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<document id="556CCFADDCFAE700DEC772DC5E591945" ID-CLB-Dataset="86427" ID-DOI="10.5281/zenodo.6789060" ID-GBIF-Dataset="11bee6ab-227a-4dec-8d51-250b7067cc1b" ID-ISSN="1175­5334" ID-Zenodo-Dep="6789060" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625624496800" checkinUser="felipe" docAuthor="Published, First" docDate="2007" docId="A77887C2FFC6FFE6FF02846244CAB4E7" docLanguage="en" docName="zt01408p078.pdf" docOrigin="Zootaxa 1408" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Leptotyphlops tanae Published 2007, sp. nov." docType="treatment" docVersion="6" lastPageNumber="29" masterDocId="5B41FFBAFFDDFFFAFF95804D4142B735" masterDocTitle="A revision of the genus Leptotyphlops in northeastern Africa and southwestern Arabia (Serpentes: Leptotyphlopidae)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="28" updateTime="1699025959295" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6D34D93AC0CCAEACC06A3D7D02BD13B4">A revision of the genus Leptotyphlops in northeastern Africa and southwestern Arabia (Serpentes: Leptotyphlopidae)</mods:title>
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<mods:title id="E34A8610C9047440F9F50BBB614A3436">Zootaxa</mods:title>
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<mods:part id="522286B10BBA0B975E411F34C83A3D88">
<mods:date id="723B1DDEAF9F745FD0175159B7FCCFBA">2007</mods:date>
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<mods:number id="546C05EE7F00D1377DE5F99D74CB7F81">2007-02-15</mods:number>
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<mods:number id="1D070FEDC1F75A1E21E7C36086E821C5">1408</mods:number>
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<treatment id="A77887C2FFC6FFE6FF02846244CAB4E7" ID-DOI="http://doi.org/10.5281/zenodo.6789119" ID-GBIF-Taxon="196374657" ID-Zenodo-Dep="6789119" LSID="urn:lsid:plazi:treatment:A77887C2FFC6FFE6FF02846244CAB4E7" httpUri="http://treatment.plazi.org/id/A77887C2FFC6FFE6FF02846244CAB4E7" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="67CB655FFFC6FFE1FF028462402AB359" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="2F6E36D4FFC6FFE1FF02846243F6B37C" blockId="27.[151,692,1071,1132]" box="[151,692,1071,1097]" pageId="27" pageNumber="28">
<heading id="742681B8FFC6FFE1FF02846243F6B37C" bold="true" box="[151,692,1071,1097]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="1DA5EAC6FFC6FFE1FF02846243F6B37C" bold="true" box="[151,692,1071,1097]" pageId="27" pageNumber="28">
<taxonomicName id="E8D14D57FFC6FFE1FF02846240CBB37C" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[151,393,1071,1097]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="tanae" status="sp. nov.">
<emphasis id="1DA5EAC6FFC6FFE1FF02846240CBB37C" bold="true" box="[151,393,1071,1097]" italics="true" pageId="27" pageNumber="28">Leptotyphlops tanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="069657BDFFC6FFE1FE04846240ACB37C" box="[401,494,1071,1097]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="B7EA2A51FFC6FFE1FE6B84624314B37C" box="[510,598,1071,1097]" captionStart="PLATE 8" captionStartId="24.[151,238,1805,1829]" captionTargetBox="[286,1295,357,1792]" captionTargetId="figure-187@24.[250,1338,326,1792]" captionTargetPageId="24" captionText="PLATE 8. Figure 1. Leptotyphlops cairi. Dorsal, lateral and ventral views of the head of SK 2124 — Siwa Oasis, Egypt, after Kramer &amp; Schnurrenberger (1963) [as Leptotyphlops macrorhynchus]. The line represents 1 mm to scale. Figure 2. Leptotyphlops braccianii. Dorsal, lateral and ventral views of the head of the holotype of G. variabilis, MSNM 3348b — Adi Ugri, Eritrea, after Scortecci (1928). Figure 3. Leptotyphlops ionidesi sp. nov. Dorsal and lateral views of the head of the holotype, MCZ 57440 — Liwale, Tanzania. The line represents 1 mm to scale. Figure 4. Leptotyphlops tanae sp. nov. Dorsal and lateral views of the head of the holotype, MCZ 40099 — Ngatana, Kenya. The line represents 1 mm to scale." figureDoi="http://doi.org/10.5281/zenodo.6789090" httpUri="https://zenodo.org/record/6789090/files/figure.png" pageId="27" pageNumber="28">Plate 8</figureCitation>
, Fig. 4)
</emphasis>
</heading>
</paragraph>
<paragraph id="2F6E36D4FFC6FFE1FF02841F402AB359" blockId="27.[151,692,1071,1132]" box="[151,360,1106,1132]" pageId="27" pageNumber="28">
<collectingRegion id="ED15F836FFC6FFE1FF02841F418DB359" box="[151,207,1106,1132]" country="Kenya" name="Tana River" pageId="27" pageNumber="28">Tana</collectingRegion>
worm-snake
</paragraph>
</subSubSection>
<subSubSection id="67CB655FFFC6FFE1FF0284EF427AB20C" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="2F6E36D4FFC6FFE1FF0284EF4568B38C" blockId="27.[151,1435,1186,1337]" box="[151,1066,1186,1209]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="0FC111FAFFC6FFE1FF0284EF4568B38C" box="[151,1066,1186,1209]" pageId="27" pageNumber="28">
<emphasis id="1DA5EAC6FFC6FFE1FF0284EF4089B38C" box="[151,459,1186,1209]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="E8D14D57FFC6FFE1FF0284EF40EEB38C" authorityName="Loveridge" authorityYear="1953" box="[151,428,1186,1209]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="longicauda">Leptotyphlops longicauda</taxonomicName>
—
</emphasis>
<treatmentCitation id="AE7010C5FFC6FFE1FE4784EF43F8B38C" author="Loveridge, A." box="[466,698,1186,1209]" page="231" pageId="27" pageNumber="28" year="1936">
<bibRefCitation id="4B404B25FFC6FFE1FE4784EF43F8B38C" author="Loveridge" box="[466,698,1186,1209]" firstAuthor="Loveridge" page="231" pageId="27" pageNumber="28" pagination="209 - 337" refId="ref60644" refString="Loveridge, A. (1936) Scientific results of an expedition to rain forest regions in eastern Africa. V. Reptiles. Bulletin of the Museum of Comparative Zoology at Harvard College (Cambridge), 79 (5), 209 - 337, Pls. 1 - 9." type="journal article" year="1936">Loveridge, 1936: 231</bibRefCitation>
</treatmentCitation>
,
<bibRefCitation id="4B404B25FFC6FFE1FD5084EF4272B38C" author="Loveridge, A." box="[709,816,1186,1209]" pageId="27" pageNumber="28" pagination="1 - 19" refId="ref60934" refString="Loveridge, A. (1957 a) On a third collection of reptiles taken in Tanganyika by C. J. P. Ionides, Esq. Tanganyika Notes and Records, (43), 1 - 19." type="journal article" year="1957">1957: 247</bibRefCitation>
(part); List, 1966: 107.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F6E36D4FFC6FFE1FF02848F4353B3CC" blockId="27.[151,1435,1186,1337]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="0FC111FAFFC6FFE1FF02848F4353B3CC" pageId="27" pageNumber="28">
<emphasis id="1DA5EAC6FFC6FFE1FF02848F409AB3EC" box="[151,472,1218,1241]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="E8D14D57FFC6FFE1FF02848F40FBB3EC" authorityName="Hahn" authorityYear="1980" box="[151,441,1218,1241]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="longicaudus">Leptotyphlops longicaudus</taxonomicName>
—
</emphasis>
<treatmentCitation id="AE7010C5FFC6FFE1FE75848F43CFB3EC" author="Spawls, S." box="[480,653,1218,1241]" page="3" pageId="27" pageNumber="28" year="1978">
<bibRefCitation id="4B404B25FFC6FFE1FE75848F43CFB3EC" author="Spawls" box="[480,653,1218,1241]" firstAuthor="Spawls" page="3" pageId="27" pageNumber="28" pagination="1 - 18" refId="ref62430" refString="Spawls, S. (1978) A checklist of the snakes of Kenya. The Journal of the East Africa Natural History Society and National Museum (Nairobi), 31 (167), 1 - 18." type="journal article" year="1978">Spawls, 1978: 3</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="AE7010C5FFC6FFE1FD74848F42DDB3EC" author="Lanza, B." box="[737,927,1218,1241]" page="220" pageId="27" pageNumber="28" year="1983">
<bibRefCitation id="4B404B25FFC6FFE1FD74848F42DDB3EC" author="Lanza" box="[737,927,1218,1241]" firstAuthor="Lanza" page="220" pageId="27" pageNumber="28" pagination="193 - 247" refId="ref59808" refString="Lanza, B. (1983) A list of the Somali amphibians and reptiles. Monitore Zoologico Italiano, Firenze, (N. S.) Supplemento, 18 (8), 193 - 247." type="journal article" year="1983">Lanza, 1983: 220</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="AE7010C5FFC6FFE1FC39848F455EB3EC" author="Lanza, B." box="[940,1052,1218,1241]" page="431" pageId="27" pageNumber="28" year="1990">
<bibRefCitation id="4B404B25FFC6FFE1FC39848F455EB3EC" author="Lanza" box="[940,1052,1218,1241]" firstAuthor="Lanza" page="431" pageId="27" pageNumber="28" pagination="407 - 465" refId="ref59847" refString="Lanza, B. (1990) Amphibians and reptiles of the Somali Democratic Republic: check list and bibliography. Biogeographia (Bologna), (1988) 14, 407 - 465." type="journal article" year="1990">1990: 431</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AE7010C5FFC6FFE1FBBD848F4413B3EC" author="McDiarmid, R. W. &amp; Campbell, J. A. &amp; Toure, T. A." box="[1064,1361,1218,1241]" page="34" pageId="27" pageNumber="28" year="1999">
<bibRefCitation id="4B404B25FFC6FFE1FBBD848F4413B3EC" author="McDiarmid" box="[1064,1361,1218,1241]" etAl="et al." firstAuthor="McDiarmid" page="34" pageId="27" pageNumber="28" refId="ref61066" refString="McDiarmid, R. W., Campbell, J. A. &amp; Toure, T. A. (1999) Snake species of the world: a taxonomic and geographic reference, Volume 1. The Herpetologists' League, Washington, 511 pp." type="book" year="1999">
McDiarmid
<emphasis id="1DA5EAC6FFC6FFE1FB39848F45A8B3EC" box="[1196,1258,1218,1241]" italics="true" pageId="27" pageNumber="28">et al.,</emphasis>
1999: 34
</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="AE7010C5FFC6FFE1FF5284AF408AB3CC" author="Spawls, S. &amp; Howell, K. &amp; Drewes, R. &amp; Ashe, J." box="[199,456,1250,1273]" page="304" pageId="27" pageNumber="28" year="2002">
<bibRefCitation id="4B404B25FFC6FFE1FF5284AF408AB3CC" author="Spawls" box="[199,456,1250,1273]" etAl="et al." firstAuthor="Spawls" page="304" pageId="27" pageNumber="28" refId="ref62470" refString="Spawls, S., Howell, K., Drewes, R. &amp; Ashe, J. (2002) A field guide to the reptiles of East Africa: Kenya, Tanzania, Uganda, Rwanda and Burundi. Academic Press, San Diego, 543 pp." type="book" year="2002">
Spawls
<emphasis id="1DA5EAC6FFC6FFE1FE8E84AF4015B3CC" box="[283,343,1250,1273]" italics="true" pageId="27" pageNumber="28">et al.,</emphasis>
2002: 304
</bibRefCitation>
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F6E36D4FFC6FFE1FF02854F4577B22C" blockId="27.[151,1435,1186,1337]" box="[151,1077,1282,1305]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="0FC111FAFFC6FFE1FF02854F4577B22C" box="[151,1077,1282,1305]" pageId="27" pageNumber="28">
<emphasis id="1DA5EAC6FFC6FFE1FF02854F40C7B22C" box="[151,389,1282,1305]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="E8D14D57FFC6FFE1FF02854F4024B22C" box="[151,358,1282,1305]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cairi">Leptotyphlops cairi</taxonomicName>
—
</emphasis>
<treatmentCitation id="AE7010C5FFC6FFE1FE19854F4395B22C" author="Hoevers, L. G. &amp; Johnson, P. M." box="[396,727,1282,1305]" page="182" pageId="27" pageNumber="28" year="1982">
<bibRefCitation id="4B404B25FFC6FFE1FE19854F4395B22C" author="Hoevers &amp; Johnson" box="[396,727,1282,1305]" firstAuthor="Hoevers" page="182" pageId="27" pageNumber="28" pagination="173 - 203" refId="ref59151" refString="Hoevers, L. G. &amp; Johnson, P. M. (1982) Notes on a collection of snakes from the Middle Juba region, Somalia. Monitore Zoologico Italiano (Firenze), (N. S.) Supplement, 16 (8), 173 - 203." type="journal article" year="1982">Hoevers &amp; Johnson, 1982: 182</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="AE7010C5FFC6FFE1FCB2854F42AFB22C" author="Lambert, M. R. K." box="[807,1005,1282,1305]" page="66" pageId="27" pageNumber="28" year="1985">
<bibRefCitation id="4B404B25FFC6FFE1FCB2854F42AFB22C" author="Lambert" box="[807,1005,1282,1305]" firstAuthor="Lambert" page="66" pageId="27" pageNumber="28" pagination="58 - 71" refId="ref59710" refString="Lambert, M. R. K. (1985) Reptiles. In: Report on the species of vertebrates in riverine and coastal forest habitats of southern Somalia and the lower Tana in Kenya. University College London &amp; Tropical Development and Research Institute, London, pp. 58 - 71." type="book chapter" year="1985">Lambert, 1985: 66</bibRefCitation>
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F6E36D4FFC6FFE1FF02856F427AB20C" blockId="27.[151,1435,1186,1337]" box="[151,824,1314,1337]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="0FC111FAFFC6FFE1FF02856F427AB20C" box="[151,824,1314,1337]" pageId="27" pageNumber="28">
<taxonomicName id="E8D14D57FFC6FFE1FF02856F408EB20C" authorityName="Hahn" authorityYear="1980" box="[151,460,1314,1337]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="longicaudatus">
<emphasis id="1DA5EAC6FFC6FFE1FF02856F408EB20C" box="[151,460,1314,1337]" italics="true" pageId="27" pageNumber="28">Leptotyphlops longicaudatus</emphasis>
</taxonomicName>
(sic) —
<treatmentCitation id="AE7010C5FFC6FFE1FDBF856F43B2B20C" author="Lambert, M. R. K." box="[554,752,1314,1337]" page="66" pageId="27" pageNumber="28" year="1985">
<bibRefCitation id="4B404B25FFC6FFE1FDBF856F43B2B20C" author="Lambert" box="[554,752,1314,1337]" firstAuthor="Lambert" page="66" pageId="27" pageNumber="28" pagination="58 - 71" refId="ref59710" refString="Lambert, M. R. K. (1985) Reptiles. In: Report on the species of vertebrates in riverine and coastal forest habitats of southern Somalia and the lower Tana in Kenya. University College London &amp; Tropical Development and Research Institute, London, pp. 58 - 71." type="book chapter" year="1985">Lambert, 1985: 66</bibRefCitation>
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="67CB655FFFC6FFE1FF02852740CAB111" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="2F6E36D4FFC6FFE1FF02852740ACB2E1" blockId="27.[151,1437,1386,2012]" pageId="27" pageNumber="28">
<materialsCitation id="9FB93C89FFC6FFE1FF02852740ACB2E1" ID-GBIF-Occurrence="3827581422" accessionNumber="MCZ 40099" collectingDate="1934-06-14" collectingDateMax="1934-06-21" collectingDateMin="1934-06-14" collectionCode="MCZ" collectorName="A. Loveridge" country="Kenya" county="Coast Region" elevation="50" latitude="-2.5" location="Ngatana" longLatPrecision="1307" longitude="40.25" municipality="Wema" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Tana River" typeStatus="holotype">
<emphasis id="1DA5EAC6FFC6FFE1FF0285274052B2B1" bold="true" box="[151,272,1386,1412]" pageId="27" pageNumber="28">
<typeStatus id="F06A8876FFC6FFE1FF0285274049B2B1" box="[151,267,1386,1412]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="49C0AE11FFC6FFE1FE8985274019B2B1" box="[284,347,1386,1412]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">MCZ</collectionCode>
40099, a male from old village of
<location id="2A0E600FFFC6FFE1FC868527423AB2B1" LSID="urn:lsid:plazi:treatment:A77887C2FFC6FFE6FF02846244CAB4E7:2A0E600FFFC6FFE1FC868527423AB2B1" box="[787,888,1386,1412]" country="Kenya" county="Coast Region" latitude="-2.5" longLatPrecision="1307" longitude="40.25" municipality="Wema" name="Ngatana" pageId="27" pageNumber="28" stateProvince="Tana River">Ngatana</location>
, ca. one mile northwest from new village of
<collectingMunicipality id="CF0AACAEFFC6FFE1FF0285DF41A6B299" box="[151,228,1426,1452]" pageId="27" pageNumber="28">Wema</collectingMunicipality>
, in the lower
<collectingRegion id="ED15F836FFC6FFE1FE1885DF4350B299" box="[397,530,1426,1452]" country="Kenya" name="Tana River" pageId="27" pageNumber="28">Tana River</collectingRegion>
,
<collectingCounty id="C60F4E58FFC6FFE1FDB585DF4386B299" box="[544,708,1426,1452]" pageId="27" pageNumber="28">Coast Region</collectingCounty>
,
<collectingCountry id="57C67644FFC6FFE1FD4785DF425CB299" box="[722,798,1426,1452]" name="Kenya" pageId="27" pageNumber="28">Kenya</collectingCountry>
(
<geoCoordinate id="4AE55013FFC6FFE1FCA785DF42D7B299" box="[818,917,1426,1452]" degrees="02" direction="south" minutes="30" orientation="latitude" pageId="27" pageNumber="28" precision="925" value="-2.5">02°30’S</geoCoordinate>
,
<geoCoordinate id="4AE55013FFC6FFE1FC3185DF454AB299" box="[932,1032,1426,1452]" degrees="40" direction="east" minutes="15" orientation="longitude" pageId="27" pageNumber="28" precision="925" value="40.25">40°15’E</geoCoordinate>
,
<elevation id="A4FCD1E7FFC6FFE1FB8285DF458FB299" box="[1047,1229,1426,1452]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="28" unit="m" value="50.0">
elevation
<quantity id="E8299B31FFC6FFE1FB1985DF458FB299" box="[1164,1229,1426,1452]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="28" unit="m" value="50.0">50 m</quantity>
</elevation>
), collected by
<collectorName id="82245302FFC6FFE1FAEA85DF4050B2E1" pageId="27" pageNumber="28">A. Loveridge</collectorName>
,
<date id="5B6F1014FFC6FFE1FE8B85F740ABB2E1" box="[286,489,1466,1492]" pageId="27" pageNumber="28" value="1934-06-14" valueMax="1934-06-21" valueMin="1934-06-14">
<collectingDate id="4B2BE9FCFFC6FFE1FE8B85F740ABB2E1" box="[286,489,1466,1492]" pageId="27" pageNumber="28" value="1934-06-14" valueMax="1934-06-21" valueMin="1934-06-14">14–21 June 1934</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="2F6E36D4FFC6FFE1FF5385AF40CAB111" blockId="27.[151,1437,1386,2012]" pageId="27" pageNumber="28">
<materialsCitation id="9FB93C89FFC6FFE1FF5385AF423DB2C9" ID-GBIF-Occurrence="3827581503" accessionNumber="MCZ 40093" box="[198,895,1506,1532]" collectionCode="NMW" pageId="27" pageNumber="28" specimenCode="MCZ 40093-98, 40100-08, 40110-11" specimenCount="1" typeStatus="paratype">
<emphasis id="1DA5EAC6FFC6FFE1FF5385AF4009B2C9" bold="true" box="[198,331,1506,1532]" pageId="27" pageNumber="28">
<typeStatus id="F06A8876FFC6FFE1FF5385AF4004B2C9" box="[198,326,1506,1532]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="7F779EAFFFC6FFE1FEC385AF435AB2C9" box="[342,536,1506,1532]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">MCZ 40093–98</specimenCode>
,
<specimenCode id="7F779EAFFFC6FFE1FDB285AF43DDB2C9" box="[551,671,1506,1532]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">40100–08</specimenCode>
,
<specimenCode id="7F779EAFFFC6FFE1FD3B85AF4267B2C9" box="[686,805,1506,1532]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="27" pageNumber="28" type="Museum">40110–11</specimenCode>
;
<collectionCode id="49C0AE11FFC6FFE1FCA185AF423DB2C9" box="[820,895,1506,1532]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28">NMW</collectionCode>
</materialsCitation>
15464 (2);
<collectionCode id="49C0AE11FFC6FFE1FB8685AF452AB2C9" box="[1043,1128,1506,1532]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="27" pageNumber="28" type="Museum">USNM</collectionCode>
12081–82
<materialsCitation id="9FB93C89FFC6FFE1FB6785AF40C6B111" ID-GBIF-Occurrence="3827581530" accessionNumber="MCZ 40099" collectingDate="1934-06-14" collectingDateMax="1934-06-21" collectingDateMin="1934-06-14" collectionCode="USNM" collectorName="A. Loveridge" country="Kenya" county="Coast Region" elevation="50" latitude="-2.5" location="Ngatana" longLatPrecision="1307" longitude="40.25" municipality="Wema" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Tana River" typeStatus="paratype">with the same data as the holotype</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="67CB655FFFC6FFE1FF53867F4080B1A9" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="2F6E36D4FFC6FFE1FF53867F4080B1A9" blockId="27.[151,1437,1386,2012]" pageId="27" pageNumber="28">
<emphasis id="1DA5EAC6FFC6FFE1FF53867F4004B179" bold="true" box="[198,326,1586,1612]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
<taxonomicName id="E8D14D57FFC6FFE1FEDB867E437CB179" authorityName="Published" authorityYear="2007" box="[334,574,1587,1612]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="tanae">
<emphasis id="1DA5EAC6FFC6FFE1FEDB867E437CB179" box="[334,574,1587,1612]" italics="true" pageId="27" pageNumber="28">Leptotyphlops tanae</emphasis>
</taxonomicName>
differs from all East African members of the
<taxonomicName id="E8D14D57FFC6FFE1FBCA867E45F5B179" box="[1119,1207,1587,1612]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cairi">
<emphasis id="1DA5EAC6FFC6FFE1FBCA867E45F5B179" box="[1119,1207,1587,1612]" italics="true" pageId="27" pageNumber="28">L. cairi</emphasis>
</taxonomicName>
complex in its low counts for middorsals (227–260) and subcaudals (25–30) and its very small size. Skull with a large frontoparietal foramen like
<emphasis id="1DA5EAC6FFC6FFE1FEF186CE4080B1A9" box="[356,450,1667,1692]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="E8D14D57FFC6FFE1FEF186CE40FCB1A9" box="[356,446,1667,1692]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cairi">L. cairi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="67CB655FFFC6FFE1FF5386E74536B1F1" box="[198,1140,1706,1732]" pageId="27" pageNumber="28" type="etymology">
<paragraph id="2F6E36D4FFC6FFE1FF5386E74536B1F1" blockId="27.[151,1437,1386,2012]" box="[198,1140,1706,1732]" pageId="27" pageNumber="28">
<emphasis id="1DA5EAC6FFC6FFE1FF5386E74011B1F1" bold="true" box="[198,339,1706,1732]" pageId="27" pageNumber="28">Etymology.</emphasis>
Named for the lower
<collectingRegion id="ED15F836FFC6FFE1FDC086E7439AB1F1" box="[597,728,1706,1732]" country="Kenya" name="Tana River" pageId="27" pageNumber="28">Tana River</collectingRegion>
, where the
<typeStatus id="F06A8876FFC6FFE1FCC886E742CDB1F1" box="[861,911,1706,1732]" pageId="27" pageNumber="28">type</typeStatus>
locality is situated.
</paragraph>
</subSubSection>
<subSubSection id="67CB655FFFC6FFE6FF53869F425DB46F" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="2F6E36D4FFC6FFE1FF53869F45DEB021" blockId="27.[151,1437,1386,2012]" pageId="27" pageNumber="28">
<emphasis id="1DA5EAC6FFC6FFE1FF53869F4015B1D9" bold="true" box="[198,343,1746,1772]" pageId="27" pageNumber="28">Description</emphasis>
(
<typeStatus id="F06A8876FFC6FFE1FEFC869F4092B1D9" box="[361,464,1746,1772]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
variations in parentheses). Body cylindrical, with head broadened and flattened, head tubercles prominent, the moderate tail tapers to a stout downturned terminal spine.
</paragraph>
<paragraph id="2F6E36D4FFC6FFE6FF53876F45AFB61F" blockId="27.[151,1437,1386,2012]" lastBlockId="28.[151,1436,152,978]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<materialsCitation id="9FB93C89FFC6FFE6FF53876F45A8B61F" ID-GBIF-Occurrence="3827581429" accessionNumber="MCZ 40100, MCZ 40097" collectionCode="MCZ" country="Kenya" county="Parietals" lastPageId="28" lastPageNumber="29" location="Snout" municipality="Supraoculars" pageId="27" pageNumber="28" specimenCount="1">
<location id="2A0E600FFFC6FFE1FF53876F4048B009" LSID="urn:lsid:plazi:treatment:A77887C2FFC6FFE6FF02846244CAB4E7:2A0E600FFFC6FFE1FF53876F4048B009" box="[198,266,1826,1852]" country="Kenya" county="Parietals" municipality="Supraoculars" name="Snout" pageId="27" pageNumber="28">Snout</location>
rounded, rostral moderate (0.36–0.50 head width, mean = 0.40), narrower than nasals and not extending back to the level of the eyes, a weak preoral groove inferiorly.
<location id="2A0E600FFFC6FFE1FC7E87074502B051" LSID="urn:lsid:plazi:treatment:A77887C2FFC6FFE6FF02846244CAB4E7:2A0E600FFFC6FFE1FC7E87074502B051" box="[1003,1088,1866,1892]" country="Kenya" county="Parietals" municipality="Supraoculars" name="Behind" pageId="27" pageNumber="28">Behind</location>
rostral, upper lip bordered by infranasal (nostril midway between rostral and supralabial along nasal suture), small anterior supralabial with width along lip equal to that of infranasal, large ocular with eye near upper anterior edge, and moderate posterior supralabial.
<collectingMunicipality id="CF0AACAEFFC6FFE1FEC9878F40B7B0E9" box="[348,501,1986,2012]" pageId="27" pageNumber="28">Supraoculars</collectingMunicipality>
slightly longer than wide, anteriorly wedged between upper nasal and ocular, posteriorly wedged between the subequal frontal and postfrontal, which are smaller than the interparietal and interoccipital.
<collectingCounty id="C60F4E58FFC1FFE6FED6808D40EAB7EF" box="[323,424,192,218]" pageId="28" pageNumber="29">Parietals</collectingCounty>
transverse, in contact with the posterior supralabials, occipitals not fused in
<collectingCountry id="57C67644FFC1FFE6FADA808D44D9B7EF" box="[1359,1435,192,218]" name="Kenya" pageId="28" pageNumber="29">Kenya</collectingCountry>
specimens (fused in
<collectionCode id="49C0AE11FFC1FFE6FE1980A54089B637" box="[396,459,232,258]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="29" type="Museum">MCZ</collectionCode>
40100 and on one side of
<collectionCode id="49C0AE11FFC1FFE6FC9A80A5420CB637" box="[783,846,232,258]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="28" pageNumber="29" type="Museum">MCZ</collectionCode>
40097) but fused in Somali and Ethiopian specimens. Temporal single. Prominent tubercles on rostral and nasals. No mental, four infralabials
</materialsCitation>
.
</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF53817545CBB64F" blockId="28.[151,1436,152,978]" pageId="28" pageNumber="29">Body covered with 14 rows of smooth, imbricate subequal scales, reducing to 10 rows on the tail. Middorsals 249 (227–260, mean 242.86, n = 21); subcaudals 30 (25–30, mean 27.71, n = 21);</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF5381C545D8B697" blockId="28.[151,1436,152,978]" box="[198,1178,392,418]" pageId="28" pageNumber="29">Total length/diameter ratio 47 (46–87); total length/tail length ratio 9.8 (10.2–12.4).</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF0281FD433FB6FF" blockId="28.[151,1436,152,978]" box="[151,637,432,458]" pageId="28" pageNumber="29">
Length of
<typeStatus id="F06A8876FFC1FFE6FE8481FD4035B6FF" box="[273,375,432,458]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
88.5 + 10 =
<quantity id="E8299B31FFC1FFE6FD9881FD433BB6FF" box="[525,633,432,458]" metricMagnitude="-2" metricUnit="m" metricValue="9.85" pageId="28" pageNumber="29" unit="mm" value="98.5">98.5 mm</quantity>
.
</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF5381954235B52F" blockId="28.[151,1436,152,978]" pageId="28" pageNumber="29">
Three to nine (usually seven) middorsal scale rows pigmented light brown to tan (in preservative, flesh pink in life
<emphasis id="1DA5EAC6FFC1FFE6FE8A824D4009B52C" box="[287,331,512,537]" italics="true" pageId="28" pageNumber="29">fide</emphasis>
<bibRefCitation id="4B404B25FFC1FFE6FEC7824D4355B52F" author="Loveridge" box="[338,535,512,538]" firstAuthor="Loveridge" pageId="28" pageNumber="29" pagination="209 - 337" refId="ref60644" refString="Loveridge, A. (1936) Scientific results of an expedition to rain forest regions in eastern Africa. V. Reptiles. Bulletin of the Museum of Comparative Zoology at Harvard College (Cambridge), 79 (5), 209 - 337, Pls. 1 - 9." type="journal article" year="1936">Loveridge, 1936</bibRefCitation>
), venter cream to pale yellow.
</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF538265422EB574" blockId="28.[151,1436,152,978]" box="[198,876,552,578]" pageId="28" pageNumber="29">
Skull (MCZ 40105) similar to that of
<emphasis id="1DA5EAC6FFC1FFE6FD158265422EB574" box="[640,876,552,577]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="E8D14D57FFC1FFE6FD158265422AB574" box="[640,872,552,577]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cairi">Leptotyphlops cairi</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF53821D45FDB58F" blockId="28.[151,1436,152,978]" pageId="28" pageNumber="29">
Everted hemipenis (MCZ 40108, SVL
<quantity id="E8299B31FFC1FFE6FD04821D43A4B55F" box="[657,742,592,618]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="28" pageNumber="29" unit="mm" value="76.0">76 mm</quantity>
, tail
<quantity id="E8299B31FFC1FFE6FC8A821D4224B55F" box="[799,870,592,618]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="28" pageNumber="29" unit="mm" value="7.0">7 mm</quantity>
, 27 subcaudals) three subcaudals (
<quantity id="E8299B31FFC1FFE6FB69821D441AB55F" box="[1276,1368,592,618]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="29" unit="mm" value="1.2">1.2 mm</quantity>
) long and 1/4 subcaudal (
<quantity id="E8299B31FFC1FFE6FEEF82354095B5A7" box="[378,471,632,658]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="mm" value="0.1">0.1 mm</quantity>
) wide, nude, uniformly narrow with a pair of lateral longitudinal flaps along basal 2/5 of organ, each flap approximately half as wide as the organ, apical tip with swollen lip,
</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF53828A404BB43F" blockId="28.[151,1436,152,978]" pageId="28" pageNumber="29">
<emphasis id="1DA5EAC6FFC1FFE6FF53828A4042B5D4" bold="true" box="[198,256,711,737]" pageId="28" pageNumber="29">Size.</emphasis>
Largest specimen (MCZ 40105) 93 + 9.5 =
<quantity id="E8299B31FFC1FFE6FC93828542C2B5D7" box="[774,896,712,738]" metricMagnitude="-1" metricUnit="m" metricValue="1.025" pageId="28" pageNumber="29" unit="mm" value="102.5">102.5 mm</quantity>
; smallest specimen (MZUF 35209) 43.5 + 4.5 =
<quantity id="E8299B31FFC1FFE6FF3A82BD4046B43F" box="[175,260,752,778]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="28" pageNumber="29" unit="mm" value="48.0">48 mm</quantity>
.
</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF53835A425DB46F" blockId="28.[151,1436,152,978]" pageId="28" pageNumber="29">
<emphasis id="1DA5EAC6FFC1FFE6FF53835A406CB404" bold="true" box="[198,302,791,817]" pageId="28" pageNumber="29">Habitat.</emphasis>
Termite mound at edge of rice swamp and within six inches of the surface of black cotton soil forming banks in flooded rice fields (
<bibRefCitation id="4B404B25FFC1FFE6FDDE830D4250B46F" author="Loveridge" box="[587,786,832,858]" firstAuthor="Loveridge" pageId="28" pageNumber="29" pagination="209 - 337" refId="ref60644" refString="Loveridge, A. (1936) Scientific results of an expedition to rain forest regions in eastern Africa. V. Reptiles. Bulletin of the Museum of Comparative Zoology at Harvard College (Cambridge), 79 (5), 209 - 337, Pls. 1 - 9." type="journal article" year="1936">Loveridge, 1936</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="67CB655FFFC1FFE6FF53832A44CAB4E7" pageId="28" pageNumber="29" type="distribution">
<paragraph id="2F6E36D4FFC1FFE6FF53832A4461B4B7" blockId="28.[151,1436,152,978]" box="[198,1315,871,898]" pageId="28" pageNumber="29">
<emphasis id="1DA5EAC6FFC1FFE6FF53832A4025B4B4" bold="true" box="[198,359,871,897]" pageId="28" pageNumber="29">Distribution.</emphasis>
Southern
<collectingCountry id="57C67644FFC1FFE6FE4A83254307B4B7" box="[479,581,872,898]" name="Ethiopia" pageId="28" pageNumber="29">Ethiopia</collectingCountry>
, southern
<collectingCountry id="57C67644FFC1FFE6FD288325425FB4B7" box="[701,797,872,898]" name="Somalia" pageId="28" pageNumber="29">Somalia</collectingCountry>
and northeastern
<collectingCountry id="57C67644FFC1FFE6FC7B8325457FB4B7" box="[1006,1085,872,898]" name="Kenya" pageId="28" pageNumber="29">Kenya</collectingCountry>
,
<quantity id="E8299B31FFC1FFE6FBDC832545F2B4B7" box="[1097,1200,872,898]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="4.0" metricValueMin="0.0" pageId="28" pageNumber="29" unit="m" value="200.0" valueMax="400.0" valueMin="0.0">0–400 m</quantity>
(
<figureCitation id="B7EA2A51FFC1FFE6FB2A83254456B4B7" box="[1215,1300,872,898]" captionStart="PLATE 5" captionStartId="13.[151,238,1737,1761]" captionTargetBox="[268,1322,271,1704]" captionTargetId="figure-152@13.[264,1323,266,1713]" captionTargetPageId="13" captionText="PLATE 5. Leptotyphlops. Distribution of the Leptotyphlops parkeri, L. reticulatus, L. longicaudus and L. nigricans species groups in northeastern Africa and southwestern Arabia." figureDoi="http://doi.org/10.5281/zenodo.6789078" httpUri="https://zenodo.org/record/6789078/files/figure.png" pageId="28" pageNumber="29">Plate 5</figureCitation>
).
</paragraph>
<paragraph id="2F6E36D4FFC1FFE6FF5383C244CAB4E7" blockId="28.[151,1436,152,978]" pageId="28" pageNumber="29">
<emphasis id="1DA5EAC6FFC1FFE6FF5383C24085B49C" bold="true" box="[198,455,911,937]" pageId="28" pageNumber="29">Additional material.</emphasis>
<materialsCitation id="9FB93C89FFC1FFE6FE4783DD4222B49F" ID-GBIF-Occurrence="3827581431" box="[466,864,912,938]" country="Ethiopia" location="Murle" pageId="28" pageNumber="29" specimenCount="1">
<collectingCountry id="57C67644FFC1FFE6FE4783DD431FB49F" box="[466,605,912,938]" name="Ethiopia" pageId="28" pageNumber="29">ETHIOPIA</collectingCountry>
.
<location id="2A0E600FFFC1FFE6FDFE83DD43F0B49F" LSID="urn:lsid:plazi:treatment:A77887C2FFC6FFE6FF02846244CAB4E7:2A0E600FFFC1FFE6FDFE83DD43F0B49F" box="[619,690,912,938]" country="Ethiopia" name="Murle" pageId="28" pageNumber="29">Murle</location>
MSNM 2979
</materialsCitation>
.
<materialsCitation id="9FB93C89FFC1FFE6FCFB83DD407EB4E7" ID-GBIF-Occurrence="3827581468" collectionCode="MZUF" country="Somalia" elevation="100" latitude="2.1333334" location="Afgoi" longLatPrecision="1307" longitude="45.133335" pageId="28" pageNumber="29" specimenCode="MZUF 35209" specimenCount="1">
<collectingCountry id="57C67644FFC1FFE6FCFB83DD42B5B49F" box="[878,1015,912,938]" name="Somalia" pageId="28" pageNumber="29">SOMALIA</collectingCountry>
.
<location id="2A0E600FFFC1FFE6FB9383DD450FB49F" LSID="urn:lsid:plazi:treatment:A77887C2FFC6FFE6FF02846244CAB4E7:2A0E600FFFC1FFE6FB9383DD450FB49F" box="[1030,1101,912,938]" country="Somalia" latitude="2.1333334" longLatPrecision="1307" longitude="45.133335" name="Afgoi" pageId="28" pageNumber="29">Afgoi</location>
,
<geoCoordinate id="4AE55013FFC1FFE6FBC983DD4586B49F" box="[1116,1220,912,938]" degrees="02" direction="north" minutes="08" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="2.1333334">02°08’N</geoCoordinate>
,
<geoCoordinate id="4AE55013FFC1FFE6FB4683DD4475B49F" box="[1235,1335,912,938]" degrees="45" direction="east" minutes="08" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="45.133335">45°08’E</geoCoordinate>
,
<quantity id="E8299B31FFC1FFE6FAD283DD44D5B49F" box="[1351,1431,912,938]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="m" value="100.0">
<elevation id="A4FCD1E7FFC1FFE6FAD283DD44D5B49F" box="[1351,1431,912,938]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<specimenCode id="7F779EAFFFC1FFE6FF0283F5407EB4E7" box="[151,316,952,978]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="28" pageNumber="29">MZUF 35209</specimenCode>
</materialsCitation>
;
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E of
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