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<document id="C49A094A6502F9C15BDF7C5D142B0226" ID-CLB-Dataset="4737" ID-DOI="10.11646/zootaxa.4500.3.4" ID-GBIF-Dataset="d44cea23-cf29-4e99-b971-7b36ea9ce911" ID-ISSN="1175-5326" ID-Zenodo-Dep="5297691" ID-ZooBank="CAC076FD-030D-4851-AAE9-44068F7B7A75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630085217845" checkinUser="felipe" docAuthor="Grebennikov, Vasily V." docDate="2018" docId="AC2887EFFFAFFF87FF56F9D1FBA69BA7" docLanguage="en" docName="zootaxa.4500.3.4.pdf" docOrigin="Zootaxa 4500 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Devernodes drimo Grebennikov 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="370" masterDocId="5011FF97FFA8FF80FFC1FF9DFFAF9C0F" masterDocTitle="Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia" masterLastPageNumber="380" masterPageNumber="363" pageNumber="370" updateTime="1699146272368" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="431309C053417B207A1AC65FEA0AB6DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C81E1C65586247BF5A8EB0A569A167BE">
<mods:title id="800608E9050CFA086F0E5B55BAED9319">Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="F6542F353CC36210D439C6A73193ADE1" type="personal">
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<mods:namePart id="12E09E1E7E20FDF6A175348AB66CCAEE">Grebennikov, Vasily V.</mods:namePart>
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<mods:title id="FEF92065459E232259F31F76A1458899">Zootaxa</mods:title>
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<mods:date id="869C20B202DD8584CCBFF34EDA40C944">2018</mods:date>
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<mods:number id="FC051AF27A1FDD0D6D19A79CF28BA36D">2018-10-17</mods:number>
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<mods:classification id="E0511B4A6B87A212E49ACDEE557586D0">journal article</mods:classification>
<mods:identifier id="04941F99E9E666777DF3157EB47C397D" type="CLB-Dataset">4737</mods:identifier>
<mods:identifier id="D5E3F4D4F293112C0F0D8B0057053849" type="DOI">10.11646/zootaxa.4500.3.4</mods:identifier>
<mods:identifier id="E12FD7A11CD8E0416D743E978F43D342" type="GBIF-Dataset">d44cea23-cf29-4e99-b971-7b36ea9ce911</mods:identifier>
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<treatment id="AC2887EFFFAFFF87FF56F9D1FBA69BA7" ID-DOI="http://doi.org/10.5281/zenodo.5304359" ID-GBIF-Taxon="185119317" ID-Zenodo-Dep="5304359" LSID="urn:lsid:plazi:treatment:AC2887EFFFAFFF87FF56F9D1FBA69BA7" httpUri="http://treatment.plazi.org/id/AC2887EFFFAFFF87FF56F9D1FBA69BA7" lastPageNumber="370" pageId="7" pageNumber="370">
<subSubSection id="6C9B6572FFAFFF87FF56F9D1FE129A69" box="[151,445,1612,1638]" pageId="7" pageNumber="370" type="nomenclature">
<paragraph id="243E36F9FFAFFF87FF56F9D1FE129A69" blockId="7.[151,509,1612,1672]" box="[151,445,1612,1638]" pageId="7" pageNumber="370">
<heading id="7F768195FFAFFF87FF56F9D1FE129A69" bold="true" box="[151,445,1612,1638]" fontSize="11" level="1" pageId="7" pageNumber="370" reason="1">
<emphasis id="16F5EAEBFFAFFF87FF56F9D1FE129A69" bold="true" box="[151,445,1612,1638]" pageId="7" pageNumber="370">
<taxonomicName id="E3814D7AFFAFFF87FF56F9D1FEDB9A69" authority="Grebennikov, 2018" authorityName="Grebennikov" authorityYear="2018" box="[151,372,1612,1638]" class="Insecta" family="Curculionidae" genus="Devernodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="drimo" status="sp. nov.">
<emphasis id="16F5EAEBFFAFFF87FF56F9D1FEDB9A69" bold="true" box="[151,372,1612,1638]" italics="true" pageId="7" pageNumber="370">Devernodes drimo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DC65790FFAFFF87FEBAF9D1FE129A69" box="[379,445,1612,1638]" pageId="7" pageNumber="370" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6C9B6572FFAFFF87FF56F9F2FBD09AFB" pageId="7" pageNumber="370" type="description">
<paragraph id="243E36F9FFAFFF87FF56F9F2FE529A87" blockId="7.[151,509,1612,1672]" box="[151,509,1647,1672]" pageId="7" pageNumber="370">
<figureCitation id="BCBA2A7CFFAFFF87FF56F9F2FEBA9A87" box="[151,277,1647,1672]" captionStart="FIGURE 5" captionStartId="9.[151,250,1134,1156]" captionTargetBox="[170,1408,195,1120]" captionTargetId="figure-793@9.[170,1408,195,1120]" captionTargetPageId="9" captionText="FIGURE 5. Devernodes drimo sp. n., holotype. A–D: habitus, dorsal (A), lateral (B), ventral (C), fronto-lateral (D)." figureDoi="http://doi.org/10.5281/zenodo.5297703" httpUri="https://zenodo.org/record/5297703/files/figure.png" pageId="7" pageNumber="370">Figs 5A–D</figureCitation>
,
<figureCitation id="BCBA2A7CFFAFFF87FEE1F9EDFEF29A88" box="[288,349,1648,1671]" captionStart="FIGURE 7" captionStartId="11.[151,250,1590,1612]" captionTargetBox="[151,1436,195,1566]" captionTargetId="figure-194@11.[151,1436,195,1566]" captionTargetPageId="11" captionText="FIGURE 7. Devernodes, clades A–F, heads dorsal (A–F) and lateral (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n." figureDoi="http://doi.org/10.5281/zenodo.5297709" httpUri="https://zenodo.org/record/5297709/files/figure.png" pageId="7" pageNumber="370">7E,K</figureCitation>
,
<figureCitation id="BCBA2A7CFFAFFF87FEA8F9F2FE0A9A87" box="[361,421,1647,1672]" captionStart="FIGURE 8" captionStartId="12.[151,250,1271,1293]" captionTargetBox="[151,1436,189,1209]" captionTargetId="figure-356@12.[151,1436,189,1209]" captionTargetPageId="12" captionText="FIGURE 8. Devernodes, clades A–F, right antennae (A–F) and declivity of left elytron (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n. Numbering of elytral striae does not necessarily imply homology." figureDoi="http://doi.org/10.5281/zenodo.5297711" httpUri="https://zenodo.org/record/5297711/files/figure.png" pageId="7" pageNumber="370">8E,K</figureCitation>
,
<figureCitation id="BCBA2A7CFFAFFF87FE70F9F2FE609A87" box="[433,463,1647,1672]" captionStart="FIGURE 10" captionStartId="13.[151,250,1898,1920]" captionTargetBox="[155,1433,1263,1873]" captionTargetId="figure-145@13.[151,1436,1261,1876]" captionTargetPageId="13" captionText="FIGURE 10. Neighbour-joining clustering of 26 DNA barcoded Devernodes type specimens performed using the online BOLD tree-building engine and Kimura 2-parameter. Branch length shown is proportional to relative divergence with scale bars indicating 0.02 units. Red dots denote illustrated specimens from each of six clusters; “HT” denotes holotypes; images are to scale. Unless otherwise indicated, all type specimens are stored in CNC." figureDoi="http://doi.org/10.5281/zenodo.5297717" httpUri="https://zenodo.org/record/5297717/files/figure.png" pageId="7" pageNumber="370">10</figureCitation>
,
<figureCitation id="BCBA2A7CFFAFFF87FE1AF9F2FE569A87" box="[475,505,1647,1672]" captionStart="FIGURE 11" captionStartId="14.[151,250,1548,1570]" captionTargetBox="[151,1436,196,1523]" captionTargetId="figure-185@14.[151,1436,193,1526]" captionTargetPageId="14" captionText="FIGURE 11. Maximum Likelihood inference phylogram of Devernodes weevils from a three fragment dataset with 3223 aligned positions. Digits at internodes are bootstrap values of 60% and more. Branch length shown is proportional to relative divergence with scale bars indicating 0.2 units." figureDoi="http://doi.org/10.5281/zenodo.5297719" httpUri="https://zenodo.org/record/5297719/files/figure.png" pageId="7" pageNumber="370">11</figureCitation>
.
</paragraph>
<paragraph id="243E36F9FFAFFF87FF56F92AFBD09AFB" blockId="7.[151,1436,1719,1960]" pageId="7" pageNumber="370">
<emphasis id="16F5EAEBFFAFFF87FF56F92AFE069ADF" bold="true" box="[151,425,1719,1744]" inLineHeading="true" pageId="7" pageNumber="370" reason="1">Diagnostic description.</emphasis>
<typeStatus id="FB3A885BFFAFFF87FE71F92AFDB79ADF" box="[432,536,1719,1744]" pageId="7" pageNumber="370" type="holotype">Holotype</typeStatus>
female (
<figureCitation id="BCBA2A7CFFAFFF87FDBBF92AFD559ADF" box="[634,762,1719,1744]" captionStart="FIGURE 5" captionStartId="9.[151,250,1134,1156]" captionTargetBox="[170,1408,195,1120]" captionTargetId="figure-793@9.[170,1408,195,1120]" captionTargetPageId="9" captionText="FIGURE 5. Devernodes drimo sp. n., holotype. A–D: habitus, dorsal (A), lateral (B), ventral (C), fronto-lateral (D)." figureDoi="http://doi.org/10.5281/zenodo.5297703" httpUri="https://zenodo.org/record/5297703/files/figure.png" pageId="7" pageNumber="370">Figs 5A–D</figureCitation>
,
<figureCitation id="BCBA2A7CFFAFFF87FCC4F925FCED9AC0" box="[773,834,1720,1743]" captionStart="FIGURE 7" captionStartId="11.[151,250,1590,1612]" captionTargetBox="[151,1436,195,1566]" captionTargetId="figure-194@11.[151,1436,195,1566]" captionTargetPageId="11" captionText="FIGURE 7. Devernodes, clades A–F, heads dorsal (A–F) and lateral (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n." figureDoi="http://doi.org/10.5281/zenodo.5297709" httpUri="https://zenodo.org/record/5297709/files/figure.png" pageId="7" pageNumber="370">7E,K</figureCitation>
,
<figureCitation id="BCBA2A7CFFAFFF87FC8FF92AFC259ADF" box="[846,906,1719,1744]" captionStart="FIGURE 8" captionStartId="12.[151,250,1271,1293]" captionTargetBox="[151,1436,189,1209]" captionTargetId="figure-356@12.[151,1436,189,1209]" captionTargetPageId="12" captionText="FIGURE 8. Devernodes, clades A–F, right antennae (A–F) and declivity of left elytron (G–L), holotypes (A, C–F) and paratype 2345 (B). A,B,G,H: D. alkippe sp. n., C, I: D. asteria sp. n., D, J: D. chthonia sp. n., E, K: D. drimo sp. n., F, L: D. methone sp. n. Numbering of elytral striae does not necessarily imply homology." figureDoi="http://doi.org/10.5281/zenodo.5297711" httpUri="https://zenodo.org/record/5297711/files/figure.png" pageId="7" pageNumber="370">8E,K</figureCitation>
), body length 4.0 mm, DNA barcode GenBank accession number in
<figureCitation id="BCBA2A7CFFAFFF87FE43F946FE7A9AFB" box="[386,469,1755,1780]" captionStart="FIGURE 10" captionStartId="13.[151,250,1898,1920]" captionTargetBox="[155,1433,1263,1873]" captionTargetId="figure-145@13.[151,1436,1261,1876]" captionTargetPageId="13" captionText="FIGURE 10. Neighbour-joining clustering of 26 DNA barcoded Devernodes type specimens performed using the online BOLD tree-building engine and Kimura 2-parameter. Branch length shown is proportional to relative divergence with scale bars indicating 0.02 units. Red dots denote illustrated specimens from each of six clusters; “HT” denotes holotypes; images are to scale. Unless otherwise indicated, all type specimens are stored in CNC." figureDoi="http://doi.org/10.5281/zenodo.5297717" httpUri="https://zenodo.org/record/5297717/files/figure.png" pageId="7" pageNumber="370">Fig. 10</figureCitation>
; unique combination of morphological characters in
<tableCitation id="69030342FFAFFF87FBE4F946FBD59AFB" box="[1061,1146,1755,1780]" captionStart="TABLE 3" captionStartId="15.[151,239,1210,1233]" captionText="TABLE 3. Discrete characters for morphological diagnostics of Devernodes weevils" pageId="7" pageNumber="370">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6C9B6572FFAFFF87FF06F962FC129B33" pageId="7" pageNumber="370" type="materials_examined">
<paragraph id="243E36F9FFAFFF87FF06F962FC129B33" blockId="7.[151,1436,1719,1960]" pageId="7" pageNumber="370">
<emphasis id="16F5EAEBFFAFFF87FF06F962FE159B17" bold="true" box="[199,442,1791,1816]" inLineHeading="true" pageId="7" pageNumber="370" reason="1">Material examined.</emphasis>
<materialsCitation id="94E93CA4FFAFFF87FE0BF962FC129B33" ID-GBIF-Occurrence="3339719303" collectingDate="2013-06-10" collectingDateMax="2013-06-21" collectingDateMin="2013-06-10" collectionCode="CNC" collectorName="E. Jendek &amp; O. Sausa" country="Malaysia" latitude="2.97" location="Pasoh forest" longLatPrecision="784" longitude="102.3" pageId="7" pageNumber="370" specimenCount="1" stateProvince="Negeri Sembilan" typeStatus="holotype">
<typeStatus id="FB3A885BFFAFFF87FE0BF962FD9D9B17" box="[458,562,1791,1816]" pageId="7" pageNumber="370" type="holotype">Holotype</typeStatus>
(
<collectionCode id="4290AE3CFFAFFF87FD8AF962FD249B17" box="[587,651,1791,1816]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="7" pageNumber="370">CNC</collectionCode>
): “
<collectingCountry id="5C967669FFAFFF87FD73F962FCB29B17" box="[690,797,1791,1816]" name="Malaysia" pageId="7" pageNumber="370">Malaysia</collectingCountry>
,
<collectingRegion id="E645F81BFFAFFF87FCF3F962FC569B17" box="[818,1017,1791,1816]" country="Malaysia" name="Negeri Sembilan" pageId="7" pageNumber="370">Negeri Sembilan</collectingRegion>
<location id="215E6022FFAFFF87FBCBF962FB329B17" LSID="urn:lsid:plazi:treatment:AC2887EFFFAFFF87FF56F9D1FBA69BA7:215E6022FFAFFF87FBCBF962FB329B17" box="[1034,1181,1791,1816]" country="Malaysia" latitude="2.97" longLatPrecision="784" longitude="102.3" name="Pasoh forest" pageId="7" pageNumber="370" stateProvince="Negeri Sembilan">Pasoh forest</location>
<geoCoordinate id="41B5503EFFAFFF87FB6FF962FB5C9B17" box="[1198,1267,1791,1816]" degrees="2.97" direction="north" orientation="latitude" pageId="7" pageNumber="370" precision="555" value="2.97">N2.97</geoCoordinate>
<geoCoordinate id="41B5503EFFAFFF87FAC2F962FACD9B17" box="[1283,1378,1791,1816]" degrees="102.30" direction="east" orientation="longitude" pageId="7" pageNumber="370" precision="555" value="102.3">E102.30</geoCoordinate>
<date id="503F1039FFAFFF87FAB3F962FEBE9B33" pageId="7" pageNumber="370" value="2013-06-10" valueMax="2013-06-21" valueMin="2013-06-10">
<collectingDate id="407BE9D1FFAFFF87FAB3F962FEBE9B33" pageId="7" pageNumber="370" value="2013-06-10" valueMax="2013-06-21" valueMin="2013-06-10">10– 21.vi.2013</collectingDate>
</date>
, 80 m
<collectorName id="8974532FFFAFFF87FE9CF8BEFE659B33" box="[349,458,1827,1852]" pageId="7" pageNumber="370">E. Jendek</collectorName>
&amp;
<collectorName id="8974532FFFAFFF87FE2CF8BEFDFF9B33" box="[493,592,1827,1852]" pageId="7" pageNumber="370">O. Šauša</collectorName>
leg”, “CNCCOLVG00005954”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6C9B6572FFAFFF87FF06F8DAFD399B8D" pageId="7" pageNumber="370" type="distribution">
<paragraph id="243E36F9FFAFFF87FF06F8DAFD399B8D" blockId="7.[151,1436,1719,1960]" pageId="7" pageNumber="370">
<emphasis id="16F5EAEBFFAFFF87FF06F8DAFECF9B6F" bold="true" box="[199,352,1863,1888]" inLineHeading="true" pageId="7" pageNumber="370" reason="1">Distribution.</emphasis>
Known only from the
<typeStatus id="FB3A885BFFAFFF87FDB7F8D5FD099B6F" box="[630,678,1864,1888]" pageId="7" pageNumber="370">type</typeStatus>
locality, Pasoh Forest Reserve, about
<quantity id="E3799B1CFFAFFF87FBADF8DAFB179B6F" box="[1132,1208,1863,1888]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="370" unit="km" value="70.0">70 km</quantity>
southeast of
<collectingRegion id="E645F81BFFAFFF87FA98F8DAFF5C9B8D" country="Malaysia" name="Kuala Lumpur" pageId="7" pageNumber="370">Kuala Lumpur</collectingRegion>
,
<collectingCountry id="5C967669FFAFFF87FF3FF8F6FECA9B8B" box="[254,357,1899,1924]" name="Malaysia" pageId="7" pageNumber="370">Malaysia</collectingCountry>
(
<figureCitation id="BCBA2A7CFFAFFF87FEB5F8F6FE689B8B" box="[372,455,1899,1924]" captionStart="FIGURE 12" captionStartId="15.[151,250,1052,1074]" captionTargetBox="[242,1361,205,1024]" captionTargetId="figure-399@15.[229,1372,195,1032]" captionTargetPageId="15" captionText="FIGURE 12. Geographical distribution of recent Lymantini. Northernmost limits in North America follow Anderson (2016); southernmost limits are based on country records (Alonso-Zarazaga &amp; Lyal 1999) and are fuzzy to reflect uncertainty. New country records (Peru, Guyana, Surinam) are based on unpublished records of Ithaura (Robert Anderson, personal communication). Map design: Jessica Hsiung." figureDoi="http://doi.org/10.5281/zenodo.5297721" httpUri="https://zenodo.org/record/5297721/files/figure.png" pageId="7" pageNumber="370">Fig. 12</figureCitation>
). Elevation:
<specimenCount id="3287FD70FFAFFF87FD97F8F6FD399B8D" box="[598,662,1899,1924]" pageId="7" pageNumber="370" type="generic">
<quantity id="E3799B1CFFAFFF87FD97F8F6FD3D9B8D" box="[598,658,1899,1924]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="370" unit="m" value="80.0">80 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="6C9B6572FFAFFF87FF06F812FBA69BA7" box="[199,1033,1935,1960]" pageId="7" pageNumber="370" type="etymology">
<paragraph id="243E36F9FFAFFF87FF06F812FBA69BA7" blockId="7.[151,1436,1719,1960]" box="[199,1033,1935,1960]" pageId="7" pageNumber="370">
<emphasis id="16F5EAEBFFAFFF87FF06F812FEE19BA7" bold="true" box="[199,334,1935,1960]" inLineHeading="true" pageId="7" pageNumber="370" reason="1">Etymology.</emphasis>
One of the seven daughters of Alcyoneus; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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