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<document id="DE67BC7E745D2D768744332B3F004D6C" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFDE6D6FD6DDDF54BB9B01A1" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xanthonomus Bernhauer 1926" docType="key" docVersion="3" lastPageNumber="306" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="306" updateTime="1710782037732" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="E0F07EDCA085A5050518854923C53A61">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<treatment id="A15C878DFFDE6D6FD6DDDF54BB9B01A1" LSID="urn:lsid:plazi:treatment:A15C878DFFDE6D6FD6DDDF54BB9B01A1" httpUri="http://treatment.plazi.org/id/A15C878DFFDE6D6FD6DDDF54BB9B01A1" lastPageNumber="306" pageId="55" pageNumber="306">
<subSubSection id="61EF6510FFDE6D6FD6DDDF54BC90003E" box="[151,663,1634,1661]" pageId="55" pageNumber="306" type="nomenclature">
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<heading id="720281F7FFDE6D6FD6DDDF54BC90003E" bold="true" box="[151,663,1634,1661]" fontSize="11" level="1" pageId="55" pageNumber="306" reason="1">
<emphasis id="1B81EA89FFDE6D6FD6DDDF54BC90003E" bold="true" box="[151,663,1634,1661]" pageId="55" pageNumber="306">
Key to species of the genus
<taxonomicName id="EEF54D18FFDE6D6FD7A6DF55BC90003E" ID-CoL="8XQQ7" authorityName="Bernhauer" authorityYear="1926" box="[492,663,1634,1661]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFDE6D6FD7A6DF55BC90003E" bold="true" box="[492,663,1634,1661]" italics="true" pageId="55" pageNumber="306">Xanthonomus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFDE6D6FD6DDDF9FBB9B01A1" pageId="55" pageNumber="306" type="key">
<keyStep id="92012E3EFFDE6D6FD6DDDF9FBB9B00B4" pageId="55" pageNumber="306">
<paragraph id="294A369BFFDE6D6FD6DDDF9FBB9B0098" blockId="55.[151,1437,1703,2019]" pageId="55" pageNumber="306">
<keyLead id="920495AEFFDE6D6FD6DDDF9FBB9B0098" pageId="55" pageNumber="306">
1 Pronotum with distinct obtuse hind angles. Abdomen very wide. Habitus as in
<figureCitation id="B1CE2A1EFFDE6D6FD595DF9FBA1F00FE" box="[991,1048,1704,1726]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="55" pageNumber="306">
Fig.
<quantity id="EE0D9B7EFFDE6D6FD246DF9FBA1F00FE" box="[1036,1048,1704,1725]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="55" pageNumber="306" unit="in" value="2.0">2</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDE6D6FD272DF9FBAB600FE" author="Steel, W. O." box="[1080,1201,1703,1726]" pageId="55" pageNumber="306" pagination="275 - 278" refId="ref36054" refString="Steel, W. O. (1955) Notes on the Omaliinae (Col., Staphylinidae). 7. The genus Xanthonomus Bernhauer. The Entomologist's Monthly Magazine, 91, 275 - 278." type="journal article" year="1955">Steel (1955)</bibRefCitation>
. Body length:
<quantity id="EE0D9B7EFFDE6D6FD30ADF90BB9E00FE" box="[1344,1433,1703,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="55" pageNumber="306" unit="mm" value="2.5">2.50 mm</quantity>
.
<collectingCountry id="51E2760BFFDE6D6FD6A8DFF2BF3D0098" box="[226,314,1733,1755]" name="Malaysia" pageId="55" pageNumber="306">Malaysia</collectingCountry>
............................................................................
<taxonomicName id="EEF54D18FFDE6D6FD2ABDFF1BB9B0098" ID-CoL="5C6HX" authority="Steel, 1955" authorityName="Steel" authorityYear="1955" box="[1249,1436,1733,1755]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="306" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="1B81EA89FFDE6D6FD2ABDFF1BAF70098" box="[1249,1264,1734,1755]" italics="true" pageId="55" pageNumber="306">X</emphasis>
.
<emphasis id="1B81EA89FFDE6D6FD2B6DFF2BB2C0099" box="[1276,1323,1733,1754]" italics="true" pageId="55" pageNumber="306">latus</emphasis>
<bibRefCitation id="4D644B6AFFDE6D6FD37ADFF2BB9B0098" author="Steel, W. O." box="[1328,1436,1733,1755]" pageId="55" pageNumber="306" pagination="275 - 278" refId="ref36054" refString="Steel, W. O. (1955) Notes on the Omaliinae (Col., Staphylinidae). 7. The genus Xanthonomus Bernhauer. The Entomologist's Monthly Magazine, 91, 275 - 278." type="journal article" year="1955">Steel, 1955</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="294A369BFFDE6D6FD6DDDFD4BB9B00B4" blockId="55.[151,1437,1703,2019]" box="[151,1436,1762,1784]" pageId="55" pageNumber="306">
<keyLead id="920495AEFFDE6D6FD6DDDFD4BB9B00B4" box="[151,1436,1762,1784]" pageId="55" pageNumber="306">- Pronotum with widely rounded hind angles. Abdomen distinctly narrower........................................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="92012E3EFFDE6D6FD6DDDE37BB9B012E" pageId="55" pageNumber="306">
<paragraph id="294A369BFFDE6D6FD6DDDE37BB9B0113" blockId="55.[151,1437,1703,2019]" pageId="55" pageNumber="306">
<keyLead id="920495AEFFDE6D6FD6DDDE37BB9B0113" pageId="55" pageNumber="306">
2 Pronotum slightly transverse, with two median, shallow and oval impressions. Elytra yellow. Habitus as in
<figureCitation id="B1CE2A1EFFDE6D6FD344DE37BB4F0156" box="[1294,1352,1792,1814]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="55" pageNumber="306">
Fig.
<quantity id="EE0D9B7EFFDE6D6FD376DE37BB4F0156" box="[1340,1352,1792,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="55" pageNumber="306" unit="in" value="1.0">1</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDE6D6FD326DE37BF230170" author="Steel, W. O." pageId="55" pageNumber="306" pagination="275 - 278" refId="ref36054" refString="Steel, W. O. (1955) Notes on the Omaliinae (Col., Staphylinidae). 7. The genus Xanthonomus Bernhauer. The Entomologist's Monthly Magazine, 91, 275 - 278." type="journal article" year="1955">Steel (1955)</bibRefCitation>
. Body larger:
<quantity id="EE0D9B7EFFDE6D6FD7E6DE2ABC3D0171" box="[428,570,1821,1843]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="55" pageNumber="306" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.50–2.80 mm</quantity>
. Aedeagus as in
<figureCitation id="B1CE2A1EFFDE6D6FD493DE2ABD160170" box="[729,785,1821,1843]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="55" pageNumber="306">
Fig.
<quantity id="EE0D9B7EFFDE6D6FD54FDE2ABD160170" box="[773,785,1821,1843]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="55" pageNumber="306" unit="in" value="5.0">5</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFDE6D6FD57ADE2ABDAE0170" author="Steel, W. O." box="[816,937,1821,1843]" pageId="55" pageNumber="306" pagination="275 - 278" refId="ref36054" refString="Steel, W. O. (1955) Notes on the Omaliinae (Col., Staphylinidae). 7. The genus Xanthonomus Bernhauer. The Entomologist's Monthly Magazine, 91, 275 - 278." type="journal article" year="1955">Steel (1955)</bibRefCitation>
.
<collectingCountry id="51E2760BFFDE6D6FD5F8DE2ABA080170" box="[946,1039,1821,1843]" name="Indonesia" pageId="55" pageNumber="306">Indonesia</collectingCountry>
(Buru)...................................................................................................
<taxonomicName id="EEF54D18FFDE6D6FD215DE0CBB9B0113" ID-CoL="4GGFH" authority="(Bernhauer, 1926)" baseAuthorityName="Bernhauer" baseAuthorityYear="1926" box="[1119,1436,1850,1872]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="306" phylum="Arthropoda" rank="species" species="toxopeanus">
<emphasis id="1B81EA89FFDE6D6FD215DE0CBA690113" box="[1119,1134,1851,1872]" italics="true" pageId="55" pageNumber="306">X</emphasis>
.
<emphasis id="1B81EA89FFDE6D6FD230DE0CBAE00113" box="[1146,1255,1851,1872]" italics="true" pageId="55" pageNumber="306">toxopeanus</emphasis>
(
<bibRefCitation id="4D644B6AFFDE6D6FD2BEDE0DBB920113" author="Bernhauer, M." box="[1268,1429,1850,1872]" pageId="55" pageNumber="306" pagination="311 - 327" refId="ref35348" refString="Bernhauer, M. (1926) Fauna Buruana Staphylinidae. Zur Staphylinidenfauna der Molukken. Treubia, 7 (3), 311 - 327." type="journal article" year="1926">Bernhauer, 1926</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="294A369BFFDE6D6FD6DDDE6FBB9B012E" blockId="55.[151,1437,1703,2019]" box="[151,1436,1880,1902]" pageId="55" pageNumber="306">
<keyLead id="920495AEFFDE6D6FD6DDDE6FBB9B012E" box="[151,1436,1880,1902]" pageId="55" pageNumber="306">- Pronotum distinctly transverse, without median impressions. Elytra darker. Body smaller............................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="92012E3EFFDE6D6FD6DDDE42BB9B01A1" pageId="55" pageNumber="306">
<paragraph id="294A369BFFDE6D6FD6DDDE42BB9B01E4" blockId="55.[151,1437,1703,2019]" pageId="55" pageNumber="306">
<keyLead id="920495AEFFDE6D6FD6DDDE42BB9B01E4" pageId="55" pageNumber="306">
3 Median lobe moderately wide; parameres distinctly not exceeding apex of median lobe (
<figureCitation id="B1CE2A1EFFDE6D6FD263DE42BA7B01C8" box="[1065,1148,1909,1931]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="55" pageNumber="306">Fig. 152</figureCitation>
). Habitus asi n
<figureCitation id="B1CE2A1EFFDE6D6FD344DE42BB6701C9" box="[1294,1376,1909,1931]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="55" pageNumber="306">Fig. 142</figureCitation>
. Body larger:
<quantity id="EE0D9B7EFFDE6D6FD76CDEA5BFB301EB" box="[294,436,1938,1960]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" metricValueMax="2.25" metricValueMin="1.87" pageId="55" pageNumber="306" unit="mm" value="2.06" valueMax="2.25" valueMin="1.87">1.87–2.25 mm</quantity>
.
<collectingCountry id="51E2760BFFDE6D6FD7F7DEA5BC1D01EB" box="[445,538,1938,1960]" name="Indonesia" pageId="55" pageNumber="306">Indonesia</collectingCountry>
(New
<collectingCountry id="51E2760BFFDE6D6FD416DEA5BCA401EB" box="[604,675,1938,1960]" name="Guinea-Bissau" pageId="55" pageNumber="306">Guinea</collectingCountry>
)..............................................
<taxonomicName id="EEF54D18FFDE6D6FD294DEA4BB6701EB" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1246,1376,1939,1960]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="305" phylum="Arthropoda" rank="species" species="guineensis" status="sp. n.">
X.
<emphasis id="1B81EA89FFDE6D6FD2B1DEA4BB6701EB" box="[1275,1376,1939,1960]" italics="true" pageId="55" pageNumber="306">guineensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDE6D6FD32CDEA5BB9B01E4" bold="true" box="[1382,1436,1938,1959]" pageId="55" pageNumber="306">
<taxonomicNameLabel id="00B257F2FFDE6D6FD32CDEA5BB9B01E4" box="[1382,1436,1938,1959]" pageId="55" pageNumber="306" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="294A369BFFDE6D6FD6DDDE87BB9B01A1" blockId="55.[151,1437,1703,2019]" pageId="55" pageNumber="306">
<keyLead id="920495AEFFDE6D6FD6DDDE87BB9B01A1" pageId="55" pageNumber="306">
- Median lobe narrow; parameres reaching apex of median lobe (
<figureCitation id="B1CE2A1EFFDE6D6FD574DE87BD950186" box="[830,914,1967,1990]" captionStart="FIGURES 159–165" captionStartId="55.[151,264,1456,1480]" captionTargetBox="[228,1379,199,1423]" captionTargetId="figure-16@55.[201,1386,181,1432]" captionTargetPageId="55" captionText="FIGURES 159–165. Xanthonomus thayerae: 159–aedeagus, parameral view (holotype), 160–aedeagus, lateral view (holotype), 161–male abdominal tergite VIII, 162–male abdominal sternite VIII, 163–female abdominal tergite VIII, 164–female abdominal sternite VIII, 165–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821301" httpUri="https://zenodo.org/record/10821301/files/figure.png" pageId="55" pageNumber="306">Fig. 159</figureCitation>
). Habitus as in
<figureCitation id="B1CE2A1EFFDE6D6FD26EDE87BA710186" box="[1060,1142,1968,1990]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="55" pageNumber="306">Fig. 143</figureCitation>
. Body smaller:
<quantity id="EE0D9B7EFFDE6D6FD341DE87BB9E0186" box="[1291,1433,1968,1990]" metricMagnitude="-3" metricUnit="m" metricValue="1.705" metricValueMax="1.8" metricValueMin="1.61" pageId="55" pageNumber="306" unit="mm" value="1.705" valueMax="1.8" valueMin="1.61">1.61–1.80 mm</quantity>
.
<collectingCountry id="51E2760BFFDE6D6FD6A8DEFABF9C01A0" box="[226,411,1997,2019]" name="Papua New Guinea" pageId="55" pageNumber="306">Papua New Guinea</collectingCountry>
......................................................................
<taxonomicName id="EEF54D18FFDE6D6FD2B8DEF9BB6701A1" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1266,1376,1997,2019]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="306" phylum="Arthropoda" rank="species" species="thayerae" status="sp. n.">
<emphasis id="1B81EA89FFDE6D6FD2B8DEF9BB0601A0" box="[1266,1281,1998,2019]" italics="true" pageId="55" pageNumber="306">X</emphasis>
.
<emphasis id="1B81EA89FFDE6D6FD346DEFABB6701A1" box="[1292,1376,1997,2018]" italics="true" pageId="55" pageNumber="306">thayerae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDE6D6FD32CDEFABB9B01A1" bold="true" box="[1382,1436,1997,2018]" pageId="55" pageNumber="306">
<taxonomicNameLabel id="00B257F2FFDE6D6FD32CDEFABB9B01A1" box="[1382,1436,1997,2018]" pageId="55" pageNumber="306" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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