307 lines
29 KiB
XML
307 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1002.58017" ID-GBIF-Dataset="8bedb86f-0c38-4738-b1b5-1dab270c95d2" ID-PMC="PMC7746671" ID-Pensoft-Pub="1313-2970-1002-1" ID-Pensoft-UUID="92722606C9CB50B6A63A1345E68167F9" ID-PubMed="33363429" ID-ZooBank="C9EAF9DCD87D402FAC35CC4983D2E142" ModsDocID="1313-2970-1002-1" checkinTime="1607628106955" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Garces, Jhoana M. & Gattolliat, Jean-Luc" docDate="2020" docId="AC1E7D76CA6656BBA87D934B80CBD129" docLanguage="en" docName="ZooKeys 1002: 1-114" docOrigin="ZooKeys 1002" docPubDate="2020-12-10" docSource="http://dx.doi.org/10.3897/zookeys.1002.58017" docTitle="Labiobaetis dalisay Kaltenbach & Garces & Gattolliat 2020, sp. nov." docType="treatment" docUuid="0CF1B9F8-0208-4EF9-BF95-379DBFAA2FF0" docUuidSource="ZooBank" docVersion="4" id="92722606C9CB50B6A63A1345E68167F9" lastPageNumber="1" masterDocId="92722606C9CB50B6A63A1345E68167F9" masterDocTitle="The success story of Labiobaetis Novikova & Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species" masterLastPageNumber="114" masterPageNumber="1" pageNumber="1" updateTime="1668146493680" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The success story of Labiobaetis Novikova & Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kaltenbach, Thomas</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Garces, Jhoana M.</mods:namePart>
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<mods:affiliation>Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Metro Manila, Philippines</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-12-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1002</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>114</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1002.58017</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1002.58017</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1002-1</mods:identifier>
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<mods:identifier type="ZooBank">C9EAF9DCD87D402FAC35CC4983D2E142</mods:identifier>
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<mods:identifier type="Pensoft-UUID">92722606C9CB50B6A63A1345E68167F9</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="170725672" LSID="urn:lsid:zoobank.org:act:0CF1B9F8-0208-4EF9-BF95-379DBFAA2FF0" httpUri="http://treatment.plazi.org/id/AC1E7D76CA6656BBA87D934B80CBD129" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/0CF1B9F8-0208-4EF9-BF95-379DBFAA2FF0" authority="Kaltenbach & Garces & Gattolliat, 2020" authorityName="Kaltenbach & Garces & Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis dalisay" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dalisay" status="sp. nov.">Labiobaetis dalisay</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">Figures 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">, 4</figureCitation>
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Habitus, larvae, dorsal view a Labiobaetis dalisay sp. nov. b Labiobaetis acei sp. nov. c Labiobaetis aldabae sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure41" httpUri="https://binary.pensoft.net/fig/484706" pageId="0" pageNumber="1">, 41a</figureCitation>
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<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Distribution of Labiobaetis in the Philippines a Philippines, overview b Group Labiobaetis dendrisetis c Group Labiobaetis numeratus." figureDoi="10.3897/zookeys.1002.58017.figure48" httpUri="https://binary.pensoft.net/fig/484713" pageId="0" pageNumber="1">, 48b</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
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Following combination of characters: A) dorsal surface of labrum with 6-8 long, dendritic setae; B) labial palp segment II with a slender thumb-like distomedial protuberance, segment III subrectangular; C) right mandible without row of thin setae at inner margin of innermost denticle; D) fore femur length 3.4
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<normalizedToken originalValue="×">x</normalizedToken>
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maximum width, dorsal margin with 10-19 curved, spine-like setae; E) hind protoptera well developed; F) paraproct distally not expanded, with ca. 15 stout, marginal spines.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
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(Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">3</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">4</figureCitation>
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,
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Habitus, larvae, dorsal view a Labiobaetis dalisay sp. nov. b Labiobaetis acei sp. nov. c Labiobaetis aldabae sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure41" httpUri="https://binary.pensoft.net/fig/484706" pageId="0" pageNumber="1">41a</figureCitation>
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). Body length 4.8-6.5 mm. Cerci ca.
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<normalizedToken originalValue="½">1/2</normalizedToken>
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of body length. Paracercus ca. 2/3 of cerci length. Antenna approx. twice as long as head length.
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</paragraph>
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<caption doi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
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<taxonomicName authorityName="Kaltenbach & Garces & Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis dalisay" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dalisay">
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<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis dalisay</emphasis>
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</taxonomicName>
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sp. nov., larva morphology
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<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
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labrum
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<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
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seta of arc on dorsal surface of labrum
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<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
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right mandible
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<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
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right prostheca
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<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
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right incisor and kinetodontium
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<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
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left mandible
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<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
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left prostheca
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<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
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hypopharynx and superlinguae
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<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
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maxilla
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<emphasis bold="true" pageId="0" pageNumber="1">j</emphasis>
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labium
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<emphasis bold="true" pageId="0" pageNumber="1">k</emphasis>
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apex of paraglossa.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
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.
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</emphasis>
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Head, thorax, and abdomen dorsally brown with bright pattern as in Fig.
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Habitus, larvae, dorsal view a Labiobaetis dalisay sp. nov. b Labiobaetis acei sp. nov. c Labiobaetis aldabae sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure41" httpUri="https://binary.pensoft.net/fig/484706" pageId="0" pageNumber="1">41a</figureCitation>
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, fore protoptera light brown with darker striation. Head, thorax, and abdomen ventrally light brown. Legs light brown, femur basally and apically brown with a distomedial brown spot and a brown streak distomedially along dorsal margin, tibia basally with brown area, tarsus medially brown. Caudalii light brown with a brown band both at base and at ca. 1/3 of cerci length, cerci distally brown.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">4g</figureCitation>
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) with scape and pedicel subcylindrical, with well-developed distolateral process at scape.
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</paragraph>
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<caption doi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
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<taxonomicName authorityName="Kaltenbach & Garces & Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis dalisay" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dalisay">
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<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis dalisay</emphasis>
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</taxonomicName>
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sp. nov., larva morphology
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<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
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foreleg
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<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
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seta at apex of femur
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<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
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fore claw
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<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
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tergum IV
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<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
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gill IV
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<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
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paraproct
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<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
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antennal scape
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<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
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metanotum.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">3a, b</figureCitation>
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). Rectangular, length 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 6-8 long, dendritic setae. Ventrally with marginal row of setae composed of anterolateral long, feathered setae and medial long, bifid, pectinate setae; ventral surface with ca. six short, spine-like setae near lateral and anterolateral margin.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">3c, d, e</figureCitation>
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). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles, inner margin of innermost denticle without a row of thin setae; denticles of both incisor and kinetodontium of unused mandibles with secondary dentation. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex. Tuft of setae at apex of mola present.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">3f, g</figureCitation>
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). Incisor and kinetodontium fused. Incisor with four denticles; kinetodontium with three denticles. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola slightly convex. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola present.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins slightly convex. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hypopharynx and superlinguae</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">3h</figureCitation>
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). Lingua approx. as long as superlinguae. Lingua longer than broad; medial tuft of stout setae poorly developed; distal half laterally expanded. Superlinguae distally rounded; lateral margin rounded; fine, long, simple setae along distal margin.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">3i</figureCitation>
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). Galea-lacinia ventrally with two simple, apical setae under canines. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and five medium to long, simple setae. Maxillary palp approx. as long as length of galea-lacinia; 2-segmented; palp segment II 1.6
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<normalizedToken originalValue="×">x</normalizedToken>
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length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment slightly pointed, with slight excavation at inner distolateral margin.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labium</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labiobaetis dalisay sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h hypopharynx and superlinguae i maxilla j labium k apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure3" httpUri="https://binary.pensoft.net/fig/484668" pageId="0" pageNumber="1">3j, k</figureCitation>
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). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca. ten spine-like setae increasing in length distally; apex with two long, robust, pectinate setae and one short, robust seta; outer margin with three spine-like setae; ventral surface with fine, simple, scattered setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and 4-6 medium, simple setae in anteromedial area; outer, anterolateral margin with some long, spine-like setae; dorsally with a row of three long, spine-like setae near inner margin. Labial palp with segment I 0.9
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<normalizedToken originalValue="×">x</normalizedToken>
|
||
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with slender, thumb-like distomedial protuberance; distomedial protuberance 0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with two or three spine-like, simple setae near outer margin. Segment III subrectangular; length 0.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hind protoptera</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">4h</figureCitation>
|
||
) well developed.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">4a-c</figureCitation>
|
||
). Ratio of foreleg segments 1.4:1.0:0.7:0.3.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Femur</emphasis>
|
||
. Length ca. 3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
maximum width. Dorsal margin with a row of 10-19 curved, spine-like setae, apically rounded and sometimes with minute dentation; length of setae 0.13
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
maximum width of femur. Apex rounded, with a pair of spine-like setae, apically rounded and sometimes with minute dentation and some short, stout setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch present.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Tibia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dorsal margin with two rows of short, spine-like setae. Ventral margin with a row of short, curved, spine-like setae, on apex some longer, partly bipectinate, spine-like setae and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 2/3 area.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Tarsus</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dorsal margin with one or two rows of short, stout setae. Ventral margin with a row of curved, spine-like setae. Claw with one row of 13-15 denticles; distally pointed; with five or six stripes; subapical setae absent.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Terga</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">4d</figureCitation>
|
||
). Surface with irregular rows of W-shaped scale bases and scattered fine, simple setae. Posterior margin of tergum IV with triangular spines, longer than wide.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gills</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">4e</figureCitation>
|
||
). Present on segments I-VII. Margin with small denticles intercalating fine simple setae. Tracheae extending from main trunk to inner and outer margins. Gill I ca. 2/3 length of segment II. Gill IV as long as length of segments V and 2/3 VI combined. Gill VII as long as length of segments VIII and 1/3 IX combined.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Labiobaetis dalisay sp. nov., larva morphology a foreleg b seta at apex of femur c fore claw d tergum IV e gill IV f paraproct g antennal scape h metanotum." figureDoi="10.3897/zookeys.1002.58017.figure4" httpUri="https://binary.pensoft.net/fig/484669" pageId="0" pageNumber="1">4f</figureCitation>
|
||
). Distally not expanded, with ca. 15 stout, marginal spines. Surface scattered with U-shaped scale bases and fine, simple setae. Cercotractor with numerous small, marginal spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Named after the Filipino word
|
||
<taxonomicName authorityName="Kaltenbach & Garces & Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis dalisay" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dalisay">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis dalisay</emphasis>
|
||
</taxonomicName>
|
||
meaning pristine, which describes the localities where the species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Philippines: Luzon (Fig.
|
||
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Distribution of Labiobaetis in the Philippines a Philippines, overview b Group Labiobaetis dendrisetis c Group Labiobaetis numeratus." figureDoi="10.3897/zookeys.1002.58017.figure48" httpUri="https://binary.pensoft.net/fig/484713" pageId="0" pageNumber="1">48b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biological aspects">
|
||
<paragraph pageId="0" pageNumber="1">Biological aspects.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The specimens were collected at altitudes from 60 m to 400 m, mainly in pristine areas.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||
.
|
||
</emphasis>
|
||
Philippines • larva; Luzon, Maria Aurora, Wenceslao, Bingwangan River;
|
||
<geoCoordinate degrees="15" direction="north" minutes="45" orientation="latitude" precision="15" seconds="48" value="15.763333">15°45'48"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="121" direction="east" minutes="25" orientation="longitude" precision="15" seconds="21" value="121.4225">121°25'21"E</geoCoordinate>
|
||
; 60 m; 05.II.1998; leg. Mendoza; on slide; GBIFCH 00592279; PNM.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||
.
|
||
</emphasis>
|
||
Philippines • larva; Luzon, Nueva Ecija, Pantabangan, Candaclan River;
|
||
<geoCoordinate degrees="15" direction="north" minutes="46" orientation="latitude" precision="15" seconds="48" value="15.78">15°46'48"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="121" direction="east" minutes="13" orientation="longitude" precision="15" seconds="17" value="121.22139">121°13'17"E</geoCoordinate>
|
||
; 240 m; 05.II.1998; leg. Mendoza; on slide; GBIFCH 00654909; ZSM • 6 larvae; Luzon, Benguet, Tuba, Taloy Sur;
|
||
<geoCoordinate degrees="16" direction="north" minutes="21" orientation="latitude" precision="15" seconds="33" value="16.359167">16°21'33"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="120" direction="east" minutes="30" orientation="longitude" precision="15" seconds="31" value="120.50861">120°30'31"E</geoCoordinate>
|
||
; 400 m; XI. 1997; leg. Mey; 1 on slide; GenBank: MT830940; GBIFCH 00763649; AdMU; 5 in alcohol; GBIFCH 00515405; MZL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |