 6−13" captionStartId="6.[151,264,1857,1881]" captionTargetBox="[171,1411,197,1822]" captionTargetId="figure-29@6.[165,1422,181,1834]" captionTargetPageId="6" captionText="FIGURES 6−13. Alycus augrabiensis sp. nov. 6—Rutellum, broad shaft, ventrodistal lobes, and apical point; 7—Subcapitulum, 3 pairs of adorals; 8—Palpal tarsi with 3 pairs of pseudacanthoid eupathids; 9—Genital valve of male, 16 genital setae per valve, 7 (pairs of) eugenitals; 10—Tarsus I (2 solenidia), famulus I (arrow); 11—Tibia I (1 solenidion), Genu I (4 solenidia), Femur I (1 solenidion); 12—Tarsus II (1 solenidion), Tibia II (2 solenidia), Genu II (2 solenidia); 13—Two claws with barbed shafts and a claw-like, oligosetulate empodium." figureDoi="http://doi.org/10.5281/zenodo.4412327" httpUri="https://zenodo.org/record/4412327/files/figure.png" pageId="2" pageNumber="303">Figs. 13</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFC37649D13A5D078" box="[956,982,1774,1795]" captionStart="FIGURES 16−22" captionStartId="8.[151,264,1897,1921]" captionTargetBox="[167,1414,190,1864]" captionTargetId="figure-29@8.[163,1425,181,1873]" captionTargetPageId="8" captionText="FIGURES 16−22. Pachygnathus nasutus sp. nov. 16—Prodorsum, sclerotized crista, filamentous sensilla ve and sci ciliated, setae exp small; 17—Swollen naso; 18—Beak-like chelicerae, typical to genus; 19—Palp tarsus with 4 pseudacanthoid eupathids; 20—Subcapitulum, 1 pair of adoral setae; 21—Genital valve of female, 13 genital setae, 2 eugenital setae; 22—Tarsus I (2 solenidia), famulus I (arrow), Tibia I (2 solenidia)." figureDoi="http://doi.org/10.5281/zenodo.4412331" httpUri="https://zenodo.org/record/4412331/files/figure.png" pageId="2" pageNumber="303">22</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFC6B649D1389D078" box="[992,1018,1774,1795]" captionStart="FIGURES 36−42" captionStartId="12.[151,264,1860,1884]" captionTargetBox="[184,1399,192,1831]" captionTargetId="figure-29@12.[176,1411,181,1837]" captionTargetPageId="12" captionText="FIGURES 36−42. Amphialycus (Amphialycus) acaciae sp. nov. 36—Subcapitulum, rutellum with broad shaft and ventrodistal lobes, 2 pairs of adorals; 37—2 claws and multisetulate, claw-like empodium (cf. oligosetulate in Fig. 13); 38—Tarsus I (2 solenidia), famulus I (arrow); 39—Tibia I (2 solenidia), Genu I (3 solenidia), Femur I (2 solenidia); 40—Tarsus II (1 solenidion), famulus II (arrow), Tibia II (2 solenidia), Genu II (2 solenidia); 41—Tarsus III (no solenidia), Tibia III (2 solenidia), Genu III (2 solenidia); 42—Leg IV, Genu IV (1 solenidion)." figureDoi="http://doi.org/10.5281/zenodo.4412339" httpUri="https://zenodo.org/record/4412339/files/figure.png" pageId="2" pageNumber="303">37</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFB8F649E146DD078" box="[1028,1054,1773,1795]" captionStart="FIGURES 54−59" captionStartId="16.[151,264,1881,1905]" captionTargetBox="[157,1428,189,1845]" captionTargetId="figure-29@16.[151,1436,181,1857]" captionTargetPageId="16" captionText="FIGURES 54−59. Amphialycus (Amphialycus) adustus sp. nov. 54—Genital valve of male with 19 genital setae, 10 (pairs of) eugenital setae; 55—Tarsus I (2 solenidia); 56—Tarsus II (2 solenidia), famulus II (arrow); 57—2 slightly bent claws and densely setulated, claw-like empodium; 58—Tritonymph, venter, 12–15 genital setae per valve; 59—Deutonymph, venter, 6 genital setae per valve." figureDoi="http://doi.org/10.5281/zenodo.4412347" httpUri="https://zenodo.org/record/4412347/files/figure.png" pageId="2" pageNumber="303">57</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFBA3649D1431D078" box="[1064,1090,1774,1795]" captionStart="FIGURES 68−75" captionStartId="19.[151,264,1783,1807]" captionTargetBox="[162,1422,193,1749]" captionTargetId="figure-18@19.[151,1436,181,1759]" captionTargetPageId="19" captionText="FIGURES 68−75. Amphialycus (Amphialycus) mayteni sp. nov. 68—Chelicera, robust, elongated, without cheliceral setae; 69—Subcapitulum, rutella with broad shaft, ventrodistal lobes,1 pair of adorals; 70—Rutellum, without apical point; 71—Palp, tarsus with 4 barbed eupathids in 2 groups; 72—Genital valve of female, 24 genital setae, 1 (pair of) eugenital setae; 73—2 slightly bent claws and densely setulated, claw-like empodium; 74—Probably a male tritonymph, genital valve, 15 genital setae per valve, 4 eugenitals, see Kethley (1991); 75—Protonymph, venter, 1 genital seta per valve. 63−75 from RSA, near Potchef- stroom." figureDoi="http://doi.org/10.5281/zenodo.4412355" httpUri="https://zenodo.org/record/4412355/files/figure.png" pageId="2" pageNumber="303">73</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFBC7649D1416D07F" box="[1100,1125,1774,1796]" captionStart="FIGURES 76−83" captionStartId="21.[151,264,1868,1892]" captionTargetBox="[182,1405,191,1833]" captionTargetId="figure-18@21.[171,1416,181,1844]" captionTargetPageId="21" captionText="FIGURES 76−83. Amphialycus (Orthacarus) oblongus (Halbert). 76—Prodorsum, naso missing, distance between counterparts of setae vi and sensilla ve smaller than between setae in, setae vi, sce and exp reduced; 77—Dorsal habitus, neotrichy; 78—Ventral habitus, 17 genital setae per valve, 8 anal setae per valve (cf. Fig. 77); 79—Genital valve of female, 16–18 genital setae, 1 pair of eugenital setae; 80—Subcapitulum, 1 pair of adorals; 81—Rutellum; 82—Palp, tarsus with 4 barbed and basally fused eupathids; 83—2 slightly bent claws and densely setulated, claw-like empodium." figureDoi="http://doi.org/10.5281/zenodo.4412359" httpUri="https://zenodo.org/record/4412359/files/figure.png" pageId="2" pageNumber="303">83</figureCitation>
|
||
)...................
|
||
<taxonomicName id="E3C54D2FC466EB6EFADF649E15EFD078" authorityName="Canestrini & Fanzago" authorityYear="1877" box="[1364,1436,1773,1795]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="2" pageNumber="303" phylum="Arthropoda" rank="tribe" tribe="Alycini">Alycini</taxonomicName>
|
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</keyLead>
|
||
</paragraph>
|
||
<paragraph id="247A36ACC466EB6EFF1C657815EFD020" blockId="2.[151,1437,1480,2001]" pageId="2" pageNumber="303">
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<keyLead id="9F349599C466EB6EFF1C657815EFD020" pageId="2" pageNumber="303">
|
||
- Anterior pair of sensilla globular (
|
||
<figureCitation id="BCFE2A29C466EB6EFDBF657812B2D05B" box="[564,705,1803,1825]" captionStart="FIGURE 135" captionStartId="38.[151,250,1690,1714]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure-29@38.[151,1436,181,1666]" captionTargetPageId="38" captionText="FIGURE 135. Details of Alycidae. A—prodorsum of Petralycus longicornis, B—prodorsum of P. brevicornis, C—palpal tarsi with solenidion ω extending partly over a pair of distal mucronate eupathids, Petralycus sp., D—strongly elongated chelicerae with 5 teeth similar in size on each fixed digit and 2 claws and a clawless empodium (arrow), Petralycus sp., E—cheliceral shafts abruptly tapering into chelae with 2 strong teeth on each fixed digit, Alycus sp., F—cheliceral shafts evenly tapering, elongated, Amphialycus sp. Abbreviations: Na—naso, Ch—chelicera, chb—cheliceral seta, P—palp, I—first leg." figureDoi="http://doi.org/10.5281/zenodo.4412391" httpUri="https://zenodo.org/record/4412391/files/figure.png" pageId="2" pageNumber="303">Figs. 135A, B</figureCitation>
|
||
); chelicerae with 3−6 teeth similar in size on each fixed digit (
|
||
<figureCitation id="BCFE2A29C466EB6EFABA657815EBD05A" box="[1329,1432,1803,1825]" captionStart="FIGURE 135" captionStartId="38.[151,250,1690,1714]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure-29@38.[151,1436,181,1666]" captionTargetPageId="38" captionText="FIGURE 135. Details of Alycidae. A—prodorsum of Petralycus longicornis, B—prodorsum of P. brevicornis, C—palpal tarsi with solenidion ω extending partly over a pair of distal mucronate eupathids, Petralycus sp., D—strongly elongated chelicerae with 5 teeth similar in size on each fixed digit and 2 claws and a clawless empodium (arrow), Petralycus sp., E—cheliceral shafts abruptly tapering into chelae with 2 strong teeth on each fixed digit, Alycus sp., F—cheliceral shafts evenly tapering, elongated, Amphialycus sp. Abbreviations: Na—naso, Ch—chelicera, chb—cheliceral seta, P—palp, I—first leg." figureDoi="http://doi.org/10.5281/zenodo.4412391" httpUri="https://zenodo.org/record/4412391/files/figure.png" pageId="2" pageNumber="303">Fig. 135D</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFF69655B1111D046" author="Theron, P. D." box="[226,354,1832,1854]" pageId="2" pageNumber="303" pagination="38 - 45" refId="ref18602" refString="Theron, P. D. (1977) New species of the genus Petralycus Grandjean (Acari: Endeostigmata) from South Africa. Acarologia, 19, 38 - 45." type="journal article" year="1977">Theron 1977</bibRefCitation>
|
||
: fig. 8;
|
||
<bibRefCitation id="40544B5DC466EB6EFE25655B1249D046" author="Uusitalo, M." box="[430,570,1832,1854]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: fig. 143); eyes absent (
|
||
<figureCitation id="BCFE2A29C466EB6EFCA8655B13DDD045" box="[803,942,1832,1854]" captionStart="FIGURE 135" captionStartId="38.[151,250,1690,1714]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure-29@38.[151,1436,181,1666]" captionTargetPageId="38" captionText="FIGURE 135. Details of Alycidae. A—prodorsum of Petralycus longicornis, B—prodorsum of P. brevicornis, C—palpal tarsi with solenidion ω extending partly over a pair of distal mucronate eupathids, Petralycus sp., D—strongly elongated chelicerae with 5 teeth similar in size on each fixed digit and 2 claws and a clawless empodium (arrow), Petralycus sp., E—cheliceral shafts abruptly tapering into chelae with 2 strong teeth on each fixed digit, Alycus sp., F—cheliceral shafts evenly tapering, elongated, Amphialycus sp. Abbreviations: Na—naso, Ch—chelicera, chb—cheliceral seta, P—palp, I—first leg." figureDoi="http://doi.org/10.5281/zenodo.4412391" httpUri="https://zenodo.org/record/4412391/files/figure.png" pageId="2" pageNumber="303">Figs. 135A, B</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFC31655B1449D046" author="Theron, P. D." box="[954,1082,1832,1854]" pageId="2" pageNumber="303" pagination="38 - 45" refId="ref18602" refString="Theron, P. D. (1977) New species of the genus Petralycus Grandjean (Acari: Endeostigmata) from South Africa. Acarologia, 19, 38 - 45." type="journal article" year="1977">Theron 1977</bibRefCitation>
|
||
: figs. 2, 16); empodia clawless (
|
||
<figureCitation id="BCFE2A29C466EB6EFAFE655B1169D020" captionStart="FIGURE 135" captionStartId="38.[151,250,1690,1714]" captionTargetBox="[151,1436,181,1666]" captionTargetId="figure-29@38.[151,1436,181,1666]" captionTargetPageId="38" captionText="FIGURE 135. Details of Alycidae. A—prodorsum of Petralycus longicornis, B—prodorsum of P. brevicornis, C—palpal tarsi with solenidion ω extending partly over a pair of distal mucronate eupathids, Petralycus sp., D—strongly elongated chelicerae with 5 teeth similar in size on each fixed digit and 2 claws and a clawless empodium (arrow), Petralycus sp., E—cheliceral shafts abruptly tapering into chelae with 2 strong teeth on each fixed digit, Alycus sp., F—cheliceral shafts evenly tapering, elongated, Amphialycus sp. Abbreviations: Na—naso, Ch—chelicera, chb—cheliceral seta, P—palp, I—first leg." figureDoi="http://doi.org/10.5281/zenodo.4412391" httpUri="https://zenodo.org/record/4412391/files/figure.png" pageId="2" pageNumber="303">Fig. 135D</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFEA8653611D0D020" author="Theron, P. D." box="[291,419,1861,1883]" pageId="2" pageNumber="303" pagination="38 - 45" refId="ref18602" refString="Theron, P. D. (1977) New species of the genus Petralycus Grandjean (Acari: Endeostigmata) from South Africa. Acarologia, 19, 38 - 45." type="journal article" year="1977">Theron 1977</bibRefCitation>
|
||
: figs. 11, 27).................................................................
|
||
<taxonomicName id="E3C54D2FC466EB6EFABF653615EFD020" authorityName="Uusitalo" authorityYear="2010" box="[1332,1436,1861,1883]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="2" pageNumber="303" phylum="Arthropoda" rank="tribe" tribe="Petralycini">Petralycini</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="9F312E09C466EB6EFF1C651015EED0AA" pageId="2" pageNumber="303">
|
||
<paragraph id="247A36ACC466EB6EFF1C651015EFD0EE" blockId="2.[151,1437,1480,2001]" pageId="2" pageNumber="303">
|
||
<keyLead id="9F349599C466EB6EFF1C651015EFD0EE" pageId="2" pageNumber="303">
|
||
3. Large lamellae in rosettes and in clumps, secondary pattern absent i.e. ridges glabrous (
|
||
<figureCitation id="BCFE2A29C466EB6EFBA065101408D003" box="[1067,1147,1891,1913]" captionStart="FIGURES 90−100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90−100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 9–10 genital setae per valve, 4–6 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="2" pageNumber="303">Figs. 92</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFB0E651014AAD003" box="[1157,1241,1891,1912]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="2" pageNumber="303">134A, B</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFB6F6510151CD003" author="Uusitalo, M." box="[1252,1391,1891,1913]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: Appendix 3) cheliceral seta present (
|
||
<figureCitation id="BCFE2A29C466EB6EFDAF65F3121FD0ED" box="[548,620,1920,1942]" captionStart="FIGURES 90−100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90−100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 9–10 genital setae per valve, 4–6 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="2" pageNumber="303">Fig. 94</figureCitation>
|
||
); prodorsum with a pair of eyes (
|
||
<figureCitation id="BCFE2A29C466EB6EFC2165F3138FD0ED" box="[938,1020,1920,1942]" captionStart="FIGURES 101−104" captionStartId="26.[151,264,1888,1912]" captionTargetBox="[160,1426,188,1854]" captionTargetId="figure-29@26.[151,1436,181,1864]" captionTargetPageId="26" captionText="FIGURES 101−104. Proteromichaelia sila sp. nov. 101—Deutonymph, venter, 4 genital setae per valve; 102—Protonymph, venter, 1 genital seta per valve; 103—Larva, ventral habitus, holotrichy, genital valves absent; 104—Larva, dorsal habitus, neotrichy. 90−104 from RSA, near Potchefstroom." figureDoi="http://doi.org/10.5281/zenodo.4412367" httpUri="https://zenodo.org/record/4412367/files/figure.png" pageId="2" pageNumber="303">Fig. 104</figureCitation>
|
||
)...........
|
||
<taxonomicName id="E3C54D2FC466EB6EFB0C65F31549D0ED" authority="Uusitalo & Ueckermann & Theron, 2020" authorityName="Uusitalo & Ueckermann & Theron" authorityYear="2020" box="[1159,1338,1920,1942]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Actinedida" pageId="2" pageNumber="303" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Proteromichaeliini">Proteromichaeliini</taxonomicName>
|
||
<emphasis id="16B1EABEC466EB6EFACB65F315EFD0EE" bold="true" box="[1344,1436,1920,1942]" pageId="2" pageNumber="303">
|
||
<taxonomicNameLabel id="0D8257C5C466EB6EFACB65F315EFD0EE" box="[1344,1436,1920,1942]" pageId="2" pageNumber="303" rank="tribe">trib. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="247A36ACC466EB6EFF1C65ED15EED0AA" blockId="2.[151,1437,1480,2001]" pageId="2" pageNumber="303">
|
||
<keyLead id="9F349599C466EB6EFF1C65ED15EED0AA" pageId="2" pageNumber="303">
|
||
- Large lamellae in rosettes or in clumps, secondary pattern granular or by small lamellae (
|
||
<figureCitation id="BCFE2A29C466EB6EFBC865EE14D3D0C8" box="[1091,1184,1949,1971]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">Figs. 105</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFB2065ED14A3D0C8" box="[1195,1232,1950,1971]" captionStart="FIGURES 119−125" captionStartId="32.[151,264,1868,1892]" captionTargetBox="[175,1414,193,1835]" captionTargetId="figure-29@32.[164,1423,181,1844]" captionTargetPageId="32" captionText="FIGURES 119−125. Laminamichaelia furcula sp. nov. 119—Prodorsum, neotrichous soft integument and in-area, sensilla ve branched, button-like naso, reticulated pattern of large lamellae on crista between strips; 120—Dorsal habitus, strong neotrichy; 121—Vental habitus, neotrichy, 9–10 genital setae per valve, 7–8 anal setae per valve; 122—Palp tarsus; 123—Subcapitulum, neotrichy; 124—Genital area, 10 genital setae per valve, 3 pairs of eugenital setae; 125—Tarsus I (3B), famulus I (arrow), Tibia I (3B1P)." figureDoi="http://doi.org/10.5281/zenodo.4412377" httpUri="https://zenodo.org/record/4412377/files/figure.png" pageId="2" pageNumber="303">119</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFB5065ED148CD0CF" box="[1243,1279,1950,1972]" captionStart="FIGURE 133" captionStartId="36.[151,250,1715,1739]" captionTargetBox="[151,1436,181,1692]" captionTargetId="figure-29@36.[151,1436,181,1692]" captionTargetPageId="36" captionText="FIGURE 133. Primary and secondary patterns of cuticle in the tribe Bimichaeliini (Bimichaeliinae). A, B—stellate primary pattern by large lamellae, secondary pattern by small lamellae, setae prominent in Laminamichaelia sp. of arbusculosa-group, C, D—primary pattern by clumps of large lamellae connected by small lamellae, setae prominent in Laminamichaelia sp. of subnuda-group, E, F—primary pattern by roundish cells of large lamellae, secondary pattern by small granulae, setae inconspicuous in Bimichaelia sp. Abbreviations: Ch—chelicera, P—palp, Gen—genitalia, C, D, E, F, H, PS—segments of opisthosoma." figureDoi="http://doi.org/10.5281/zenodo.4412385" httpUri="https://zenodo.org/record/4412385/files/figure.png" pageId="2" pageNumber="303">133</figureCitation>
|
||
A–F); cheliceral seta absent (
|
||
<figureCitation id="BCFE2A29C466EB6EFED365C811C7D0AB" box="[344,436,1979,2001]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">Figs. 108</figureCitation>
|
||
,
|
||
<figureCitation id="BCFE2A29C466EB6EFE3565C81196D0AB" box="[446,485,1979,2000]" captionStart="FIGURES 119−125" captionStartId="32.[151,264,1868,1892]" captionTargetBox="[175,1414,193,1835]" captionTargetId="figure-29@32.[164,1423,181,1844]" captionTargetPageId="32" captionText="FIGURES 119−125. Laminamichaelia furcula sp. nov. 119—Prodorsum, neotrichous soft integument and in-area, sensilla ve branched, button-like naso, reticulated pattern of large lamellae on crista between strips; 120—Dorsal habitus, strong neotrichy; 121—Vental habitus, neotrichy, 9–10 genital setae per valve, 7–8 anal setae per valve; 122—Palp tarsus; 123—Subcapitulum, neotrichy; 124—Genital area, 10 genital setae per valve, 3 pairs of eugenital setae; 125—Tarsus I (3B), famulus I (arrow), Tibia I (3B1P)." figureDoi="http://doi.org/10.5281/zenodo.4412377" httpUri="https://zenodo.org/record/4412377/files/figure.png" pageId="2" pageNumber="303">120</figureCitation>
|
||
;
|
||
<bibRefCitation id="40544B5DC466EB6EFE6465C81209D0AB" author="Uusitalo, M." box="[495,634,1979,2001]" pageId="2" pageNumber="303" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
|
||
: fig. 102); prodorsum without eyes (
|
||
<figureCitation id="BCFE2A29C466EB6EFC5C65C8145AD0AA" box="[983,1065,1979,2001]" captionStart="FIGURES 105−111" captionStartId="28.[151,264,1874,1898]" captionTargetBox="[204,1378,187,1839]" captionTargetId="figure-29@28.[198,1390,181,1850]" captionTargetPageId="28" captionText="FIGURES 105−111. Laminamichaelia shibai nom. nov. 105—Prodorsum, holotrichy, ciliated setae vi, sce and in, barbed sensilla ve, button-like naso, reticulated pattern of large lamellae on crista; 106—Dorsal habitus, strong neotrichy; 107—Vental habitus, neotrichy, 4–5 genital setae per valve, 5 anal setae per valve; 108—Straight chelicera; 109—Subcapitulum; 110—Palp tarsus, solenidion supported by 1 eupathid, other one is free; 111—Genital area of female, 5 genital setae per valve, 2 pairs of eugenital setae." figureDoi="http://doi.org/10.5281/zenodo.4412369" httpUri="https://zenodo.org/record/4412369/files/figure.png" pageId="2" pageNumber="303">Fig. 105</figureCitation>
|
||
)...................
|
||
<taxonomicName id="E3C54D2FC466EB6EFA9165C815EED0AA" authorityName="Womersley" authorityYear="1944" box="[1306,1437,1979,2001]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="2" pageNumber="303" phylum="Arthropoda" rank="tribe" tribe="Bimichaeliini">Bimichaeliini</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |