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<document id="573B6737CE766C607E33EA7CCA09FAC0" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFF36D45D6DDDAC1BA550703" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba latissima Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="280" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="277" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C5619849356063CDC0176C86CCDB5093" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AF8EA03A24C243196CCF42E1919CE9A1">
<mods:title id="34BB81A7FCC014AFBC39FDA2450E2684">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
</mods:titleInfo>
<mods:name id="8441C1E88CFD3A37C98E9DAFE3C53499" type="personal">
<mods:role id="3DE3E512A640EB92196B1F937EBB5E16">
<mods:roleTerm id="E64AA570303CF91896E4FF9E1B8D51E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2D1360DC5101DD7125D64C30D3EA9B4A">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="7FD591063C595528D159AA5658B9214F">Zootaxa</mods:title>
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<mods:part id="51B393DC60BF96DEDED962CD7C80C68D">
<mods:date id="3D5CAE03FEB134C168DCF9D34EACE213">2024</mods:date>
<mods:detail id="FEE5FACB85F895D7726379DD3FD557FE" type="pubDate">
<mods:number id="21CE408C3E65021231B99EC95F058D95">2024-03-14</mods:number>
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<mods:number id="73CAE0D9A9E8B98D3C0A33FE4EC7B160">5424</mods:number>
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<mods:number id="7F29D20FFB0FA420526AF8B76F618BA2">3</mods:number>
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<mods:url id="11EB1A65AEC32C16FAA09F5F60A857E4">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
</mods:location>
<mods:classification id="23BE9277A95E3D3C560305FD937DBE11">journal article</mods:classification>
<mods:identifier id="53B99C0A706BC0852E88FFE9FA2844DC" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="9531E2F34317D0A42F750405A8227210" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CB31DC273347EFBC88581E0D2E457305" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="B65C104D647957209503E64A3A755277" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFF36D45D6DDDAC1BA550703" ID-DOI="http://doi.org/10.5281/zenodo.10821279" ID-Zenodo-Dep="10821279" LSID="urn:lsid:plazi:treatment:A15C878DFFF36D45D6DDDAC1BA550703" httpUri="http://treatment.plazi.org/id/A15C878DFFF36D45D6DDDAC1BA550703" lastPageId="29" lastPageNumber="280" pageId="26" pageNumber="277">
<subSubSection id="61EF6510FFF36D42D6DDDAC1BC1E0252" box="[151,537,1014,1041]" pageId="26" pageNumber="277" type="nomenclature">
<paragraph id="294A369BFFF36D42D6DDDAC1BC1E0252" blockId="26.[151,537,1014,1077]" box="[151,537,1014,1041]" pageId="26" pageNumber="277">
<heading id="720281F7FFF36D42D6DDDAC1BC1E0252" bold="true" box="[151,537,1014,1041]" fontSize="11" level="1" pageId="26" pageNumber="277" reason="1">
<emphasis id="1B81EA89FFF36D42D6DDDAC1BC1E0252" bold="true" box="[151,537,1014,1041]" pageId="26" pageNumber="277">
<taxonomicName id="EEF54D18FFF36D42D6DDDAC1BFD70252" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,464,1014,1041]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF36D42D6DDDAC1BFD70252" bold="true" box="[151,464,1014,1041]" italics="true" pageId="26" pageNumber="277">Paraphloeostiba latissima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFF36D42D79DDAC0BC1E0252" box="[471,537,1015,1041]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D42D6DDDD2CBF560276" box="[151,337,1051,1077]" pageId="26" pageNumber="277" type="description">
<paragraph id="294A369BFFF36D42D6DDDD2CBF560276" blockId="26.[151,537,1014,1077]" box="[151,337,1051,1077]" pageId="26" pageNumber="277">
(
<figureCitation id="B1CE2A1EFFF36D42D6D5DD2CBEF10276" box="[159,246,1051,1077]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="26" pageNumber="277">Figs 41</figureCitation>
,
<figureCitation id="B1CE2A1EFFF36D42D748DD2CBF4E0276" box="[258,329,1051,1077]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="26" pageNumber="277">68–74</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D42D6DDDD54BB1B033A" pageId="26" pageNumber="277" type="materials_examined">
<paragraph id="294A369BFFF36D42D6DDDD54BFAA0286" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D6DDDD54BF42023F" bold="true" box="[151,325,1122,1149]" pageId="26" pageNumber="277">Type material.</emphasis>
<typeStatus id="F64E8839FFF36D42D707DD54BFB2023E" box="[333,437,1123,1149]" pageId="26" pageNumber="277" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFF36D42D7F6DD54BFD6023E" box="[444,465,1123,1149]" count="1" pageId="26" pageNumber="277" type="male">♂</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFF36D42D42CDD54BCAD023E" box="[614,682,1123,1149]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="26" pageNumber="277">NEW</collectionCode>
<collectingCountry id="51E2760BFFF36D42D4FBDD54BD1C023E" box="[689,795,1123,1149]" name="Guinea-Bissau" pageId="26" pageNumber="277">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFF36D42D5CDDD54BA12023E" box="[903,1045,1122,1149]" pageId="26" pageNumber="277" value="1965-06-16">16. VI. 1965</date>
| flying in daylight’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF36D42D335DD54BB9F023E" box="[1407,1432,1123,1149]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="277">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFF36D42D428DDB0BCFC02E2" box="[610,763,1159,1185]" pageId="26" pageNumber="277" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF36D42D547DDB0BDCF02E2" authorityName="Steel" authorityYear="1960" box="[781,968,1159,1185]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF36D42D547DDB0BDCF02E2" box="[781,968,1159,1185]" italics="true" pageId="26" pageNumber="277">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF36D42D590DDB0BA3802E2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[986,1087,1159,1185]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF36D42D590DDB0BA3802E2" box="[986,1087,1159,1185]" italics="true" pageId="26" pageNumber="277">latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D20FDDB0BA8302E2" bold="true" box="[1093,1156,1159,1185]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D20FDDB0BA8302E2" box="[1093,1156,1159,1185]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF36D42D341DDB0BB2602E2" box="[1291,1313,1159,1185]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="277" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFF36D42D70DDD9CBFA60286" box="[327,417,1195,1221]" pageId="26" pageNumber="277">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFF36D42D68DDDF8BB1B033A" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<typeStatus id="F64E8839FFF36D42D68DDDF8BF3F02AA" box="[199,312,1231,1257]" pageId="26" pageNumber="277" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFF36D42D70CDDF8BF7402AA" box="[326,371,1231,1257]" count="1" pageId="26" pageNumber="277" type="male">1 ♂</specimenCount>
(underside of the card with handwritten ‘Okapo 1164’),
<specimenCount id="3FF3FD12FFF36D42D241DDF8BA4A02AA" box="[1035,1101,1231,1257]" count="2" pageId="26" pageNumber="277" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFF36D42D215DDF8BAF902AA" box="[1119,1278,1231,1257]" count="1" pageId="26" pageNumber="277" type="generic">one specimen</specimenCount>
without right antennomeres 3–11 dissected;
<specimenCount id="3FF3FD12FFF36D42D7A6DDC4BC82034E" box="[492,645,1267,1293]" count="1" pageId="26" pageNumber="277" type="generic">one specimen</specimenCount>
with underside of the card with handwritten ‘Okapo 1164’): same data as the holotype (
<collectionCode id="4FE4AE5EFFF36D42D719DC20BFAA0372" box="[339,429,1303,1329]" pageId="26" pageNumber="277">BMNH</collectionCode>
);
<specimenCount id="3FF3FD12FFF36D42D78BDC20BFEA0372" box="[449,493,1303,1329]" count="1" pageId="26" pageNumber="277" type="male">1 ♂</specimenCount>
(dissected),
<specimenCount id="3FF3FD12FFF36D42D437DC20BCAF0372" box="[637,680,1303,1329]" count="1" pageId="26" pageNumber="277" type="female">1 ♀</specimenCount>
(underside of the card with handwritten ‘90H’): ‘
<collectionCode id="4FE4AE5EFFF36D42D292DC20BB1B0372" box="[1240,1308,1303,1329]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="26" pageNumber="277">NEW</collectionCode>
<collectingCountry id="51E2760BFFF36D42D36FDC20BB880372" box="[1317,1423,1303,1329]" name="Guinea-Bissau" pageId="26" pageNumber="277">GUINEA</collectingCountry>
| Okapa,
<date id="5D4B105BFFF36D42D6A5DC0DBF870316" box="[239,384,1338,1365]" pageId="26" pageNumber="277" value="1965-09-25">25. IX. 1965</date>
, | refuse’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFF36D42D424DC0CBC800316" box="[622,647,1339,1365]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="277">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFF36D42D22BDC0CBABC0316" box="[1121,1211,1339,1365]" pageId="26" pageNumber="277">BMNH</collectionCode>
). All
<typeStatus id="F64E8839FFF36D42D2B2DC0CBB640316" box="[1272,1379,1339,1365]" pageId="26" pageNumber="277" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFF36D42D792DC68BC6F033A" box="[472,616,1375,1401]" pageId="26" pageNumber="277" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFF36D42D436DC68BD30033A" authorityName="Steel" authorityYear="1960" box="[636,823,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF36D42D436DC68BD30033A" box="[636,823,1375,1401]" italics="true" pageId="26" pageNumber="277">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFF36D42D501DC68BDB7033A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[843,944,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="277" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF36D42D501DC68BDB7033A" box="[843,944,1375,1401]" italics="true" pageId="26" pageNumber="277">latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF36D42D5FDDC68BDF1033A" bold="true" box="[951,1014,1375,1401]" pageId="26" pageNumber="277">
<taxonomicNameLabel id="00B257F2FFF36D42D5FDDC68BDF1033A" box="[951,1014,1375,1401]" pageId="26" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFF36D42D2CADC68BA91033A" box="[1152,1174,1375,1401]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="277" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF36D44D68DDCB5BDD701FC" lastPageId="28" lastPageNumber="279" pageId="26" pageNumber="277" type="description">
<paragraph id="294A369BFFF36D42D68DDCB5BEDE004A" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<emphasis id="1B81EA89FFF36D42D68DDCB5BF5E03DF" bold="true" box="[199,345,1410,1436]" pageId="26" pageNumber="277">Description.</emphasis>
Measurements (n=6): HW: 0.37–0.43; HL: 0.25–0.26;OL: 0.12–0.14; TL: 0.02; AL (
<typeStatus id="F64E8839FFF36D42D362DCB4BB9603DE" box="[1320,1425,1411,1437]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
): 0.60; PL: 0.30–0.36; PWmax: 0.46–0.53; PWmin: 0.42–0.48; ESL: 0.47–0.53; EW: 0.60–0.62; MTbL (
<typeStatus id="F64E8839FFF36D42D362DC90BB960382" box="[1320,1425,1447,1473]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFF36D42D76FDCFCBF8903A6" box="[293,398,1483,1509]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
): 0.16 (MTrL 1–4: 0.05; MTrL 5: 0.11); AW: 0.57–0.60; AedL: 0.30; BL: 1.67–2.10 (
<typeStatus id="F64E8839FFF36D42D37BDCFCBB9E03A6" box="[1329,1433,1483,1509]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
: 1.82).
</paragraph>
<paragraph id="294A369BFFF36D42D68DDF24BA3C0132" blockId="26.[151,1437,1122,2013]" pageId="26" pageNumber="277">
<materialsCitation id="999D3CC6FFF36D42D68DDF24BA300132" collectionCode="V" pageId="26" pageNumber="277" specimenCount="9" typeStatus="holotype">
Habitus as in
<figureCitation id="B1CE2A1EFFF36D42D72FDF24BFBD006E" box="[357,442,1555,1581]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="26" pageNumber="277">Fig. 41</figureCitation>
. Body reddish brown, with yellowish to yellow-brown elytra, with darker medioapical, laterobasal and basal portions (laterobasal portions of pronotum of
<typeStatus id="F64E8839FFF36D42D5CBDF00BDE40012" box="[897,995,1591,1617]" pageId="26" pageNumber="277" type="holotype">holotype</typeStatus>
and
<specimenCount id="3FF3FD12FFF36D42D252DF00BAB30012" box="[1048,1204,1591,1617]" count="2" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">two paratypes</specimenCount>
yellowish; abdomen slightly paler in
<specimenCount id="3FF3FD12FFF36D42D717DF6CBC150036" box="[349,530,1627,1653]" count="3" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">three paratypes</specimenCount>
); antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow. Head with fine punctation, denser in middle and sparser on infraorbital portions (
<specimenCount id="3FF3FD12FFF36D42D262DF48BABD00DA" box="[1064,1210,1663,1689]" count="1" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">one paratype</specimenCount>
with very indistinct punctation); neck without visible punctures; pronotum with fine and moderately dense punctation, similar to that in middle portion of head, finer and sparser in middle; elytra with fine punctation, slightly denser and deeper than that in pronotum, finer around scutellum and along suture; abdominal tergites with very fine and moderately dense punctation, indistinct in abdominal tergites IV–
<collectionCode id="4FE4AE5EFFF36D42D4FBDE38BCC2016A" box="[689,709,1807,1833]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="277" type="Museum">V</collectionCode>
in
<specimenCount id="3FF3FD12FFF36D42D4A1DE38BD9D016A" box="[747,922,1807,1833]" count="3" pageId="26" pageNumber="277" type="generic" typeStatus="paratype">three paratypes</specimenCount>
. Head with dense and coarse microsculpture, transverse on vertex, coarser and distinctly isodiametric in middle, isodiametric and elongate on infraorbital portions; microsculpture of elytra very dense, about as that in pronotum, but slightly coarser
</materialsCitation>
.
</paragraph>
<paragraph id="294A369BFFF36D43D68DDE4CBA1E06BA" blockId="26.[151,1437,1122,2013]" lastBlockId="27.[151,1437,151,609]" lastPageId="27" lastPageNumber="278" pageId="26" pageNumber="277">Head with slightly elevated middle and infraorbital portions, with deep and moderately wide anteriomedian depressions, 1.4–1.5 times as broad as head, with narrow nad deep linear anteocellar foveae, almost reaching level of middle length of eyes; postocular carina absent. Ocelli moderately large, located at level of posterior margins of eyes; distance between ocelli about as long as distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about 1.4 times as long as preceding segment. Antennomere 6 slightly longer and significantly broader than 5, 7–9 slightly longer and broader than 6, 10 slightly longer than 9.</paragraph>
<paragraph id="294A369BFFF26D43D68DD834BD140702" blockId="27.[151,1437,151,609]" pageId="27" pageNumber="278">Pronotum 1.4–1.5 times as broad as long, 1.2 times as broad as head, widest in or slightly above middle, strongly narrowed posteriad toward subacute hing angle.</paragraph>
<paragraph id="294A369BFFF26D43D68DD87CBDFA0726" blockId="27.[151,1437,151,609]" box="[199,1021,330,357]" pageId="27" pageNumber="278">Elytra 1.1–1.2 times as broad as long, 1.4–1.5 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFF26D43D68DD858BC8507CA" blockId="27.[151,1437,151,609]" box="[199,642,367,393]" pageId="27" pageNumber="278">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="294A369BFFF26D43D68DD8A4BDB00422" blockId="27.[151,1437,151,609]" pageId="27" pageNumber="278">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFF26D43D51ED8A4BDA007EE" box="[852,935,403,429]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 70</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFF26D43D794D880BC340792" box="[478,563,439,465]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 71</figureCitation>
). Aedeagus with wide median lobe, indistinctly narrowed to widely rounded apex; parameres moderately short, significantly not reaching apex of median lobe, with narrow apical portion, with two long apical and two to three preapical setae, and two small setae in middle part of inner margin of each paramere; internal sac wide and moderately short, with two narrow and moderately long sclerotized structures in preapical portion (
<figureCitation id="B1CE2A1EFFF26D43D72FDB70BFBE0422" box="[357,441,583,609]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 68</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFF26D43D52ADB70BDB40422" box="[864,947,583,609]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278">Fig. 69</figureCitation>
.
</paragraph>
<caption id="7D8A6613FFF26D43D6DDDE4EBC80019A" ID-DOI="http://doi.org/10.5281/zenodo.10821251" ID-Zenodo-Dep="10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="27" pageNumber="278" startId="27.[151,264,1913,1937]" targetBox="[172,1428,642,1882]" targetPageId="27" targetType="figure">
<paragraph id="294A369BFFF26D43D6DDDE4EBC80019A" blockId="27.[151,1436,1913,2010]" pageId="27" pageNumber="278">
<emphasis id="1B81EA89FFF26D43D6DDDE4EBF5101D2" bold="true" box="[151,342,1913,1937]" pageId="27" pageNumber="278">FIGURES 68–74.</emphasis>
<taxonomicName id="EEF54D18FFF26D43D717DE4DBC6D01D1" authorityName="Shavrin" authorityYear="2024" box="[349,618,1914,1938]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="278" phylum="Arthropoda" rank="species" species="latissima">
<emphasis id="1B81EA89FFF26D43D717DE4DBC6D01D1" box="[349,618,1914,1938]" italics="true" pageId="27" pageNumber="278">Paraphloeostiba latissima</emphasis>
</taxonomicName>
(paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="7D8A6613FFF56D44D6DDDF09BA920039" ID-DOI="http://doi.org/10.5281/zenodo.10821253" ID-Zenodo-Dep="10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="28" pageNumber="279" startId="28.[151,264,1598,1622]" targetBox="[212,1375,188,1560]" targetPageId="28" targetType="figure">
<paragraph id="294A369BFFF56D44D6DDDF09BA920039" blockId="28.[151,1436,1598,1659]" pageId="28" pageNumber="279">
<emphasis id="1B81EA89FFF56D44D6DDDF09BF5C0015" bold="true" box="[151,347,1598,1622]" pageId="28" pageNumber="279">FIGURES 75–80.</emphasis>
Habitus of
<taxonomicName id="EEF54D18FFF56D44D7ACDF09BC880015" authorityName="Steel" authorityYear="1960" box="[486,655,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFF56D44D7ACDF09BC880015" box="[486,655,1598,1622]" italics="true" pageId="28" pageNumber="279">Paraphloeostiba</emphasis>
</taxonomicName>
: 75–
<taxonomicName id="EEF54D18FFF56D44D480DF08BD4A0015" authorityName="Shavrin" authorityYear="2024" box="[714,845,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="okapensis">
<emphasis id="1B81EA89FFF56D44D480DF08BD4A0015" box="[714,845,1598,1622]" italics="true" pageId="28" pageNumber="279">P. okapensis</emphasis>
</taxonomicName>
(paratype), 76–
<taxonomicName id="EEF54D18FFF56D44D5B6DF08BA880015" authorityName="Shavrin" authorityYear="2024" box="[1020,1167,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="penelopeae">
<emphasis id="1B81EA89FFF56D44D5B6DF08BA880015" box="[1020,1167,1598,1622]" italics="true" pageId="28" pageNumber="279">P. penelopeae</emphasis>
</taxonomicName>
(paratype), 77–
<taxonomicName id="EEF54D18FFF56D44D374DF08BB9B0015" authorityName="Shavrin" authorityYear="2024" box="[1342,1436,1598,1622]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="1B81EA89FFF56D44D374DF08BB9B0015" box="[1342,1436,1598,1622]" italics="true" pageId="28" pageNumber="279">P. pilosa</emphasis>
</taxonomicName>
(holotype), 78–
<taxonomicName id="EEF54D18FFF56D44D779DF54BF890039" authorityName="Shavrin" authorityYear="2024" box="[307,398,1634,1658]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="1B81EA89FFF56D44D779DF54BF890039" box="[307,398,1634,1658]" italics="true" pageId="28" pageNumber="279">P. riedeli</emphasis>
</taxonomicName>
(paratype), 79–
<taxonomicName id="EEF54D18FFF56D44D465DF54BCC20039" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[559,709,1634,1658]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFF56D44D465DF54BCC20039" box="[559,709,1634,1658]" italics="true" pageId="28" pageNumber="279">P. rhopalocera</emphasis>
</taxonomicName>
, 80–
<taxonomicName id="EEF54D18FFF56D44D4B2DF54BD480039" authorityName="Shavrin" authorityYear="2024" box="[760,847,1634,1658]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="rufula">
<emphasis id="1B81EA89FFF56D44D4B2DF54BD480039" box="[760,847,1634,1658]" italics="true" pageId="28" pageNumber="279">P. rufula</emphasis>
</taxonomicName>
(holotype). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFF56D44D68DDFFABCCB0148" blockId="28.[151,1437,1741,1983]" pageId="28" pageNumber="279">
Female. Posterior margin of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFF56D44D56EDFFABD7F00A4" box="[804,888,1741,1767]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="28" pageNumber="279">Fig. 72</figureCitation>
) and sternite VIII (
<figureCitation id="B1CE2A1EFFF56D44D213DFFABAAA00A4" box="[1113,1197,1741,1767]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="28" pageNumber="279">Fig. 73</figureCitation>
) truncate. Accessory sclerite as in
<figureCitation id="B1CE2A1EFFF56D44D760DFC6BF7A0148" box="[298,381,1777,1803]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="28" pageNumber="279">Fig. 74</figureCitation>
. Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFF56D44D68DDE23BDD701FC" blockId="28.[151,1437,1741,1983]" pageId="28" pageNumber="279">
<emphasis id="1B81EA89FFF56D44D68DDE23BFB7016C" bold="true" box="[199,432,1812,1839]" pageId="28" pageNumber="279">Comparative notes.</emphasis>
Based on the presence of longitudinal impressions on the pronotum and the general shape of the aedeagus with narrow and short parameres,
<taxonomicName id="EEF54D18FFF56D44D4AADE0EBD640110" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[736,867,1849,1875]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="latissima" status="sp. n.">
<emphasis id="1B81EA89FFF56D44D4AADE0EBD640110" box="[736,867,1849,1875]" italics="true" pageId="28" pageNumber="279">P. latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF56D44D527DE0EBDA80110" bold="true" box="[877,943,1849,1875]" pageId="28" pageNumber="279">
<taxonomicNameLabel id="00B257F2FFF56D44D527DE0EBDA80110" box="[877,943,1849,1875]" pageId="28" pageNumber="279" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is somewhat similar to
<taxonomicName id="EEF54D18FFF56D44D28DDE0EBED50134" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="279" phylum="Arthropoda" rank="species" species="cooteri">
<emphasis id="1B81EA89FFF56D44D28DDE0EBB330111" box="[1223,1332,1849,1874]" italics="true" pageId="28" pageNumber="279">P. cooteri</emphasis>
Shavrin, 2020
</taxonomicName>
, recently described from
<collectingRegion id="EB31F879FFF56D44D7A6DE6ABC540134" box="[492,595,1885,1911]" country="China" name="Zhejiang" pageId="28" pageNumber="279">Zhejiang</collectingRegion>
,
<collectingCountry id="51E2760BFFF56D44D417DE6ABCA70134" box="[605,672,1885,1911]" name="China" pageId="28" pageNumber="279">China</collectingCountry>
(Shavrin 2020). The new species can be distinguished from it by the less transverse pronotum, finer punctation of the head and the pronotum, broader apical portion of the median lobe, narrower and shorter parameres and different structure of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF46D45D68DD9A1BF2B0697" pageId="29" pageNumber="280" type="distribution">
<paragraph id="294A369BFFF46D45D68DD9A1BF2B0697" blockId="29.[151,1436,150,321]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D68DD9A1BF6606F3" bold="true" box="[199,353,150,176]" pageId="29" pageNumber="280">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFF46D45D723D9A1BC9506F3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[361,658,150,176]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="280" phylum="Arthropoda" rank="species" species="latissima" status="sp. nov.">
<emphasis id="1B81EA89FFF46D45D723D9A1BC9506F3" box="[361,658,150,176]" italics="true" pageId="29" pageNumber="280">Paraphloeostiba latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFF46D45D4D1D9A1BCDB06F3" bold="true" box="[667,732,150,176]" pageId="29" pageNumber="280">
<taxonomicNameLabel id="00B257F2FFF46D45D4D1D9A1BCDB06F3" box="[667,732,150,176]" pageId="29" pageNumber="280" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFF46D45D5B0D9A0BA2D06F2" box="[1018,1066,151,177]" pageId="29" pageNumber="280">type</typeStatus>
locality in central part of
<collectingCountry id="51E2760BFFF46D45D312D9A0BF2F0697" name="Papua New Guinea" pageId="29" pageNumber="280">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFF46D45D68DD9E9BA550703" pageId="29" pageNumber="280" type="etymology">
<paragraph id="294A369BFFF46D45D68DD9E9BF4A075F" blockId="29.[151,1436,150,321]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D68DD9E9BF4906BB" bold="true" box="[199,334,222,248]" pageId="29" pageNumber="280">Etymology.</emphasis>
The specific epithet is the Latin
<emphasis id="1B81EA89FFF46D45D489D9E9BD3406BB" box="[707,819,222,248]" italics="true" pageId="29" pageNumber="280">latissimus</emphasis>
, -
<emphasis id="1B81EA89FFF46D45D501D9E8BD9406BB" box="[843,915,223,248]" italics="true" pageId="29" pageNumber="280">a, -um</emphasis>
(broadest). It alludes to the wide median lobe of the aedeagus.
</paragraph>
<paragraph id="294A369BFFF46D45D68DD811BA550703" blockId="29.[151,1436,150,321]" box="[199,1106,294,321]" pageId="29" pageNumber="280">
<emphasis id="1B81EA89FFF46D45D68DD811BF4C0703" bold="true" box="[199,331,294,320]" pageId="29" pageNumber="280">Bionomics.</emphasis>
Specimens were collected flying in daylight or were found in refuse.
</paragraph>
</subSubSection>
</treatment>
</document>
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