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<document id="584473CFDE854607220DBF38288A4FE5" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFE56D56D6DDDA99BF00021A" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba barclayi Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="265" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="263" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5278A3E2CBC0C7EB930F8E6E0CE0DA34" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="70F745BDA31BB770D128E2FD4B40856C">
<mods:title id="E61F6FD3B0A80786AF3C213FC01B3BF4">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
</mods:titleInfo>
<mods:name id="368AEBB7E5DF6585CA3F38738A56D2A0" type="personal">
<mods:role id="8FF7FC598EF5C3D4C919B1B2F1AD0AF6">
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</mods:role>
<mods:namePart id="7D7001FB64D281661CEEA6952A7173B6">Shavrin, Alexey V.</mods:namePart>
</mods:name>
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<mods:title id="74B731E695B33C5E7675C4B5A96E4109">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5651378A27F6F1117BDB6EB1F2BD3924">
<mods:date id="68F1FF8F3732AE785E8AF6F72E427B20">2024</mods:date>
<mods:detail id="3BF826AF1E07965EB8D74EE346207F68" type="pubDate">
<mods:number id="5C6B3699CC25EA87FD721524B50DA042">2024-03-14</mods:number>
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<mods:number id="5278E876CBA0B88AD395EBC86E372642">5424</mods:number>
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<mods:detail id="705CFD873FFA4E68A5CB6D1F5C7531D7" type="issue">
<mods:number id="960D64B2201004E1964D575ADEAD528D">3</mods:number>
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<mods:url id="931A19B5AF2A168A20F949835AB7C702">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
</mods:location>
<mods:classification id="EB859167993AF90226E70E8128EC8F05">journal article</mods:classification>
<mods:identifier id="B45E4651D4EBEA89993E4889284FF6FE" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="EBED71EA82B1B354FD96E2DFF4B7C98C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AE72C5FD10C147F3C9D24DE9D3048EAD" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="FAB6C643E35F578CE1B24BC4D449127D" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFE56D56D6DDDA99BF00021A" LSID="urn:lsid:plazi:treatment:A15C878DFFE56D56D6DDDA99BF00021A" httpUri="http://treatment.plazi.org/id/A15C878DFFE56D56D6DDDA99BF00021A" lastPageId="14" lastPageNumber="265" pageId="12" pageNumber="263">
<subSubSection id="61EF6510FFE56D54D6DDDA99BC08058A" box="[151,527,942,969]" pageId="12" pageNumber="263" type="nomenclature">
<paragraph id="294A369BFFE56D54D6DDDA99BC08058A" blockId="12.[151,527,942,1005]" box="[151,527,942,969]" pageId="12" pageNumber="263">
<heading id="720281F7FFE56D54D6DDDA99BC08058A" bold="true" box="[151,527,942,969]" fontSize="11" level="1" pageId="12" pageNumber="263" reason="1">
<emphasis id="1B81EA89FFE56D54D6DDDA99BC08058A" bold="true" box="[151,527,942,969]" pageId="12" pageNumber="263">
<taxonomicName id="EEF54D18FFE56D54D6DDDA99BFC1058A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,454,942,969]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE56D54D6DDDA99BFC1058A" bold="true" box="[151,454,942,969]" italics="true" pageId="12" pageNumber="263">Paraphloeostiba barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFE56D54D787DA98BC08058A" box="[461,527,943,969]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D54D6DDDAE4BF4405AE" box="[151,323,978,1005]" pageId="12" pageNumber="263" type="description">
<paragraph id="294A369BFFE56D54D6DDDAE4BF4405AE" blockId="12.[151,527,942,1005]" box="[151,323,978,1005]" pageId="12" pageNumber="263">
(
<figureCitation id="B1CE2A1EFFE56D54D6D5DAE4BEEF05AE" box="[159,232,978,1005]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="12" pageNumber="263">Figs 5</figureCitation>
,
<figureCitation id="B1CE2A1EFFE56D54D6BEDAE4BF3B05AE" box="[244,316,979,1005]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="12" pageNumber="263">26–31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D54D6DDDD2CBCCD02AA" pageId="12" pageNumber="263" type="materials_examined">
<paragraph id="294A369BFFE56D54D6DDDD2CBA46023E" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D6DDDD2CBF4D0277" bold="true" box="[151,330,1050,1077]" pageId="12" pageNumber="263">Type material.</emphasis>
<typeStatus id="F64E8839FFE56D54D71CDD2CBFB90276" box="[342,446,1051,1077]" pageId="12" pageNumber="263" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFE56D54D780DD2CBFD80276" box="[458,479,1051,1077]" count="1" pageId="12" pageNumber="263" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFE56D54D7B9DD2CBC3B0276" box="[499,572,1050,1077]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="12" pageNumber="263">Fig. 5</figureCitation>
; right antennomeres 3–11 missing; dissected): ‘
<collectionCode id="4FE4AE5EFFE56D54D226DD2CBAB70276" box="[1132,1200,1051,1077]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="263">NEW</collectionCode>
<collectingCountry id="51E2760BFFE56D54D2F6DD2CBB210276" box="[1212,1318,1051,1077]" name="Guinea-Bissau" pageId="12" pageNumber="263">GUINEA</collectingCountry>
| Awande (ca.
<geoCoordinate id="4CC1505CFFE56D54D684DD08BF1B021A" box="[206,284,1087,1113]" degrees="6" direction="south" minutes="31" orientation="latitude" pageId="12" pageNumber="263" precision="925" value="-6.5166664">6°31’S</geoCoordinate>
<geoCoordinate id="4CC1505CFFE56D54D760DD09BF9E021A" box="[298,409,1086,1113]" degrees="145" direction="east" minutes="34" orientation="longitude" pageId="12" pageNumber="263" precision="925" value="145.56667">145°34’E</geoCoordinate>
), |
<date id="5D4B105BFFE56D54D782DD09BC4D021A" box="[456,586,1086,1113]" pageId="12" pageNumber="263" value="1965-11-21">21.XI.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE56D54D4ACDD08BCF8021A" box="[742,767,1087,1113]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="263">R</collectionCode>
. Hornabrook BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFE56D54D2BADD08BB8E021A" box="[1264,1417,1087,1113]" pageId="12" pageNumber="263" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE56D54D6DDDD54BF55023E" authorityName="Steel" authorityYear="1960" box="[151,338,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE56D54D6DDDD54BF55023E" box="[151,338,1123,1149]" italics="true" pageId="12" pageNumber="263">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE56D54D72FDD54BFC6023E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[357,449,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE56D54D72FDD54BFC6023E" box="[357,449,1123,1149]" italics="true" pageId="12" pageNumber="263">barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE56D54D783DD54BC0F023E" bold="true" box="[457,520,1123,1149]" pageId="12" pageNumber="263">
<taxonomicNameLabel id="00B257F2FFE56D54D783DD54BC0F023E" box="[457,520,1123,1149]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE56D54D4DBDD54BCA0023E" box="[657,679,1123,1149]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="263" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFE56D54D591DD54BA32023E" box="[987,1077,1123,1149]" pageId="12" pageNumber="263">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFE56D54D68DDDB0BCCD02AA" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
<typeStatus id="F64E8839FFE56D54D68DDDB0BF2E02E2" box="[199,297,1159,1185]" pageId="12" pageNumber="263" type="paratype">Paratype</typeStatus>
<specimenCount id="3FF3FD12FFE56D54D778DDB0BF4002E2" box="[306,327,1159,1185]" count="1" pageId="12" pageNumber="263" type="female">♀</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFE56D54D797DDB0BC2602E2" box="[477,545,1159,1185]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="263">NEW</collectionCode>
<collectingCountry id="51E2760BFFE56D54D460DDB0BC9302E2" box="[554,660,1159,1185]" name="Guinea-Bissau" pageId="12" pageNumber="263">GUINEA</collectingCountry>
| Border Ranges, Miarosa | (
<quantity id="EE0D9B7EFFE56D54D597DDB0BA2802E2" box="[989,1071,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="263" unit="m" value="1900.0">1900m</quantity>
, ca.
<geoCoordinate id="4CC1505CFFE56D54D22EDDB0BAD802E2" box="[1124,1247,1159,1185]" degrees="6" direction="south" minutes="31" orientation="latitude" pageId="12" pageNumber="263" precision="15" seconds="10" value="-6.519444">6°31’10’’S</geoCoordinate>
|
<geoCoordinate id="4CC1505CFFE56D54D2BCDDB1BB8902E2" box="[1270,1422,1158,1185]" degrees="145" direction="east" minutes="33" orientation="longitude" pageId="12" pageNumber="263" precision="15" seconds="00" value="145.55">145°33’00’’E</geoCoordinate>
|
<date id="5D4B105BFFE56D54D6DDDD9DBF1D0286" box="[151,282,1194,1221]" pageId="12" pageNumber="263" value="1965-11-16">16.XI.1965</date>
’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFE56D54D7ECDD9CBFB80286" box="[422,447,1195,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="263">R</collectionCode>
. Hornabrook
<collectionCode id="4FE4AE5EFFE56D54D412DD9CBC830286" box="[600,644,1195,1221]" country="United Kingdom" name="Bristol Museum" pageId="12" pageNumber="263">BM</collectionCode>
1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFE56D54D5C1DD9CBA1C0286" box="[907,1051,1195,1221]" pageId="12" pageNumber="263" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFE56D54D264DD9CBAEE0286" authorityName="Steel" authorityYear="1960" box="[1070,1257,1195,1221]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFE56D54D264DD9CBAEE0286" box="[1070,1257,1195,1221]" italics="true" pageId="12" pageNumber="263">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFE56D54D2B1DD9CBB500286" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1275,1367,1195,1221]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE56D54D2B1DD9CBB500286" box="[1275,1367,1195,1221]" italics="true" pageId="12" pageNumber="263">barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE56D54D314DD9CBB9A0286" bold="true" box="[1374,1437,1195,1221]" pageId="12" pageNumber="263">
<taxonomicNameLabel id="00B257F2FFE56D54D314DD9CBB9A0286" box="[1374,1437,1195,1221]" pageId="12" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFE56D54D750DDF8BF3702AA" box="[282,304,1231,1257]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="263" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFE56D54D429DDF8BCBA02AA" box="[611,701,1231,1257]" pageId="12" pageNumber="263">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE56D56D68DDDC5BFDA05C2" lastPageId="14" lastPageNumber="265" pageId="12" pageNumber="263" type="description">
<paragraph id="294A369BFFE56D54D68DDDC5BA720316" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
<emphasis id="1B81EA89FFE56D54D68DDDC5BF5E034F" bold="true" box="[199,345,1266,1292]" pageId="12" pageNumber="263">Description.</emphasis>
Measurements (n=2): HW: 0.35; HL: 0.20–0.21; OL: 0.11–0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFE56D54D2ADDDC4BB57034E" box="[1255,1360,1267,1293]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
): 0.52; PL: 0.25–0.27; PW: 0.42–0.45; ESL: 0.40–0.42; EW: 0.52–0.55; MTbL (
<typeStatus id="F64E8839FFE56D54D591DC20BA430372" box="[987,1092,1303,1329]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
): 0.27; MTrL (
<typeStatus id="F64E8839FFE56D54D2A4DC20BB500372" box="[1262,1367,1303,1329]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
): 0.12 (MTrL 1–4: 0.05; MTrL 5: 0.07); AW: 0.50–0.52; AedL: 0.35; BL: 1.62(
<typeStatus id="F64E8839FFE56D54D58BDC0CBA2E0316" box="[961,1065,1339,1365]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
)–1.79.
</paragraph>
<paragraph id="294A369BFFE56D54D68DDC68BB5B00DA" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">
Habitus as in
<figureCitation id="B1CE2A1EFFE56D54D722DC68BFA9033A" box="[360,430,1374,1401]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="12" pageNumber="263">Fig. 5</figureCitation>
. Head, pronotum and abdomen reddish-brown (pronotum slightly paler, with yellowish lateroapical portions in
<typeStatus id="F64E8839FFE56D54D7F8DCB4BC1E03DE" box="[434,537,1411,1437]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
); elytra yellowish-brown, with darkened mediobasal, lateroapical and apical portions; antennomeres 5–11 brown; mouthparts, antennomeres 1–4 and legs yellow. Head with irregular sparse and relatively large punctation, denser in middle; neck without visible punctation; pronotum with relatively large and dense punctation, similar to that in middle portion of head, larger and deeper in lateral and finer in mediobasal portions; punctation of elytra similar to that in pronotum, but distinctly finer and sparser in middle portions of each elytron along suture. Head with distinct dense microsculpture: transverse on vertex, subdiagonal in middle and longitudinal in infraorbital portions; pronotum with dense transverse microreticulation, finer in middle; elytra with dense subdiagonal microreticulation, finer along suture; abdominal tergites with very fine transverse meshes.
</paragraph>
<paragraph id="294A369BFFE56D54D68DDF94BF3A01FA" blockId="12.[151,1437,1050,1977]" pageId="12" pageNumber="263">Head with distinct narrow and moderately deep anteriomedian depression, 1.6 times as broad as long, with short, narrow and relatively deep anteocellar foveae, subdiagonally stretching posteriad and reaching level of of anterior third length of eyes; postocular carina absent. Ocelli moderately large, located at level of posterior margins of eyes; distance between ocelli slightly larger than distance between ocellus and posterior margin of eyes. Antennomere 3 distinctly shorter and narrower than 2, 4 about twice as long as 3, 5 slightly broader than 4, 6 slightly longer and distinctly broader than 5, 7–9 slightly longer and broader than 6, 10 indistinctly longer than 9, apical antennomere about as long combined length of two preceeding segments, from about middle strongly narrowed apicad toward subacute apex.</paragraph>
<caption id="7D8A6613FFE46D55D6DDDF18BF5000CC" ID-DOI="http://doi.org/10.5281/zenodo.10821229" ID-Zenodo-Dep="10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="13" pageNumber="264" startId="13.[151,264,1583,1607]" targetBox="[187,1428,185,1557]" targetPageId="13" targetType="figure">
<paragraph id="294A369BFFE46D55D6DDDF18BF5000CC" blockId="13.[151,1437,1583,1680]" pageId="13" pageNumber="264">
<emphasis id="1B81EA89FFE46D55D6DDDF18BF50000B" bold="true" box="[151,343,1583,1608]" pageId="13" pageNumber="264">FIGURES 26–31.</emphasis>
<taxonomicName id="EEF54D18FFE46D55D72ADF07BC61000B" authorityName="Shavrin" authorityYear="2024" box="[352,614,1584,1608]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="264" phylum="Arthropoda" rank="species" species="barclayi">
<emphasis id="1B81EA89FFE46D55D72ADF07BC61000B" box="[352,614,1584,1608]" italics="true" pageId="13" pageNumber="264">Paraphloeostiba barclayi</emphasis>
</taxonomicName>
(holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFE46D55D68DDFD5BFD80107" blockId="13.[151,1437,1762,2040]" pageId="13" pageNumber="264">Pronotum 1.6 times as broad as long, 1.2 times as broad as head, widest in middle, more narrowed posteriad than anteriad; lateral impressions moderately deep, distinctly wider and deeper in laterobasal portions; lateral margins with fine regular crenulation.</paragraph>
<paragraph id="294A369BFFE46D55D68DDE79BB9A012B" blockId="13.[151,1437,1762,2040]" box="[199,1437,1870,1897]" pageId="13" pageNumber="264">Elytra 1.3 times as broad as long, 1.5–1.6 times as long as pronotum, with narrowly impressed lateral margins.</paragraph>
<paragraph id="294A369BFFE46D55D68DDE45BD3301CF" blockId="13.[151,1437,1762,2040]" box="[199,820,1906,1932]" pageId="13" pageNumber="264">Metatarsi slightly more than twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFE46D55D68DDEA1BC8501F3" blockId="13.[151,1437,1762,2040]" box="[199,642,1942,1968]" pageId="13" pageNumber="264">Abdomen slightly narrower than elytra.</paragraph>
<paragraph id="294A369BFFE46D56D68DDE8DBC86075E" blockId="13.[151,1437,1762,2040]" lastBlockId="14.[151,1437,151,1113]" lastPageId="14" lastPageNumber="265" pageId="13" pageNumber="264">
Male. Posterior margin of abdominal tergite VIII rounded (
<figureCitation id="B1CE2A1EFFE46D55D51FDE8DBDAF0197" box="[853,936,1978,2004]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="13" pageNumber="264">Fig. 28</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFE46D55D748DEE9BF5001BB" box="[258,343,2014,2040]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="13" pageNumber="264">Fig. 29</figureCitation>
). Aedeagus with wide basal portion, strongly narrowed toward narrow and long median lobe with small rounded apex; parameres narrow, distinctly exceeding apex of median lobe and distinctly broadened apically, each with two long apical and two preapical setae and with additional three short setae in about middle of inner margin; internal sac moderately long and narrow, with large elongate sclerotized structure in basal portion (
<figureCitation id="B1CE2A1EFFE76D56D376D9E8BB9706BA" box="[1340,1424,223,249]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 26</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFE76D56D461D834BC7A075E" box="[555,637,259,285]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 27</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFE76D56D68DD810BD2B0726" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
Female. Posterior margins of abdominal tergite VIII (
<figureCitation id="B1CE2A1EFFE76D56D562D810BD7C0702" box="[808,891,295,321]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 30</figureCitation>
) and abdominal sternite VIII (
<figureCitation id="B1CE2A1EFFE76D56D29DD810BB2D0702" box="[1239,1322,295,321]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 31</figureCitation>
) truncate. Female accessory sclerite and spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFE76D56D68DD858BA3107EE" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFE76D56D537D858BDD407CA" box="[893,979,367,393]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 30</figureCitation>
). Posterior margin of abdominal sternite VIII widely rounded (
<figureCitation id="B1CE2A1EFFE76D56D7C7D8A4BFE707EE" box="[397,480,403,429]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="14" pageNumber="265">Fig. 31</figureCitation>
). Accessory sclerite and spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFE76D56D68DD881BCFC045A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DD881BFB60792" bold="true" box="[199,433,438,465]" pageId="14" pageNumber="265">Comparative notes.</emphasis>
Based on the very narrow median lobe and the parameres,
<taxonomicName id="EEF54D18FFE76D56D21AD880BACF0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1104,1224,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D21AD880BACF0792" box="[1104,1224,439,465]" italics="true" pageId="14" pageNumber="265">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D29AD880BB170792" bold="true" box="[1232,1296,439,465]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D29AD880BB170792" box="[1232,1296,439,465]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFE76D56D6DDD8ECBF4C07B7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,331,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D6DDD8ECBEAF07B7" box="[151,168,475,500]" italics="true" pageId="14" pageNumber="265">P</emphasis>
.
<emphasis id="1B81EA89FFE76D56D6FDD8ECBF4C07B7" box="[183,331,475,500]" italics="true" pageId="14" pageNumber="265">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D718D8ECBF9607B6" bold="true" box="[338,401,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D718D8ECBF9607B6" box="[338,401,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFE76D56D7D5D8ECBC2C07B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[415,555,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D7D5D8ECBC2C07B6" box="[415,555,475,501]" italics="true" pageId="14" pageNumber="265">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D478D8ECBC7507B6" bold="true" box="[562,626,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D478D8ECBC7507B6" box="[562,626,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFE76D56D435D8ECBD1207B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[639,789,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D435D8ECBD1207B6" box="[639,789,475,501]" italics="true" pageId="14" pageNumber="265">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D556D8ECBD5B07B6" bold="true" box="[796,860,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D556D8ECBD5B07B6" box="[796,860,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="EEF54D18FFE76D56D5D3D8ECBDF707B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[921,1008,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D5D3D8ECBDF707B6" box="[921,1008,475,501]" italics="true" pageId="14" pageNumber="265">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D5BDD8ECBA3007B6" bold="true" box="[1015,1079,475,501]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D5BDD8ECBA3007B6" box="[1015,1079,475,501]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFE76D56D68DDB14BF150422" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D74DDB14BFBA047F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[263,445,547,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D74DDB14BF1F047F" box="[263,280,547,572]" italics="true" pageId="14" pageNumber="265">P</emphasis>
.
<emphasis id="1B81EA89FFE76D56D763DB14BFBA047F" box="[297,445,547,572]" italics="true" pageId="14" pageNumber="265">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D78CDB14BC0F047E" bold="true" box="[454,520,547,573]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D78CDB14BC0F047E" box="[454,520,547,573]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller body, the paler elytra, slightly broader pronotum and the longer parameres;
</paragraph>
<paragraph id="294A369BFFE76D56D68DDB5CBDC504EA" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D74EDB5CBF8904C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,398,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D74EDB5CBF8904C6" box="[260,398,619,645]" italics="true" pageId="14" pageNumber="265">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D7DEDB5CBFD404C6" bold="true" box="[404,467,619,645]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D7DEDB5CBFD404C6" box="[404,467,619,645]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the larger body, the paler elytra, the presence of the punctation on the head, the finer punctation on the pronotum, the shorter elytra and the longer parameres;
</paragraph>
<paragraph id="294A369BFFE76D56D68DDB84BA8C04B2" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D749DB84BF90048E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[259,407,691,717]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D749DB84BF90048E" box="[259,407,691,717]" italics="true" pageId="14" pageNumber="265">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D7D7DB84BFDC048E" bold="true" box="[413,475,691,717]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D7D7DB84BFDC048E" box="[413,475,691,717]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker head, pronotum and abdomen, the presence of the punctation of the head, sparser and larger punctation of the pronotum, the shorter elytra and the longer parameres;
</paragraph>
<paragraph id="294A369BFFE76D56D68DDBCCBC01057A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
from
<taxonomicName id="EEF54D18FFE76D56D74EDBCCBF5C0556" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,347,763,789]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D74EDBCCBF5C0556" box="[260,347,763,789]" italics="true" pageId="14" pageNumber="265">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D728DBCCBFA60556" bold="true" box="[354,417,763,789]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D728DBCCBFA60556" box="[354,417,763,789]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the paler elytra, the absence of elevations between punctures on the elytra and narrower apical portions of the parameres.
</paragraph>
<paragraph id="294A369BFFE76D56D68DDA74BFDA05C2" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
From all these species,
<taxonomicName id="EEF54D18FFE76D56D799DA74BC80051F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[467,647,835,861]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFE76D56D799DA74BFE3051F" box="[467,484,835,860]" italics="true" pageId="14" pageNumber="265">P</emphasis>
.
<emphasis id="1B81EA89FFE76D56D7B9DA74BC80051F" box="[499,647,835,860]" italics="true" pageId="14" pageNumber="265">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D4DADA74BCD6051E" bold="true" box="[656,721,835,861]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D4DADA74BCD6051E" box="[656,721,835,861]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the details of the external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE76D56D68DDABDBEF7058A" pageId="14" pageNumber="265" type="distribution">
<paragraph id="294A369BFFE76D56D68DDABDBEF7058A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DDABDBF6605E7" bold="true" box="[199,353,906,932]" pageId="14" pageNumber="265">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFE76D56D726DABCBC8905E6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[364,654,907,933]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="1B81EA89FFE76D56D726DABCBC8905E6" box="[364,654,907,933]" italics="true" pageId="14" pageNumber="265">Paraphloeostiba barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFE76D56D4D3DABCBCDA05E6" bold="true" box="[665,733,907,933]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="00B257F2FFE76D56D4D3DABCBCDA05E6" box="[665,733,907,933]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in central part of
<collectingCountry id="51E2760BFFE76D56D35DDABCBEEB058A" name="Papua New Guinea" pageId="14" pageNumber="265">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFE76D56D68DDAE4BF00021A" pageId="14" pageNumber="265" type="etymology">
<paragraph id="294A369BFFE76D56D68DDAE4BF0E0252" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DDAE4BF4905AE" bold="true" box="[199,334,979,1005]" pageId="14" pageNumber="265">Etymology.</emphasis>
Patronymic, the species is named to honour of well-known coleopterologist Maxwell V.L. Barclay (London).
</paragraph>
<paragraph id="294A369BFFE76D56D68DDD2DBF00021A" blockId="14.[151,1437,151,1113]" pageId="14" pageNumber="265">
<emphasis id="1B81EA89FFE76D56D68DDD2DBF4C0277" bold="true" box="[199,331,1050,1076]" pageId="14" pageNumber="265">Bionomics.</emphasis>
The
<typeStatus id="F64E8839FFE76D56D7DBDD2CBFF60276" box="[401,497,1051,1077]" pageId="14" pageNumber="265" type="paratype">paratype</typeStatus>
was collected at an elevation
<quantity id="EE0D9B7EFFE76D56D523DD2CBDC30276" box="[873,964,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="265" unit="m" value="1900.0">1900 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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