686 lines
101 KiB
XML
686 lines
101 KiB
XML
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<mods:title id="B88C304B3B8487C02FE5B797B85EA9B0">A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<mods:namePart id="E5BB226323385F1CB7A46FF0404A1207">Sands, Arthur F</mods:namePart>
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<mods:affiliation id="444362C603C8F8D74D29D2A2174CB489">Justus Liebig Univrsity Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart id="BF6DBB9A86D9893588DEBB03FCE8F76A">Gloeer, Peter</mods:namePart>
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<mods:affiliation id="862E4C913322C43F58F432F053401F3B">Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
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<mods:namePart id="3A3666E34694F2F3D284D91D8F297003">Guerlek, Mustafa E</mods:namePart>
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<mods:affiliation id="6E3B644424A4363FFF4D49AA6EA3F126">Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
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<mods:namePart id="763D994F3730E2114968D6E8C6C26673">Albrecht, Christian</mods:namePart>
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<mods:namePart id="705884A5518714D2B5D2F7BE7AEFE487">Neubauer, Thomas A</mods:namePart>
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<mods:affiliation id="5DDA8791CEA174686E0164ABF33AB55F">Justus Liebig University Giessen, Giessen, Germany & Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
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<mods:date id="63DCF002EAD462750265A18CE77BEAE9">2020</mods:date>
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<treatment id="AC6413C1AA685897BCE7FE624ABCEA19" ID-GBIF-Taxon="161939045" LSID="urn:lsid:plazi:treatment:AC6413C1AA685897BCE7FE624ABCEA19" httpUri="http://treatment.plazi.org/id/AC6413C1AA685897BCE7FE624ABCEA19" lastPageNumber="25" pageId="0" pageNumber="25" scope_class="Gastropoda" scope_family="Neritidae" scope_subFamily="Neritininae" scope_superFamily="Neritoidea">
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<subSubSection id="32DCE328D2B8B205C4597047E433F474" pageId="0" pageNumber="25" type="nomenclature">
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<paragraph id="489B267A10FE7FBFCBEACE8E9B7703DA" pageId="0" pageNumber="25">
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<taxonomicName id="AF76581E8D00F4FA4F26EC03CFC47380" ID-CoL="8GQF9" LSID="AC6413C1-AA68-5897-BCE7-FE624ABCEA19" authority="Issel, 1865" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus major" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="major">Theodoxus major Issel, 1865</taxonomicName>
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<figureCitation id="1E463B313BFC1D9A586C52162572BB02" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">Figure 19A -Zb</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C044DDD0644457E983DBC3704A7EBFC2" pageId="0" pageNumber="25" type="reference_group">
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<paragraph id="F7DF12FB5E478DF3CCED6DA983F38BBC" pageId="0" pageNumber="25">
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<taxonomicName id="62680A0958EFEADF34FE89F3B5A9526E" authorityName="Eichwald" authorityYear="1838" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina liturata" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="liturata">Neritina liturata</taxonomicName>
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<bibRefCitation id="F69AB5B96D861EDD58EF1307D5AAD7A2" author="Eichwald, E" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="0" pageNumber="25" pagination="125 - 174" publicationUrl="https://biodiversitylibrary.org/page/41342125" refId="B49" refString="Eichwald, E, 1838. Faunae Caspii Maris primitiae. Bulletin de la Societe Imperiale des Naturalistes de Moscou 11 (2): 125 - 174, https://biodiversitylibrary.org/page/41342125" title="Faunae Caspii Maris primitiae." url="https://biodiversitylibrary.org/page/41342125" volume="11" year="1838">Eichwald 1838</bibRefCitation>
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: 156-157;
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<bibRefCitation id="C23A91D903F41F7D71593CF60BAE7EE1" author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
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: 223-224, pl. 21, figs 24-26;
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<bibRefCitation id="DD40C7802308ADD781A5698659EB3B44" author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
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: 8, pl. 212, fig. 1336 (
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<emphasis id="BEC95AD99C4D2BB6C418DB7E177A1518" bold="true" pageId="0" pageNumber="25">non</emphasis>
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<taxonomicName id="FDCA108A47D271A8FC74136879CDC2AF" authorityName="Eichwald" authorityYear="1838" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina liturata" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="liturata">Neritina liturata</taxonomicName>
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Schultze, 1826).
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</paragraph>
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<paragraph id="FF4263F4B0A7EB8D720F4E0BD6AA4656" pageId="0" pageNumber="25">
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<taxonomicName id="B4231351D986366EA8D397A7E1B2DF7A" authorityName="Afs-" authorityYear="2019" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus schirazensis var. major" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="schirazensis" variety="major">Theodoxus schirazensis var. major</taxonomicName>
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<bibRefCitation id="BB1A7EBF46B950A8472313675AEDEAE5" author="Issel, A" journalOrPublisher="Stamperia Reale, Torino" pageId="0" pageNumber="25" publicationUrl="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" refId="B72" refString="Issel, A, 1865. Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Stamperia Reale, Torino, http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" title="Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note." url="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" year="1865">Issel 1865</bibRefCitation>
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: 24;
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<bibRefCitation id="BF4482CDBCD6BF3B690C7CC3BC1991F3" author="Issel, A" journalOrPublisher="serie" pageId="0" pageNumber="25" pagination="387 - 439" publicationUrl="https://biodiversitylibrary.org/page/31197934" refId="B73" refString="Issel, A, 1866. Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Memorie della Reale accademia delle scienze di Torino, 2. serie 23: 387 - 439, https://biodiversitylibrary.org/page/31197934" title="Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Memorie della Reale accademia delle scienze di Torino, 2." url="https://biodiversitylibrary.org/page/31197934" volume="23" year="1866">Issel 1866</bibRefCitation>
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: 408.
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</paragraph>
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<paragraph id="CFBCB23AEF96820DC1A33839AFF6C1D6" pageId="0" pageNumber="25">
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<taxonomicName id="40ECC8E243D2CCE56D5B6CC012BD456B" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
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Schultzii
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<bibRefCitation id="DA3E56C2B126046CBAFBBB9B18C253EC" author="Grimm, OA" journalOrPublisher="Trudy Aralo-Kaspiyskoy Ekspeditsii" pageId="0" pageNumber="25" pagination="1 - 105" publicationUrl="http://www.cawater-info.net/library/rus/hist/2_grimm.pdf" refId="B68" refString="Grimm, OA, 1877. Kaspiyskoye more i yego fauna. Tetrad' 2. Trudy Aralo-Kaspiyskoy Ekspeditsii 2 (2): 1 - 105, http://www.cawater-info.net/library/rus/hist/2_grimm.pdf" title="Kaspiyskoye more i yego fauna. Tetrad' 2." url="http://www.cawater-info.net/library/rus/hist/2_grimm.pdf" volume="2" year="1877">Grimm 1877</bibRefCitation>
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: 77-78, pl. 7, fig. 5, pl. 8, fig. 16.
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</paragraph>
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<paragraph id="A195F3033B842228425E6F2B878DFC10" pageId="0" pageNumber="25">
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<taxonomicName id="9FA328604464BBEF1295A92A299CE1EF" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
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Schulzii [sic]:
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<bibRefCitation id="E908F54295659A55646FBA9724FAF0F9" author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
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||
: 239-240, pl. 23, figs 13-16.
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</paragraph>
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<paragraph id="A6A7825CFBF362AD6FA685E08F4BB2AF" pageId="0" pageNumber="25">
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<taxonomicName id="02787FD041564258816B8C6360F2B8E5" authorityName="Lindholm" authorityYear="1924" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallasi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallasi">Theodoxus pallasi</taxonomicName>
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<bibRefCitation id="9BEF1B7B209F21929E29A53DA3117A2C" author="Lindholm, VA" journalOrPublisher="Russkiy Gidrobiologicheskiy Zhurnal" pageId="0" pageNumber="25" pagination="32 - 34" refId="B89" refString="Lindholm, VA, 1924. K nomenklature nekotorykh kaspiyskikh gastropod. Russkiy Gidrobiologicheskiy Zhurnal 3 (1-2): 32 - 34" title="K nomenklature nekotorykh kaspiyskikh gastropod." volume="3" year="1924">Lindholm 1924</bibRefCitation>
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: 33, 34 (nom. nov. pro
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<taxonomicName id="18276FBF55528D7D4984BC9ED8EFB0F0" authorityName="Eichwald" authorityYear="1838" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina liturata" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="liturata">Neritina liturata</taxonomicName>
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||
Eichwald, 1838,
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<emphasis id="8B08C6ECC9ACC62A62778F5416FB1DCC" bold="true" pageId="0" pageNumber="25">non</emphasis>
|
||
Schultze, 1826);
|
||
<bibRefCitation id="CE569838D6811086BC74138DFE226C59" author="Starobogatov, YI" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B151" refString="Starobogatov, YI, 1974. Phylum Mollusca. In Mordukhay-Boltovskoy FD (ed.). Atlas bespozvonochnykh Aral'skogo morya. Pishchevaya Promyshlennost', Moscow: pp. 237-257." title="Phylum Mollusca. In Mordukhay-Boltovskoy FD (ed.). Atlas bespozvonochnykh Aral'skogo morya. Pishchevaya Promyshlennost', Moscow: pp. 237 - 257." year="1974">Starobogatov 1974</bibRefCitation>
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||
: 255-256, text fig. 224;
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<bibRefCitation id="1A4D27AF56193596779346A182C05993" author="Akramovskiy, NN" journalOrPublisher="Izvestiya Akademii Nauk Armyanskoy SSR, Yerevan" pageId="0" pageNumber="25" refId="B3" refString="Akramovskiy, NN, 1976. Fauna Armyanskoy SSR. Mollyuski (Mollusca). Izvestiya Akademii Nauk Armyanskoy SSR, Yerevan" title="Fauna Armyanskoy SSR. Mollyuski (Mollusca)." year="1976">Akramovskiy 1976</bibRefCitation>
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||
: 88, text fig. 23, pl. 1, figs 1, 2;
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<bibRefCitation id="40D43C85A12A88A6DC0FF7459563DFD7" DOI="https://doi.org/10.1127/arch.moll/146/213-226" author="Anistratenko, VV" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="213 - 226" refId="B17" refString="Anistratenko, VV, Zettler, ML, Anistratenko, OY, 2017. On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region. Archiv fuer Molluskenkunde 146 (2): 213 - 226, DOI: https://doi.org/10.1127/arch.moll/146/213-226" title="On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region." url="https://doi.org/10.1127/arch.moll/146/213-226" volume="146" year="2017">Anistratenko et al. 2017</bibRefCitation>
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||
: 221, figs 4, 7, 10, 11;
|
||
<bibRefCitation id="93A00C165B60F6C33F931F940549D4DD" DOI="https://doi.org/10.3897/zookeys.770.25365" author="Neubauer, TA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="43 - 103" refId="B113" refString="Neubauer, TA, van de Velde, S, Yanina, T, Wesselingh, FP, 2018. A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys 770: 43 - 103, DOI: https://doi.org/10.3897/zookeys.770.25365" title="A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy." url="https://doi.org/10.3897/zookeys.770.25365" volume="770" year="2018">Neubauer et al. 2018</bibRefCitation>
|
||
: 48-51, figs 4A-F;
|
||
<bibRefCitation id="173DB18E608793471C9039D2F19A2B85" DOI="https://doi.org/10.3897/zookeys.827.31365" author="Wesselingh, FP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="31 - 124" refId="B159" refString="Wesselingh, FP, Neubauer, TA, Anistratenko, VV, Vinarski, MV, Yanina, T, ter Poorten, JJ, Kijashko, P, Albrecht, C, Anistratenko, OY, D'Hont, A, Frolov, P, Gandara, AM, Gittenberger, A, Gogaladze, A, Karpinsky, M, Lattuada, M, Popa, L, Sands, AF, van de Velde, S, Vandendorpe, J, Wilke, T, 2019. Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys 827: 31 - 124, DOI: https://doi.org/10.3897/zookeys.827.31365" title="Mollusc species from the Pontocaspian region - an expert opinion list." url="https://doi.org/10.3897/zookeys.827.31365" volume="827" year="2019">Wesselingh et al. 2019</bibRefCitation>
|
||
: 64-65;
|
||
<bibRefCitation id="8480C6426B07602F3C655FF970535963" author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
|
||
<normalizedToken id="A4C2186227A95E60B5F1F0BDE98D82EE" originalValue="Glöer">Gloeer</normalizedToken>
|
||
2019
|
||
</bibRefCitation>
|
||
: 48-49, fig. 36.
|
||
</paragraph>
|
||
<paragraph id="D74AD3BC38E69B62707735BD022D60BD" pageId="0" pageNumber="25">
|
||
<taxonomicName id="55E9AFCC9AEC2340068227EE35399CEC" authorityName="Kolesnikov" authorityYear="1947" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallasi var. nalivkini" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="pallasi" variety="nalivkini">Theodoxus (Theodoxus) pallasi var. nalivkini</taxonomicName>
|
||
<bibRefCitation id="D2F4E5CD353393E2718B0E2EF3B5E0DD" author="Kolesnikov, VP" journalOrPublisher="Byulleten' Moskovskogo Obshchestva Ispytateley Prirody, otdel geologicheskiy" pageId="0" pageNumber="25" pagination="105 - 112" refId="B84" refString="Kolesnikov, VP, 1947. Tablitsa dlya opredeleniya kaspiyskikh gastropod. Byulleten' Moskovskogo Obshchestva Ispytateley Prirody, otdel geologicheskiy 22 (1): 105 - 112" title="Tablitsa dlya opredeleniya kaspiyskikh gastropod." volume="22" year="1947">Kolesnikov 1947</bibRefCitation>
|
||
: 106, 110.
|
||
</paragraph>
|
||
<paragraph id="4D2A0615384F646DD3CEC2E6074E9085" pageId="0" pageNumber="25">
|
||
<taxonomicName id="F099AB70085D31F030777348705A55E3" baseAuthorityName="Grimm" baseAuthorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus schultzi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="schultzi">Theodoxus (Ninnia) schultzi</taxonomicName>
|
||
[sic]
|
||
<taxonomicName id="17FB28E825FB6935C5C2073BE7085338" authorityName="Kolesnikov" authorityYear="1947" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. jukovi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="schultzi" variety="jukovi">var. jukovi</taxonomicName>
|
||
<bibRefCitation id="B701AED2F8A5A7612CE89D1C016D6CFC" author="Kolesnikov, VP" journalOrPublisher="Byulleten' Moskovskogo Obshchestva Ispytateley Prirody, otdel geologicheskiy" pageId="0" pageNumber="25" pagination="105 - 112" refId="B84" refString="Kolesnikov, VP, 1947. Tablitsa dlya opredeleniya kaspiyskikh gastropod. Byulleten' Moskovskogo Obshchestva Ispytateley Prirody, otdel geologicheskiy 22 (1): 105 - 112" title="Tablitsa dlya opredeleniya kaspiyskikh gastropod." volume="22" year="1947">Kolesnikov 1947</bibRefCitation>
|
||
: 106, 110.
|
||
</paragraph>
|
||
<paragraph id="D2EAA81849662EB83F90FA82731631DA" pageId="0" pageNumber="25">
|
||
<taxonomicName id="1B6842586E7EDE0F459ACA0DAC053BF8" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus zhukovi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="zhukovi">Theodoxus zhukovi</taxonomicName>
|
||
[sic]:
|
||
<bibRefCitation id="83DB91F306B42046C18C9C19AF0F2706" author="Starobogatov, YI" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B151" refString="Starobogatov, YI, 1974. Phylum Mollusca. In Mordukhay-Boltovskoy FD (ed.). Atlas bespozvonochnykh Aral'skogo morya. Pishchevaya Promyshlennost', Moscow: pp. 237-257." title="Phylum Mollusca. In Mordukhay-Boltovskoy FD (ed.). Atlas bespozvonochnykh Aral'skogo morya. Pishchevaya Promyshlennost', Moscow: pp. 237 - 257." year="1974">Starobogatov 1974</bibRefCitation>
|
||
: 255, text fig. 223.
|
||
</paragraph>
|
||
<paragraph id="FE05EF526ECCC39816B26F77043F54A8" pageId="0" pageNumber="25">
|
||
<taxonomicName id="25EE382C91B0A2D0B5A637B3DC173548" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus astrachanicus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="astrachanicus">Theodoxus astrachanicus</taxonomicName>
|
||
Starobogatov in
|
||
<bibRefCitation id="C3D84BCB3DF43B95C90AAB47BD636931" author="Starobogatov, YI" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B152" refString="Starobogatov, YI, Filchakov, VA, Antonova, LA, Pirogov, VV, 1994. Novyye dannyye o mollyuskakh i vysshikh rakoobraznykh delty Volgi. Vestnik Zoologii 4-5: 8-12." title="Novyye dannyye o mollyuskakh i vysshikh rakoobraznykh delty Volgi. Vestnik Zoologii 4 - 5: 8 - 12." year="1994">Starobogatov et al. 1994</bibRefCitation>
|
||
: 8-9, fig. 1(1-2);
|
||
<bibRefCitation id="1E664AAE25105A3BA4B23E661D962C82" author="Degtyarenko, EV" journalOrPublisher="Zbirnyk prats' Zoolohichnoho muzeyu" pageId="0" pageNumber="25" pagination="13 - 57" refId="B44" refString="Degtyarenko, EV, Anistratenko, VV, 2013. Mollyuski kontinental'nykh vodoyemov severo-zapadnogo Priazov'ya: faunisticheskiy obzor s zamechaniyami po rasprostraneniyu i ekologii. Zbirnyk prats' Zoolohichnoho muzeyu 42: 13 - 57" title="Mollyuski kontinental'nykh vodoyemov severo-zapadnogo Priazov'ya: faunisticheskiy obzor s zamechaniyami po rasprostraneniyu i ekologii." volume="42" year="2013">Degtyarenko and Anistratenko 2013</bibRefCitation>
|
||
: 22-23, pl. 1, fig. 2a-c.
|
||
</paragraph>
|
||
<paragraph id="1DD1E9EE734BEC9A8D9631D9AA16F32E" pageId="0" pageNumber="25">
|
||
<taxonomicName id="27FC3F43EF53E73EF57C6ED8E748262F" baseAuthorityName="Grimm" baseAuthorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus schultzii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="schultzii">Theodoxus (Theodoxus) schultzii</taxonomicName>
|
||
:
|
||
<bibRefCitation id="CBD02404AF19CE9BCC421C2139AAD39E" author="Zettler, ML" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="25" pagination="245 - 252" refId="B165" refString="Zettler, ML, 2007. A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea. Journal of Conchology 39 (3): 245 - 252" title="A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea." volume="39" year="2007">Zettler 2007</bibRefCitation>
|
||
: 249-250, figs 4a-h, 5a-h;
|
||
<bibRefCitation id="4DDC2FC0BCE6BD3C7BE6F07BBC904525" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
|
||
: 157.
|
||
</paragraph>
|
||
<paragraph id="5B75459E371D85CC998D41BBB4DA4C32" pageId="0" pageNumber="25">
|
||
?
|
||
<taxonomicName id="4D2A2D9DFDB6CD3F954CDEB77FC611A8" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus doriae" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="doriae">Theodoxus doriae</taxonomicName>
|
||
:
|
||
<bibRefCitation id="9AEF0DB8A25D6F698173501B3EF103D0" author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee 2012</bibRefCitation>
|
||
: 44-45 (partim, only material from Gilan and Mazandaran provinces;
|
||
<emphasis id="F84DF3A1C3EBFD277ECF6DEF5A626573" bold="true" pageId="0" pageNumber="25">non</emphasis>
|
||
<taxonomicName id="8AE0CA1766895E7B71370394771DADA8" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus doriae" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="doriae">Theodoxus doriae</taxonomicName>
|
||
Issel, 1865 [=
|
||
<taxonomicName id="8A26A5E4F712A48FA324A9456C4D7788" class="Gastropoda" family="Neritidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. pallidus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallidus">T. pallidus</taxonomicName>
|
||
])
|
||
</paragraph>
|
||
<paragraph id="55CFA8E77BA38A1B166B403CD3AF95B0" pageId="0" pageNumber="25">
|
||
?
|
||
<taxonomicName id="5DA6DE750F3523B0583B6C87A96217C7" baseAuthorityName="Mousson" baseAuthorityYear="1874" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus euphraticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euphraticus">Theodoxus euphraticus</taxonomicName>
|
||
:
|
||
<bibRefCitation id="C755CB724A6EEB9450111B68F7B9C957" author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee 2012</bibRefCitation>
|
||
: 61 (partim, only material from the Khorasan provinces;
|
||
<emphasis id="F1189AB205D83B484CB2F6EADAAA3EFD" bold="true" pageId="0" pageNumber="25">non</emphasis>
|
||
<taxonomicName id="9AB06781CE74FA0ACA83E5B7DA69F6C0" authorityName="Mousson" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina euphratica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euphratica">Neritina euphratica</taxonomicName>
|
||
Mousson, 1874 [=
|
||
<taxonomicName id="0A549095227AEBF97D2E0AED70F949DF" class="Gastropoda" family="Neritidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">T. jordani</taxonomicName>
|
||
])
|
||
</paragraph>
|
||
<paragraph id="80776C7DE6F4B42C8D4087113D0E789E" pageId="0" pageNumber="25">
|
||
<taxonomicName id="DBEEF0FC30180A9F3AA84B4950953901" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus fluviatilis</taxonomicName>
|
||
:
|
||
<bibRefCitation id="7F7391F1E94B17706A21279A92B2E1F0" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
|
||
<normalizedToken id="2FEAEDD69FC665AE2E7AA87F3EABE78B" originalValue="Glöer">Gloeer</normalizedToken>
|
||
and
|
||
<normalizedToken id="DB3B5FDE331A55F21F6A79E2C975F3B9" originalValue="Pešić">Pesic</normalizedToken>
|
||
2012
|
||
</bibRefCitation>
|
||
: 16-17 (partim, only Khorrasan material;
|
||
<emphasis id="AB6B16F49BAC09ABC1C254D815D475D4" bold="true" pageId="0" pageNumber="25">non</emphasis>
|
||
<taxonomicName id="23376A9D0A77CA19A0E66F8EB2C740D6" authorityName="Linnaeus" authorityYear="1758" class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Nerita fluviatilis</taxonomicName>
|
||
Linnaeus, 1758).
|
||
</paragraph>
|
||
<paragraph id="7871A400FBB57781E319158052C19C9B" pageId="0" pageNumber="25">
|
||
<taxonomicName id="196EDC54A535360AD5FFA27809324DE4" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus astrachanicus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="astrachanicus">Theodoxus (Theodoxus) astrachanicus</taxonomicName>
|
||
:
|
||
<bibRefCitation id="E9DF7403FEF20C7B6B8CD6D553987CDF" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
|
||
: 155-156.
|
||
</paragraph>
|
||
<paragraph id="1621BD856122F4B7207B46FF956900BC" pageId="0" pageNumber="25">
|
||
<taxonomicName id="0362F88259CD20F4B45E25E0478A5815" authorityName="Lindholm" authorityYear="1924" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallasi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallasi">Theodoxus (Theodoxus) pallasi</taxonomicName>
|
||
:
|
||
<bibRefCitation id="22EE0FD00D6193517369291E39076D9A" author="Vinarski, MV" journalOrPublisher="Severtsov Institute of Ecology and Evolution of RAS, Moscow" pageId="0" pageNumber="25" refId="B157" refString="Vinarski, MV, Kantor, YI, 2016. Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A.N. Severtsov Institute of Ecology and Evolution of RAS, Moscow" title="Analytical catalogue of fresh and brackish water molluscs of Russia and adjacent countries. A. N." year="2016">Vinarski and Kantor 2016</bibRefCitation>
|
||
: 156-157 (and synonyms therein).
|
||
</paragraph>
|
||
<paragraph id="885263F4BFACB5F360FE79E83BD1BF5B" pageId="0" pageNumber="25">
|
||
<taxonomicName id="6F57FF9B3305C9231E4F7F666D930334" baseAuthorityName="Grimm" baseAuthorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus schultzii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="schultzii">Theodoxus schultzii</taxonomicName>
|
||
:
|
||
<bibRefCitation id="E678140B8F4C7BC25C1BF0D6230A5675" DOI="https://doi.org/10.3897/zookeys.827.31365" author="Wesselingh, FP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="31 - 124" refId="B159" refString="Wesselingh, FP, Neubauer, TA, Anistratenko, VV, Vinarski, MV, Yanina, T, ter Poorten, JJ, Kijashko, P, Albrecht, C, Anistratenko, OY, D'Hont, A, Frolov, P, Gandara, AM, Gittenberger, A, Gogaladze, A, Karpinsky, M, Lattuada, M, Popa, L, Sands, AF, van de Velde, S, Vandendorpe, J, Wilke, T, 2019. Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys 827: 31 - 124, DOI: https://doi.org/10.3897/zookeys.827.31365" title="Mollusc species from the Pontocaspian region - an expert opinion list." url="https://doi.org/10.3897/zookeys.827.31365" volume="827" year="2019">Wesselingh et al. 2019</bibRefCitation>
|
||
: 66;
|
||
<bibRefCitation id="2753B1B28E0FF621BDCE630593528F0B" author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
|
||
<normalizedToken id="D14A1DD730527CEC7A521E0E7944CB89" originalValue="Glöer">Gloeer</normalizedToken>
|
||
2019
|
||
</bibRefCitation>
|
||
: 52, fig. 40.
|
||
</paragraph>
|
||
<paragraph id="726307B7EB57AF7B6E9DE1BF30AC73CF" pageId="0" pageNumber="25">
|
||
<taxonomicName id="AB2F2825573C830A17B943B0C0E05073" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus major" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="major">Theodoxus major</taxonomicName>
|
||
:
|
||
<bibRefCitation id="3E352AA745ACA67D494F6EE04032AE12" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
|
||
: 3, fig. 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C6B3DE03A0227B699E0916AF7447A411" pageId="0" pageNumber="25" type="type locality">
|
||
<paragraph id="FF1DFA3184800CDD27F230D97BAFC246" pageId="0" pageNumber="25">
|
||
<typeStatus id="C2527053386602F3A9296550DAF44CE2">Type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="425F031D121C7310B319CC2C1C2389C1" pageId="0" pageNumber="25">
|
||
Lake Sevan, Armenia. However,
|
||
<bibRefCitation id="98B3B60812CE4A98DBA4DA5CF57DADAB" author="Akramovskiy, NN" journalOrPublisher="Izvestiya Akademii Nauk Armyanskoy SSR, Yerevan" pageId="0" pageNumber="25" refId="B3" refString="Akramovskiy, NN, 1976. Fauna Armyanskoy SSR. Mollyuski (Mollusca). Izvestiya Akademii Nauk Armyanskoy SSR, Yerevan" title="Fauna Armyanskoy SSR. Mollyuski (Mollusca)." year="1976">Akramovskiy (1976)</bibRefCitation>
|
||
suggested the
|
||
<typeStatus id="01C3B0591CD09A916700E1CD622D25CE">type</typeStatus>
|
||
locality given by
|
||
<bibRefCitation id="6D7D154B9B3E20E00CC3CBE629D180C6" author="Issel, A" journalOrPublisher="Stamperia Reale, Torino" pageId="0" pageNumber="25" publicationUrl="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" refId="B72" refString="Issel, A, 1865. Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Stamperia Reale, Torino, http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" title="Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note." url="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" year="1865">Issel (1865)</bibRefCitation>
|
||
was erroneous and should be Yerevan in Armenia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B9320D904E828B4FA149F21D8351C92F" pageId="0" pageNumber="25" type="type material">
|
||
<paragraph id="A29BFCDB88907376EA47EA5DD19A903B" pageId="0" pageNumber="25">
|
||
<typeStatus id="A42EFBE6C83C5DC0F161840C7DA51D82">Type</typeStatus>
|
||
material.
|
||
</paragraph>
|
||
<paragraph id="0ECD62CDE14353D7583408830A5789CD" pageId="0" pageNumber="25">
|
||
<materialsCitation id="239E214F85BA40AE99FF3897FB71CB9B" collectionCode="MRSN" country="Turkey" typeStatus="syntypes">
|
||
The
|
||
<typeStatus id="26C1A0F6DD7FB23343A9DFF4EE4D134D">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="5391A90E2FF83A384373921F93491B78" lsidName="T. schirazensis var. major" pageId="0" pageNumber="25" rank="variety" species="schirazensis" variety="major">T. schirazensis var. major</taxonomicName>
|
||
are supposed to be stored in
|
||
<collectionCode id="3C0A94369DEFB216C097526F7E929393" collectionName="Italy, Torino, Museo Regionale di Scienze Naturali" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15721" name="Museo Regionale di Scienze Naturali" type="Herbarium">MRSN</collectionCode>
|
||
, but the collection is inaccessible at the moment due to renovation and restoration of the museum (E. Gavetti pers. comm. 09/2019)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="1EFEDF554888466FA37BC1A17FB96F17" collectionCode="ZIN" country="Turkey" typeStatus="lectotype">
|
||
The
|
||
<typeStatus id="A572876913D7C7495107474E3CA066C6">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="1D271E46C4B301FACF5D9E0337650C80" lsidName="N. liturata" pageId="0" pageNumber="25" rank="species" species="liturata">
|
||
<emphasis id="B3C16CA605EB83FADFE316EBF403AA5D" italics="true" pageId="0" pageNumber="25">N. liturata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="83DD074CAB35D610DF043756833D1217" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="9492303D0CAFBDE692F40A8849C4ED9F" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
(designated by
|
||
<bibRefCitation id="9EA4C659A6E01E1D0206F18DB1F19EF5" author="Starobogatov, YI" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B152" refString="Starobogatov, YI, Filchakov, VA, Antonova, LA, Pirogov, VV, 1994. Novyye dannyye o mollyuskakh i vysshikh rakoobraznykh delty Volgi. Vestnik Zoologii 4-5: 8-12." title="Novyye dannyye o mollyuskakh i vysshikh rakoobraznykh delty Volgi. Vestnik Zoologii 4 - 5: 8 - 12." year="1994">Starobogatov et al. 1994</bibRefCitation>
|
||
) is stored in
|
||
<collectionCode id="9CFC66BEAA04E21B798B618F7B8A5811" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
(coll. no. 54547/63)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3ED24B77DDA40AEE439792DEC2176873" collectionCode="ZIN" country="Turkey" typeStatus="syntypes">
|
||
Twenty
|
||
<typeStatus id="37D00E6F319D6F6CECBBBCBABBF927B4">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="0FBEAE6BA37F59FBBF26FDEAA14A0E4F" lsidName="N. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
|
||
<emphasis id="AC7E63E9C64392FA38D35FF82658ECB4" italics="true" pageId="0" pageNumber="25">N. schultzii</emphasis>
|
||
</taxonomicName>
|
||
are stored in
|
||
<collectionCode id="A6208E7BB3C2625870588E6AE8C9A532" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
(coll. no. 6214/5) and a single in
|
||
<collectionCode id="F388E87D8A7F41C955DD65E3CF757722" collectionName="ZMD" country="Ukraine" httpUri="http://grbio.org/cool/x71v-fhm9" name="The Benedict Dybowski Zoological Museum" type="Museum">ZMD</collectionCode>
|
||
(coll. no. ZB-M W. Dyb. 71; see
|
||
<bibRefCitation id="94EC7550B9F7FD0FC86C98354A3B0AD8" DOI="https://doi.org/10.1127/arch.moll/147/223-236" author="Anistratenko, VV" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="223 - 236" refId="B16" refString="Anistratenko, VV, Anistratenko, OY, Shydlovskyy, I, 2018. Karl E. von Baer's collection of Caspian Sea molluscs stored in the Zoological Museum of Lviv University, Ukraine. Part 1. Catalogue and general description. Archiv fuer Molluskenkunde 147 (2): 223 - 236, DOI: https://doi.org/10.1127/arch.moll/147/223-236" title="Karl E. von Baer's collection of Caspian Sea molluscs stored in the Zoological Museum of Lviv University, Ukraine. Part 1. Catalogue and general description." url="https://doi.org/10.1127/arch.moll/147/223-236" volume="147" year="2018">Anistratenko et al. 2018</bibRefCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BB75F3147F2FAA5FB7DD0DC274D24674" pageId="0" pageNumber="25" type="remarks">
|
||
<paragraph id="6948F5D87F2BF8A6D1DB9F28B3267F3A" pageId="0" pageNumber="25">Remarks.</paragraph>
|
||
<paragraph id="9B13862DEC4E9311948E4ABB7AC2990D" pageId="0" pageNumber="25">
|
||
<bibRefCitation id="A5EB2B4B4311C5457B8F2EC5BCE96915" DOI="https://doi.org/10.3897/zookeys.827.31365" author="Wesselingh, FP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="31 - 124" refId="B159" refString="Wesselingh, FP, Neubauer, TA, Anistratenko, VV, Vinarski, MV, Yanina, T, ter Poorten, JJ, Kijashko, P, Albrecht, C, Anistratenko, OY, D'Hont, A, Frolov, P, Gandara, AM, Gittenberger, A, Gogaladze, A, Karpinsky, M, Lattuada, M, Popa, L, Sands, AF, van de Velde, S, Vandendorpe, J, Wilke, T, 2019. Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys 827: 31 - 124, DOI: https://doi.org/10.3897/zookeys.827.31365" title="Mollusc species from the Pontocaspian region - an expert opinion list." url="https://doi.org/10.3897/zookeys.827.31365" volume="827" year="2019">Wesselingh et al. (2019)</bibRefCitation>
|
||
reviewed the taxonomy of the Pontocaspian
|
||
<taxonomicName id="4B8ECF67E339F9301C19768B889EFA49" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="BCB0E24A944801C19B900E240D58389F" italics="true" pageId="0" pageNumber="25">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
spp. They showed that
|
||
<taxonomicName id="B7A13E32325C3EA7B592EFE74220E493" lsidName="N. liturata" pageId="0" pageNumber="25" rank="species" species="liturata">
|
||
<emphasis id="9C92F5C06AB4DECE10AC60FD5EC26742" italics="true" pageId="0" pageNumber="25">N. liturata</emphasis>
|
||
</taxonomicName>
|
||
Eichwald, 1838 (described from Derbent, Dagestan, Russia, north-western Caspian Sea) was invalid as a junior primary homonym of
|
||
<taxonomicName id="C203B04111BCE3584C09E8787451CEE6" lsidName="N. liturata" pageId="0" pageNumber="25" rank="species" species="liturata">
|
||
<emphasis id="14BC2342C32CA9AD967CD7F5296DC2B8" italics="true" pageId="0" pageNumber="25">N. liturata</emphasis>
|
||
</taxonomicName>
|
||
Schultze, 1826 (described from the Americas) and was replaced by
|
||
<bibRefCitation id="347A4C932DB172571C8C02B44FB2358A" author="Lindholm, VA" journalOrPublisher="Russkiy Gidrobiologicheskiy Zhurnal" pageId="0" pageNumber="25" pagination="32 - 34" refId="B89" refString="Lindholm, VA, 1924. K nomenklature nekotorykh kaspiyskikh gastropod. Russkiy Gidrobiologicheskiy Zhurnal 3 (1-2): 32 - 34" title="K nomenklature nekotorykh kaspiyskikh gastropod." volume="3" year="1924">Lindholm (1924)</bibRefCitation>
|
||
with
|
||
<taxonomicName id="DC5360A7CFCF9245D7D5692AC3B9BDB0" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="4ACE5C0C4F7F2EB910BB274609158D83" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
Lindholm, 1924. Moreover, they supported the synonymy of
|
||
<taxonomicName id="B4C38B25E4323F2413D0D8BCEA1F9CB0" lsidName="T. astrachanicus" pageId="0" pageNumber="25" rank="species" species="astrachanicus">
|
||
<emphasis id="C2E39F0E927DAA2562E8882B9CBF1921" italics="true" pageId="0" pageNumber="25">T. astrachanicus</emphasis>
|
||
</taxonomicName>
|
||
Starobogatov in
|
||
<bibRefCitation id="86C67D75D955AC47E211C2D56EE963BA" author="Starobogatov, YI" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B152" refString="Starobogatov, YI, Filchakov, VA, Antonova, LA, Pirogov, VV, 1994. Novyye dannyye o mollyuskakh i vysshikh rakoobraznykh delty Volgi. Vestnik Zoologii 4-5: 8-12." title="Novyye dannyye o mollyuskakh i vysshikh rakoobraznykh delty Volgi. Vestnik Zoologii 4 - 5: 8 - 12." year="1994">Starobogatov et al. 1994</bibRefCitation>
|
||
with
|
||
<taxonomicName id="C47633EC17E8355E3669DB05DF3C5F15" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="A901DE828C67D12EC2FBFAFA3F642D3D" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="D86D18C4481FA764FCC1A7650D542115" DOI="https://doi.org/10.1127/arch.moll/146/213-226" author="Anistratenko, VV" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="213 - 226" refId="B17" refString="Anistratenko, VV, Zettler, ML, Anistratenko, OY, 2017. On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region. Archiv fuer Molluskenkunde 146 (2): 213 - 226, DOI: https://doi.org/10.1127/arch.moll/146/213-226" title="On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region." url="https://doi.org/10.1127/arch.moll/146/213-226" volume="146" year="2017">Anistratenko et al. 2017</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="145C25CB2BCD475264B7DA8DF55E4422" DOI="https://doi.org/10.3897/zookeys.827.31365" author="Wesselingh, FP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="31 - 124" refId="B159" refString="Wesselingh, FP, Neubauer, TA, Anistratenko, VV, Vinarski, MV, Yanina, T, ter Poorten, JJ, Kijashko, P, Albrecht, C, Anistratenko, OY, D'Hont, A, Frolov, P, Gandara, AM, Gittenberger, A, Gogaladze, A, Karpinsky, M, Lattuada, M, Popa, L, Sands, AF, van de Velde, S, Vandendorpe, J, Wilke, T, 2019. Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys 827: 31 - 124, DOI: https://doi.org/10.3897/zookeys.827.31365" title="Mollusc species from the Pontocaspian region - an expert opinion list." url="https://doi.org/10.3897/zookeys.827.31365" volume="827" year="2019">Wesselingh et al. (2019)</bibRefCitation>
|
||
additionally suggested
|
||
<taxonomicName id="3F3EA6549703660CCEBB76D4155C4607" lsidName="T. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
|
||
<emphasis id="72E0A66824DD98973F9EACB671D8FA58" italics="true" pageId="0" pageNumber="25">T. schultzii</emphasis>
|
||
</taxonomicName>
|
||
(Grimm, 1877) and
|
||
<taxonomicName id="4C3483B83E08CD4354082B658477B05A" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="1815A0F266CEDE17337858C6319F16A4" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
(the latter originally described as a variety of the unavailable name
|
||
<taxonomicName id="701AB6EA8450D30D9290FA0BCEFB8EDB" lsidName="T. schirazensis" pageId="0" pageNumber="25" rank="species" species="schirazensis">
|
||
<emphasis id="5EA62E412BB010C9546C0E7D751B6CD4" italics="true" pageId="0" pageNumber="25">T. schirazensis</emphasis>
|
||
</taxonomicName>
|
||
) may additionally be conspecific with
|
||
<taxonomicName id="35F98C654C77FA6E1888C3D185E9F207" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="C898F5951255CCFC1BDD06FFAFCA2E74" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
given morphological similarities. However,
|
||
<bibRefCitation id="61B718B07A8C8C3B6C9426AC115EB79A" DOI="https://doi.org/10.3897/zookeys.827.31365" author="Wesselingh, FP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="31 - 124" refId="B159" refString="Wesselingh, FP, Neubauer, TA, Anistratenko, VV, Vinarski, MV, Yanina, T, ter Poorten, JJ, Kijashko, P, Albrecht, C, Anistratenko, OY, D'Hont, A, Frolov, P, Gandara, AM, Gittenberger, A, Gogaladze, A, Karpinsky, M, Lattuada, M, Popa, L, Sands, AF, van de Velde, S, Vandendorpe, J, Wilke, T, 2019. Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys 827: 31 - 124, DOI: https://doi.org/10.3897/zookeys.827.31365" title="Mollusc species from the Pontocaspian region - an expert opinion list." url="https://doi.org/10.3897/zookeys.827.31365" volume="827" year="2019">Wesselingh et al. (2019)</bibRefCitation>
|
||
refrained from synonymising
|
||
<taxonomicName id="32BD9E43CCCC83B55CA4B0562861B1F9" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="E3A6B51081EDBDD8CA59A312135B1380" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C76DCCFDB6385BD0764BD283901FE3F7" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="2A726E0A85F7C501AC6F85E6B5A0CF83" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="386A50EB02FA9325EA35F5DC003583A1" lsidName="T. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
|
||
<emphasis id="18DE557445AF0FC3FE0E4589B35F6361" italics="true" pageId="0" pageNumber="25">T. schultzii</emphasis>
|
||
</taxonomicName>
|
||
, pending molecular support. This was provided recently by
|
||
<bibRefCitation id="CD496C3A790A95F02524BAC548B9F0FE" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. (2019b)</bibRefCitation>
|
||
, who corroborated the synonymy of
|
||
<taxonomicName id="63E83FD40CD8585BC8663F85FB12E7E5" lsidName="T. astrachanicus" pageId="0" pageNumber="25" rank="species" species="astrachanicus">
|
||
<emphasis id="28484D2BF792EBA4A000ECD6310EA00D" italics="true" pageId="0" pageNumber="25">T. astrachanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="579B3E9210B330C374D775A8EF527C58" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="2B40C028FEC2BA924DA2B5C168245545" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="966EA4811E75AD5383ABE7D3FE812323" lsidName="T. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
|
||
<emphasis id="A0264EA918D393A1AD7AAC72595ECEFA" italics="true" pageId="0" pageNumber="25">T. schultzii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="EAFC7B192F281CCF1FA5C9B8B54A84B0" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="EA585BD813D2890A2562D8EE61763041" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
(also see Fig.
|
||
<figureCitation id="8CB2EE4D0CD780A55EC4DB11995AE050" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). As already argued by
|
||
<bibRefCitation id="D0466B90E594052A21C04D53DA4D0B7E" DOI="https://doi.org/10.3897/zookeys.827.31365" author="Wesselingh, FP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="31 - 124" refId="B159" refString="Wesselingh, FP, Neubauer, TA, Anistratenko, VV, Vinarski, MV, Yanina, T, ter Poorten, JJ, Kijashko, P, Albrecht, C, Anistratenko, OY, D'Hont, A, Frolov, P, Gandara, AM, Gittenberger, A, Gogaladze, A, Karpinsky, M, Lattuada, M, Popa, L, Sands, AF, van de Velde, S, Vandendorpe, J, Wilke, T, 2019. Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys 827: 31 - 124, DOI: https://doi.org/10.3897/zookeys.827.31365" title="Mollusc species from the Pontocaspian region - an expert opinion list." url="https://doi.org/10.3897/zookeys.827.31365" volume="827" year="2019">Wesselingh et al. (2019)</bibRefCitation>
|
||
, the name
|
||
<taxonomicName id="B1AB93C782B207BC6F7FC3C3E694C87C" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="4A6270EE832667B7E48587368401717E" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
has priority.
|
||
</paragraph>
|
||
<paragraph id="CBFA96A28AB79EAF0E8605948081D658" pageId="0" pageNumber="25">
|
||
There is large intraspecific morphological variability within this species, especially regarding the radula, shell shape and periostracum colouration (
|
||
<bibRefCitation id="F0CCA3FC411D5ED4362342927DF5999E" author="Zettler, ML" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="25" pagination="245 - 252" refId="B165" refString="Zettler, ML, 2007. A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea. Journal of Conchology 39 (3): 245 - 252" title="A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea." volume="39" year="2007">Zettler 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4B6BE7CDFF06C1A034C2A7FE0C81AD8B" DOI="https://doi.org/10.1127/arch.moll/146/213-226" author="Anistratenko, VV" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="213 - 226" refId="B17" refString="Anistratenko, VV, Zettler, ML, Anistratenko, OY, 2017. On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region. Archiv fuer Molluskenkunde 146 (2): 213 - 226, DOI: https://doi.org/10.1127/arch.moll/146/213-226" title="On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region." url="https://doi.org/10.1127/arch.moll/146/213-226" volume="146" year="2017">Anistratenko et al. 2017</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="5D1CC238EB93155B4C5A6D92AC069C1E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">19</figureCitation>
|
||
). For example, specimens from deeper parts of the Caspian Sea (
|
||
<taxonomicName id="7B8F388AF924450BA91E94E6BC720F71" lsidName="T. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
|
||
<emphasis id="ED3FD57ABD468FD5BB2C7C540945CC64" italics="true" pageId="0" pageNumber="25">T. schultzii</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) may lack colouration and have a broadened columellar plate making the shell look somewhat flattened (
|
||
<bibRefCitation id="46E2226DDF60E213284CA0559A80B6AA" author="Zettler, ML" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="25" pagination="245 - 252" refId="B165" refString="Zettler, ML, 2007. A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea. Journal of Conchology 39 (3): 245 - 252" title="A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea." volume="39" year="2007">Zettler 2007</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="8863C120CDC4C4B0852C06E89ACC457D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">19</figureCitation>
|
||
), while shoreline samples (
|
||
<taxonomicName id="ACF7327A3E2A699C531260185CB948BA" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="C7E5A6FE2B97DAD1A3708D5A75C30148" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) lack this broadened columellar plate and display clear, black diagonal banding patterning (Fig.
|
||
<figureCitation id="38341583E356DF53008E92EBE8474853" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">19</figureCitation>
|
||
). Inland specimens (
|
||
<taxonomicName id="EC227DF63B8A5667681946D79008808B" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="ADE286B96C0575C300F9485E60C2C076" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) from Iran and Armenia tend to be dark and with a less distinct banding pattern (Fig.
|
||
<figureCitation id="AA78DC5B4FA35F1AA59FE3A945296CD4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">19</figureCitation>
|
||
). The operculum of this species is somewhat similar to those of
|
||
<taxonomicName id="15D2961CCBF53613F5C3AC4C8A67AAF4" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
|
||
<emphasis id="11853C8A47191609B4F184CFD540506E" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C66500716859452F9BAFBA778E26966B" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis id="C6BA094A05B1E20D34BEF44C098AB48E" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2325DE77633233F4006E4CA32D44AD12" lsidName="T. velox" pageId="0" pageNumber="25" rank="species" species="velox">
|
||
<emphasis id="38817F7AE5D001AF9E41C3E2FFEB7886" italics="true" pageId="0" pageNumber="25">T. velox</emphasis>
|
||
</taxonomicName>
|
||
with a strong rib-shield and no pseudo-apophysis (Figs
|
||
<figureCitation id="40F1060761CD87A59C54C72D1E9DEA59" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen collected close to Oued Laabid, Morocco (UGSB 18106) (Fig. 2); E - H. Specimen from the Danube River, Woerth an der Donau, Germany (UGSB 24173); I - L. Specimen (T. euxinus - morphotype) collected in the Papuc Creek, Kiyikoey, Turkey (UGSB 24172) (Fig. 2); M - P. Specimen (T. euxinus - morphotype) from Ovidiopol, Ukraine (UGSB 18124); Q - S. Specimen (T. heldreichi - morphotype) collected in Lake Egirdir, Turkey (UGSB 20361); T, U. Specimen (T. danasteri - morphotype) collected in Krasna Kosa, Ukraine (UGSB 18417) (Fig. 2); V. Opercula of a T. danasteri - morphotype from Lake Skadar, Montenegro (UGSB 24171). Specimens A-D, I-L and T-U were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure9" httpUri="https://binary.pensoft.net/fig/376458" pageId="0" pageNumber="25">9</figureCitation>
|
||
,
|
||
<figureCitation id="A3C0C412141B0A52904E04E003896DB5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E - H. Specimen (T. subthermalis - morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I - K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL- 111736); L. Specimen (T. saulcyi - morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25">10</figureCitation>
|
||
,
|
||
<figureCitation id="E20F791E23195632691FC5DC34824738" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">19</figureCitation>
|
||
,
|
||
<figureCitation id="838962BD70DA5D662B3AECA3E87E6C16" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Theodoxus pallidus (Dunker, 1861). A - D. Specimen from Haji Abad, Fars, Iran (UGSB 22227); E - H. Syntype of N. pallida from southern Iran (ZMB 108.792); I, J; K, L. Two syntypes of N. pallida from southern Iran (ZMB 108.793); M - P. Specimen (T. doriae - morphotype) from a spring near Harat, Yazd, Iran (UGSB 21706); Q, R; S, T; U, V; W - Y. Four syntypes of N. doriae from the thermal springs of Kerman, Iran (MSNG, without coll. no.). Specimens A-D and M-P were used in the phylogeny. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure20" httpUri="https://binary.pensoft.net/fig/376469" pageId="0" pageNumber="25">20</figureCitation>
|
||
,
|
||
<figureCitation id="ECEC0D192F9CF3B6F87CC2F988F2BBA8" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Theodoxus velox V. Anistratenko in O. Anistratenko et al., 1999. A - D. Specimen from the Dnieper River, Kiev, Ukraine (UGSB 24186); E - H. Specimen collected in the Danube delta, Vylkove, Ukraine (UGSB 24184) used in the phylogeny (Fig. 2); I, J. Holotype from Dnieper Delta, Zbur'ivka Liman, Ukraine (IZAN); K - M. Specimen from Lake Sapanca, Yeni Esme, Turkey (UGSB 20691). Theodoxus sarmaticus (Lindholm, 1901). N - P; Q - S; T - U. Three syntypes from the Oskol River, near Golubino, Russia stored in ZIN (coll. no. 6099 / 24). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure22" httpUri="https://binary.pensoft.net/fig/376471" pageId="0" pageNumber="25">22</figureCitation>
|
||
). It is not surprising that molecular data of material from locations in Khorrasan province studied by
|
||
<bibRefCitation id="C031DACC86A6FDFBA7924F073F3847D3" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
|
||
<normalizedToken id="A012B212AD446908133AB243FC58822D" originalValue="Glöer">Gloeer</normalizedToken>
|
||
and
|
||
<normalizedToken id="7B1C0F0006D4495DF0554BE924A194C8" originalValue="Pešić">Pesic</normalizedToken>
|
||
(2012)
|
||
</bibRefCitation>
|
||
and identified as
|
||
<taxonomicName id="01535B5E92A48C66DA4C66A7BE88DE4A" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis id="F00060C990552FCBC10CFCAD757AD993" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
conforms to
|
||
<taxonomicName id="21ACF0AFBE608D2FAA99AAFCC9D0EE87" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="15A8A396743B46DA9F0B026C0542F3E1" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EA475A40A0CAD3A4801AFD6E1E353C01" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="EA52918F5FE2301FBA3605A699EB7138" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). Akramovskiy (1971) already suggested that specimens considered
|
||
<taxonomicName id="B09E1A05814E2FD2231849900928A0B4" lsidName="T. subthermalis" pageId="0" pageNumber="25" rank="species" species="subthermalis">
|
||
<emphasis id="CF23897904BB8BFDC69EC6D72EEFB8D5" italics="true" pageId="0" pageNumber="25">T. subthermalis</emphasis>
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="65C2A4C0278DA870449CD09D63211C9C" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis id="F87FEE495D47C3B56995EE6020D1A3B7" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
] from Armenia represented black-coloured morphotypes of
|
||
<taxonomicName id="149CCA543CF8C2E7CB0AA8CA0583D2D3" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="443E867411361A865E2EE01FB342257D" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="929D92A49B10734DEDAA4C3C1C2E1F1D" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="1A5BDDBE1F1C78485BE69F815FB0D744" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
]. However,
|
||
<taxonomicName id="DD5F08B9D55C0372DB79FEC4FE693F4B" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="57B8A3E40B77E916BEE3624C3DCEEA61" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
only shares a close phylogenetic relationship with
|
||
<taxonomicName id="245D4790276835B36DFB9D46BB0389B3" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
|
||
<emphasis id="DB8F522E5873E6F23A88C4C7F5773197" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
|
||
</taxonomicName>
|
||
and likely diverged from their common ancestor during the Pleistocene (
|
||
<bibRefCitation id="B5EA5DF206AC78553D015AE549914F53" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="93A2B3DD8BC8DCD8AD106A255D7468AB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="D06EB4A3625B8E6FD5180A28EC1A6E73" doi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25" start="Figure 19" startId="F19">
|
||
<paragraph id="04335F88AFCBD2EFC30F18CBFB97CC36" pageId="0" pageNumber="25">
|
||
<emphasis id="770E78229EF1A3D383DB1E71BBEF88E4" bold="true" pageId="0" pageNumber="25">Figure 19.</emphasis>
|
||
<taxonomicName id="88A2D8DD11351EDF1189E66E16458355" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus major" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="major">
|
||
<emphasis id="5F7CE2DF7C1B68EFA3BE56AD6C049B03" italics="true" pageId="0" pageNumber="25">Theodoxus major</emphasis>
|
||
</taxonomicName>
|
||
Issel, 1865.
|
||
<emphasis id="912DAE21DA0CE923060B497D1EE7397E" bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
,
|
||
<emphasis id="F639A547383CDB7B8B9E92799C702A48" bold="true" pageId="0" pageNumber="25">B</emphasis>
|
||
. Specimen (
|
||
<taxonomicName id="33B29DD2535EEC6E3425B22343A30995" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="E1D0CC59B681B9B2E1C861DD280C3DB9" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (
|
||
<collectionCode id="DEAEA19106B0F9C1943C8EAC58BDAB84" collectionName="UGSB">UGSB</collectionCode>
|
||
20712);
|
||
<emphasis id="1D0C93B21365D6C0B0F0D8318CE44142" bold="true" pageId="0" pageNumber="25">C</emphasis>
|
||
,
|
||
<emphasis id="69F72717FFECCD868CA76886382C7CD0" bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
. Specimen (
|
||
<taxonomicName id="5F1F50BF888966359DEBC034D154C314" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="871586DC02C35C4AC476388925EE638E" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) collected in the Aral Sea, Kazakhstan in 1979 (
|
||
<collectionCode id="216B8C9FA43B688F73D2D6E57B02C3F2" collectionName="UGSB">UGSB</collectionCode>
|
||
19129);
|
||
<emphasis id="46DBC515082FFFD617F21C5A7257D9B8" bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
-
|
||
<emphasis id="A4F283739BD3FF567FFDCD3CD6F2283E" bold="true" pageId="0" pageNumber="25">H</emphasis>
|
||
. Specimen (
|
||
<taxonomicName id="B95D40CD704E108FD79D1B6BDD2D9EE9" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="5E9310BBE1A12678C2C85E384776D230" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (
|
||
<collectionCode id="B94CF59FF1D1773495A00A00318DF291" collectionName="UGSB">UGSB</collectionCode>
|
||
18091);
|
||
<emphasis id="B75D62F91E5BD62D2E8911E166C1D3E3" bold="true" pageId="0" pageNumber="25">I</emphasis>
|
||
-
|
||
<emphasis id="5CC05133322AF93DFDBF789EB4373354" bold="true" pageId="0" pageNumber="25">K</emphasis>
|
||
. Specimen of
|
||
<taxonomicName id="6A35BD32FC5DEF85D5562C58BA569E5F" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="11EBE169BADE308F212D343E7968EF7B" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
collected at the outflow of Lake Akna, Aknalich, Armenia (
|
||
<collectionCode id="BDD15A04F6702718DD5EE6952E158537" collectionName="UGSB">UGSB</collectionCode>
|
||
20482);
|
||
<emphasis id="BA48296FFB488C0B1E5E9604CD826A3E" bold="true" pageId="0" pageNumber="25">L</emphasis>
|
||
-
|
||
<emphasis id="1A6896AD0AD4F381D7F5F071671183E4" bold="true" pageId="0" pageNumber="25">N</emphasis>
|
||
. Specimen (
|
||
<taxonomicName id="E291CF017FEC449B996630DE1E38357F" lsidName="T. astrachanicus" pageId="0" pageNumber="25" rank="species" species="astrachanicus">
|
||
<emphasis id="A3DFFFC9D539F1F538EDE2D267956AE6" italics="true" pageId="0" pageNumber="25">T. astrachanicus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) collected in Utlyukskij Liman, Ukraine (
|
||
<collectionCode id="580575D967440BD0B8923382F46BF940" collectionName="UGSB">UGSB</collectionCode>
|
||
18130);
|
||
<emphasis id="C19BCAEE0E13903104D828A9E9CEC921" bold="true" pageId="0" pageNumber="25">O</emphasis>
|
||
,
|
||
<emphasis id="C9C2A74044206AA0DE7BA70492040551" bold="true" pageId="0" pageNumber="25">P</emphasis>
|
||
. Specimen of
|
||
<taxonomicName id="CC2C5F1BDBA568C391474D120E34D709" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="4678E60BCE66775F6944D5042D3C67E3" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (
|
||
<collectionCode id="4FE7F7FE027E4EDEA8310C3A34FC1AD3" collectionName="UGSB">UGSB</collectionCode>
|
||
20496);
|
||
<emphasis id="826472E9EBE50D90FC443A8260E0A217" bold="true" pageId="0" pageNumber="25">Q</emphasis>
|
||
-
|
||
<emphasis id="2AC006FC160EF9B73F16ED410585C487" bold="true" pageId="0" pageNumber="25">T</emphasis>
|
||
. Specimen of
|
||
<taxonomicName id="1C240F36E11CE188DFC2EC687912BAEE" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="EB989A226E07F1BD85159E798CA35837" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
from Zoeram, North Khorasan, Iran (
|
||
<collectionCode id="4C743FF0C45601B6DD21CB3289400BE5" collectionName="UGSB">UGSB</collectionCode>
|
||
21661);
|
||
<emphasis id="CDE756FDD94D2DA9C3A75F0B35737931" bold="true" pageId="0" pageNumber="25">U</emphasis>
|
||
,
|
||
<emphasis id="A79B1B5D5F1D1A2934ABEF19C04F9065" bold="true" pageId="0" pageNumber="25">V</emphasis>
|
||
. Specimen of
|
||
<taxonomicName id="8B764CEB68C2106F072C6A39A7412E2F" lsidName="T. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
|
||
<emphasis id="72E7A1CEEEB11CDCD04962057904F53D" italics="true" pageId="0" pageNumber="25">T. schultzii</emphasis>
|
||
</taxonomicName>
|
||
collected in the Caspian Sea, offshore from Aktau, Kazakhstan (
|
||
<collectionCode id="3519664BA4746F8147F674B79AF03A92" collectionName="UGSB">UGSB</collectionCode>
|
||
20714);
|
||
<emphasis id="8A217DB01B921CAF9836F955877D9B3E" bold="true" pageId="0" pageNumber="25">W</emphasis>
|
||
-
|
||
<emphasis id="52DFB01A7599A776F896313D786FED9A" bold="true" pageId="0" pageNumber="25">Y</emphasis>
|
||
.
|
||
<typeStatus id="AEED0C16502D246475D512A0CB14A1A8">Lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="F85465E5D5631B8AA6D1389EECD3FB3E" lsidName="T. pallasi" pageId="0" pageNumber="25" rank="species" species="pallasi">
|
||
<emphasis id="982DB93217F23CBDDFD21019FDAC0726" italics="true" pageId="0" pageNumber="25">T. pallasi</emphasis>
|
||
</taxonomicName>
|
||
from Caspian Sea, Dagestan, Russia (
|
||
<collectionCode id="807B057E49670E676D79BCCB3240E0B6" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
54547/63);
|
||
<emphasis id="6CBF21C5ED170E4913693AAFA330EC07" bold="true" pageId="0" pageNumber="25">Za</emphasis>
|
||
,
|
||
<emphasis id="10639701786E4648292D473398206CFA" bold="true" pageId="0" pageNumber="25">Zb</emphasis>
|
||
.
|
||
<typeStatus id="0B872EA773BA4A9B974B7ED5FF245568">Syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="736AA41C0023EE3F3E7AFC9B483FB9C6" lsidName="N. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
|
||
<emphasis id="0508730AC455B09BEF4C13FCCD70E929" italics="true" pageId="0" pageNumber="25">N. schultzii</emphasis>
|
||
</taxonomicName>
|
||
from the Caspian Sea (
|
||
<collectionCode id="01DBB45C86D5265F2AE5C2C4426F40E5" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" type="Museum">ZIN</collectionCode>
|
||
, no.
|
||
<quantity id="8CB0A49124A7C160251A04EBC937299E" metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
|
||
systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig.
|
||
<figureCitation id="5B6759232FB9C21E20BC8A172A3EFB73" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). Scale bars:
|
||
<quantity id="E8C1CCB6BC9B8D01401EC305197160F4" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="2C1DE9F69A810DAE1861B8B9D961B341" pageId="0" pageNumber="25" type="distribution">
|
||
<paragraph id="290E2969AE765AAC7F4746E00556F9A6" pageId="0" pageNumber="25">Distribution.</paragraph>
|
||
<paragraph id="FBFDB92BC80571452B2156A0D8E714F5" pageId="0" pageNumber="25">
|
||
<taxonomicName id="90DE54E29A4290558E89B40041A8D189" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus major" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="major">
|
||
<emphasis id="6930264FE4669D9AB5EECA35F5D2F886" italics="true" pageId="0" pageNumber="25">Theodoxus major</emphasis>
|
||
</taxonomicName>
|
||
is found in the Caspian Sea and parts of the Azov Sea (
|
||
<bibRefCitation id="BB48E619A00C5B8F4F353FCCED8E6540" author="Roth, G" editor="Krupp, F" journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany" pageId="0" pageNumber="25" pagination="72 - 79" refId="B136" refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79" title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)." volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East." year="1987">Roth 1987</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2DF7FFB1BFEF58D04FC04E5E5862572B" author="Karpinsky, MG" journalOrPublisher="VNIRO, Moscow" pageId="0" pageNumber="25" refId="B79" refString="Karpinsky, MG, 2002. Ecology of the benthos of the Middle and Southern Caspian. VNIRO, Moscow" title="Ecology of the benthos of the Middle and Southern Caspian." year="2002">Karpinsky 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8F57555658EB170DDAB997E759F1EDE2" DOI="https://doi.org/10.1127/arch.moll/146/213-226" author="Anistratenko, VV" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="213 - 226" refId="B17" refString="Anistratenko, VV, Zettler, ML, Anistratenko, OY, 2017. On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region. Archiv fuer Molluskenkunde 146 (2): 213 - 226, DOI: https://doi.org/10.1127/arch.moll/146/213-226" title="On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region." url="https://doi.org/10.1127/arch.moll/146/213-226" volume="146" year="2017">Anistratenko et al. 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C3433215824FBE47FF564DEEB8FF3581" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
|
||
) and is also present in the Volga and Don river systems (
|
||
<bibRefCitation id="030E5B914D8DFBEB79A93E19F4221A08" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
|
||
). It has likely become regionally extinct in the Aral Sea (
|
||
<bibRefCitation id="98FC1632216BAD6D05343628CC634BE0" DOI="https://doi.org/10.1007/BF02904954" author="Andreev, NI" journalOrPublisher="International Journal of Salt Lake Research" pageId="0" pageNumber="25" pagination="103 - 110" refId="B13" refString="Andreev, NI, Andreeva, SI, Filippov, AA, Aladin, NV, 1992. The fauna of the Aral Sea in 1989. 1. The benthos. International Journal of Salt Lake Research 1: 103 - 110, DOI: https://doi.org/10.1007/BF02904954" title="The fauna of the Aral Sea in 1989. 1. The benthos." url="https://doi.org/10.1007/BF02904954" volume="1" year="1992">Andreev et al. 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="0821297D5E06E07405F95754C3A5FBE9" DOI="https://doi.org/10.1023/A:1009009924839" author="Aladin, NV" journalOrPublisher="International Journal of Salt Lake Research" pageId="0" pageNumber="25" pagination="301 - 343" refId="B5" refString="Aladin, NV, Filippov, AA, Plotnikov, IS, Orlova, MI, Williams, WD, 1998. Changes in the structure and function of biological communities in the Aral Sea, with particular reference to the northern part (Small Aral Sea), 1985-1994: a review. International Journal of Salt Lake Research 7 (4): 301 - 343, DOI: https://doi.org/10.1023/A:1009009924839" title="Changes in the structure and function of biological communities in the Aral Sea, with particular reference to the northern part (Small Aral Sea), 1985 - 1994: a review." url="https://doi.org/10.1023/A:1009009924839" volume="7" year="1998">Aladin et al. 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="64CE59CFB63327F91D4A98F2018ECA29" DOI="https://doi.org/10.1111/lre.12149" author="Plotnikov, IS" journalOrPublisher="Lakes & Reservoirs: Research & Management" pageId="0" pageNumber="25" pagination="315 - 328" refId="B123" refString="Plotnikov, IS, Ermakhanov, ZK, Aladin, NV, Micklin, P, 2016. Modern state of the Small (Northern) Aral Sea fauna. Lakes & Reservoirs: Research & Management 21 (4): 315 - 328, DOI: https://doi.org/10.1111/lre.12149" title="Modern state of the Small (Northern) Aral Sea fauna." url="https://doi.org/10.1111/lre.12149" volume="21" year="2016">Plotnikov et al. 2016</bibRefCitation>
|
||
), although it may still persist in some of its associated drainages (
|
||
<bibRefCitation id="360B6C15F8FB16EDD7F10D8A03B7E747" author="Zettler, ML" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="25" pagination="245 - 252" refId="B165" refString="Zettler, ML, 2007. A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea. Journal of Conchology 39 (3): 245 - 252" title="A redescription of Theodoxus schultzii (Grimm, 1877), an endemic neritid gastropod of the Caspian Sea." volume="39" year="2007">Zettler 2007</bibRefCitation>
|
||
). In Western Asia it can be found in the Aras River system (
|
||
<bibRefCitation id="14C50F3BDC346A440380F8C67861764A" DOI="https://doi.org/10.22092/ISFJ.2017.109948" author="Aliyev, A" journalOrPublisher="Iranian Scientific Fisheries Journal" pageId="0" pageNumber="25" pagination="131 - 142" refId="B9" refString="Aliyev, A, Ahmadi, R, 2010. Biodiversity of benetic invertebrates in Aras River. Iranian Scientific Fisheries Journal 19 (2): 131 - 142, DOI: https://doi.org/10.22092/ISFJ.2017.109948" title="Biodiversity of benetic invertebrates in Aras River." url="https://doi.org/10.22092/ISFJ.2017.109948" volume="19" year="2010">Aliyev and Ahmadi 2010</bibRefCitation>
|
||
), Lake Yerevan and its catchment systems in Armenia (
|
||
<bibRefCitation id="61A4F669B7800D7E60E167EDA4FAB15F" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
|
||
), and Masalli in Azerbaijan. Records of
|
||
<taxonomicName id="6EA6B71AEC9D02F71C1BF627D90AFEE1" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
|
||
<emphasis id="D6C006510528AE035A6AC9C9C4213197" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
|
||
</taxonomicName>
|
||
Issel, 1865 [=
|
||
<taxonomicName id="0DD1DAF676F0EF6D526DC6FFD790A2B0" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
|
||
<emphasis id="1A639B3E78FFA00E82ACC228892FF8F6" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
|
||
</taxonomicName>
|
||
],
|
||
<taxonomicName id="9DC4E6ABD9AB748EFD345A8EA69C97FE" lsidName="T. euphraticus" pageId="0" pageNumber="25" rank="species" species="euphraticus">
|
||
<emphasis id="687F713D382A54B0DE82F35F45682E8A" italics="true" pageId="0" pageNumber="25">T. euphraticus</emphasis>
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="F5A678AFE1BF184D81989828077ADFFE" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="98C84D7F677D95045F6C8484559BACEB" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
], or
|
||
<taxonomicName id="1E450ABAA857482C6571726464973549" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis id="2A54664A14EFDD467D8BD5A013918B3E" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="6D097483AB5EF26C4632DFD283F83271" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
|
||
<normalizedToken id="93807AD7B23A423F3CDCDCA6BE2C4820" originalValue="Glöer">Gloeer</normalizedToken>
|
||
and
|
||
<normalizedToken id="846F3796CABC1243D6E196E4B05AE89D" originalValue="Pešić">Pesic</normalizedToken>
|
||
(2012)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="77C4EA8CFCDE4815DE126288999FCE55" author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee (2012)</bibRefCitation>
|
||
from several mineral springs and streams in Gilan, Mazandaran, and the Khorasan provinces of northern Iran are likely misidentifications of
|
||
<taxonomicName id="100539F75E09BD44EED1186B8D26BE22" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="AF9A7F37F8F47AD211E3F3A039825F86" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="535FDA38D59049D3BE925D459714806D" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="653ECE81F6C81B7E56A2115A1BE21B20" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3B</figureCitation>
|
||
). Its presence in Lake Sevan (the supposed
|
||
<typeStatus id="F0F583C65130859DEE8674236A29D481">type</typeStatus>
|
||
locality) or the adjacent area around the lake was questioned by
|
||
<bibRefCitation id="3956A62784AFD779A4D14D7FA76F1A66" author="Akramovskiy, NN" journalOrPublisher="Izvestiya Akademii Nauk Armyanskoy SSR, Yerevan" pageId="0" pageNumber="25" refId="B3" refString="Akramovskiy, NN, 1976. Fauna Armyanskoy SSR. Mollyuski (Mollusca). Izvestiya Akademii Nauk Armyanskoy SSR, Yerevan" title="Fauna Armyanskoy SSR. Mollyuski (Mollusca)." year="1976">Akramovskiy (1976)</bibRefCitation>
|
||
(see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |