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<document id="4398A60AEFF4B5C9401BD83FCE177496" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFCB6D7CD6DDDAD1BD300422" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba penelopeae Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="287" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="285" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="AFC830D1F7C984025F9DEFB5124F4C58" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="90DF9769F30890102FB7ACC30AE7DFED">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
</mods:titleInfo>
<mods:name id="A3FCFEC78EE13C5AA249C2E6AFD0649F" type="personal">
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<mods:namePart id="61F3E6214FFA43BE43C4889440F11874">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="0C714ADD3C9C13AAFEC9A63B32EBDCE8">Zootaxa</mods:title>
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<mods:date id="45076557C82646A87B0E0E804A57236A">2024</mods:date>
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<mods:number id="052B0E0804E8EA606CE16AB3D33673CF">2024-03-14</mods:number>
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<mods:classification id="46E7B1E62305A435A87010122534B5D4">journal article</mods:classification>
<mods:identifier id="8B77A6525FC14BCB419BD6D245957D69" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="4B0F7A8DB082B51C8DA934611401B004" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B7AFEE4F0D6F55D4A281B8E1482212EF" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="1EE95A9AC399036D5AA0FD467BADE544" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFCB6D7CD6DDDAD1BD300422" LSID="urn:lsid:plazi:treatment:A15C878DFFCB6D7CD6DDDAD1BD300422" httpUri="http://treatment.plazi.org/id/A15C878DFFCB6D7CD6DDDAD1BD300422" lastPageId="36" lastPageNumber="287" pageId="34" pageNumber="285">
<subSubSection id="61EF6510FFCB6D7AD6DDDAD1BC330242" box="[151,564,998,1025]" pageId="34" pageNumber="285" type="nomenclature">
<paragraph id="294A369BFFCB6D7AD6DDDAD1BC330242" blockId="34.[151,564,998,1061]" box="[151,564,998,1025]" pageId="34" pageNumber="285">
<heading id="720281F7FFCB6D7AD6DDDAD1BC330242" bold="true" box="[151,564,998,1025]" fontSize="11" level="1" pageId="34" pageNumber="285" reason="1">
<emphasis id="1B81EA89FFCB6D7AD6DDDAD1BC330242" bold="true" box="[151,564,998,1025]" pageId="34" pageNumber="285">
<taxonomicName id="EEF54D18FFCB6D7AD6DDDAD1BFEC0242" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,491,998,1025]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCB6D7AD6DDDAD1BFEC0242" bold="true" box="[151,491,998,1025]" italics="true" pageId="34" pageNumber="285">Paraphloeostiba penelopeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFCB6D7AD7B8DAD0BC330242" box="[498,564,999,1025]" pageId="34" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCB6D7AD6DDDD3CBF580266" box="[151,351,1035,1061]" pageId="34" pageNumber="285" type="description">
<paragraph id="294A369BFFCB6D7AD6DDDD3CBF580266" blockId="34.[151,564,998,1061]" box="[151,351,1035,1061]" pageId="34" pageNumber="285">
(
<figureCitation id="B1CE2A1EFFCB6D7AD6D5DD3CBEF00266" box="[159,247,1035,1061]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="34" pageNumber="285">Figs 76</figureCitation>
,
<figureCitation id="B1CE2A1EFFCB6D7AD748DD3CBF500266" box="[258,343,1035,1061]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="34" pageNumber="285">97–103</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCB6D7AD6DDDD64BCBA0362" pageId="34" pageNumber="285" type="materials_examined">
<paragraph id="294A369BFFCB6D7AD6DDDD64BD9102F6" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
<emphasis id="1B81EA89FFCB6D7AD6DDDD64BF43022F" bold="true" box="[151,324,1106,1133]" pageId="34" pageNumber="285">Type material.</emphasis>
<typeStatus id="F64E8839FFCB6D7AD703DD64BFB6022E" box="[329,433,1107,1133]" pageId="34" pageNumber="285" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFCB6D7AD7FDDD64BFC8022E" box="[439,463,1107,1133]" count="1" pageId="34" pageNumber="285" type="male">♂</specimenCount>
: ‘SOLOMON IS. |
<collectingRegion id="EB31F879FFCB6D7AD4E9DD64BD28022E" box="[675,815,1107,1133]" country="Solomon Islands" name="Guadalcanal" pageId="34" pageNumber="285">Guadalcanal</collectingRegion>
| Mt. Austen [handwritten] | Jan [handwritten] 1966 [the last number “6” is handwritten] | 19993 [handwritten] | P. Greenslade’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFCB6D7AD263DD40BAC502D2" box="[1065,1218,1143,1169]" pageId="34" pageNumber="285" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFCB6D7AD29FDD40BB9702D2" authorityName="Steel" authorityYear="1960" box="[1237,1424,1143,1169]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFCB6D7AD29FDD40BB9702D2" box="[1237,1424,1143,1169]" italics="true" pageId="34" pageNumber="285">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFCB6D7AD6DDDDACBF1002F6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,279,1179,1205]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCB6D7AD6DDDDACBF1002F6" box="[151,279,1179,1205]" italics="true" pageId="34" pageNumber="285">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCB6D7AD754DDACBF5A02F6" bold="true" box="[286,349,1179,1205]" pageId="34" pageNumber="285">
<taxonomicNameLabel id="00B257F2FFCB6D7AD754DDACBF5A02F6" box="[286,349,1179,1205]" pageId="34" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFCB6D7AD7ACDDACBFFB02F6" box="[486,508,1179,1205]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="34" pageNumber="285" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFCB6D7AD57ADDACBD8D02F6" box="[816,906,1179,1205]" pageId="34" pageNumber="285">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDD88BCBA0362" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
<typeStatus id="F64E8839FFCB6D7AD68DDD88BF3F029A" box="[199,312,1215,1241]" pageId="34" pageNumber="285" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFCB6D7AD709DD88BF96029A" box="[323,401,1215,1241]" count="14" pageId="34" pageNumber="285" type="male">14 ♂♂</specimenCount>
(
<specimenCount id="3FF3FD12FFCB6D7AD7EADD88BC3B029A" box="[416,572,1215,1241]" count="1" pageId="34" pageNumber="285" type="generic">one specimen</specimenCount>
dissected),
<specimenCount id="3FF3FD12FFCB6D7AD48BDD88BD08029A" box="[705,783,1215,1241]" count="18" pageId="34" pageNumber="285" type="female">18 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFCB6D7AD554DD88BDBD029A" box="[798,954,1215,1241]" count="1" pageId="34" pageNumber="285" type="generic">one specimen</specimenCount>
dissected): ‘SOLOMON IS. |
<collectingRegion id="EB31F879FFCB6D7AD35ADD88BB9B029A" box="[1296,1436,1215,1241]" country="Solomon Islands" name="Guadalcanal" pageId="34" pageNumber="285">Guadalcanal</collectingRegion>
Mt. | Austen
<date id="5D4B105BFFCB6D7AD77ADDD4BF9F02BE" box="[304,408,1251,1277]" pageId="34" pageNumber="285" value="1966-01">Jan 1966</date>
| 19993 P. | Greenslade’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFCB6D7AD508DDD4BDD502BE" box="[834,978,1251,1277]" pageId="34" pageNumber="285" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFCB6D7AD5A6DDD4BAA002BE" authorityName="Steel" authorityYear="1960" box="[1004,1191,1251,1277]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFCB6D7AD5A6DDD4BAA002BE" box="[1004,1191,1251,1277]" italics="true" pageId="34" pageNumber="285">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFCB6D7AD28BDDD4BB4602BE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1217,1345,1251,1277]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="285" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCB6D7AD28BDDD4BB4602BE" box="[1217,1345,1251,1277]" italics="true" pageId="34" pageNumber="285">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCB6D7AD300DDD4BB8A02BE" bold="true" box="[1354,1421,1251,1277]" pageId="34" pageNumber="285">
<taxonomicNameLabel id="00B257F2FFCB6D7AD300DDD4BB8A02BE" box="[1354,1421,1251,1277]" pageId="34" pageNumber="285" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFCB6D7AD747DC30BF240362" box="[269,291,1287,1313]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="34" pageNumber="285" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFCB6D7AD41DDC30BCB60362" box="[599,689,1287,1313]" pageId="34" pageNumber="285">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCB6D7CD68DDC1DBC1407EE" lastPageId="36" lastPageNumber="287" pageId="34" pageNumber="285" type="description">
<paragraph id="294A369BFFCB6D7AD68DDC1DBA7B03CE" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
<emphasis id="1B81EA89FFCB6D7AD68DDC1DBF5E0307" bold="true" box="[199,345,1322,1348]" pageId="34" pageNumber="285">Description.</emphasis>
Measurements (n=33): HW: 0.40; HL: 0.17–0.22; OL: 0.10–0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFCB6D7AD2A2DC1CBB560306" box="[1256,1361,1323,1349]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
): 0.45; PL: 0.18–0.25; PW: 0.43–0.52; ESL: 0.36–0.45; EW: 0.55–0.60; MTbL (
<typeStatus id="F64E8839FFCB6D7AD591DC78BA43032A" box="[987,1092,1359,1385]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFCB6D7AD2A4DC78BB50032A" box="[1262,1367,1359,1385]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
): 0.17 (MTrL 1–4: 0.05; MTrL 5: 0.12); AW: 0.58; AedL: 0.32; BL: 1.40–2.00 (
<typeStatus id="F64E8839FFCB6D7AD582DC44BA3703CE" box="[968,1072,1395,1421]" pageId="34" pageNumber="285" type="holotype">holotype</typeStatus>
: 1.85).
</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDCA0BFDF00CA" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">
Habitus as in
<figureCitation id="B1CE2A1EFFCB6D7AD727DCA0BFC303F2" box="[365,452,1431,1457]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="34" pageNumber="285">Fig. 76</figureCitation>
. Body yellow-brown, with reddish-brown apical portions of elytra (some specimens with slightly darker neck, head (or only clypeus) and apical parts of abdominal tergites III–VI). Head and neck, pronotum and scutellum without punctation; elytra with moderately dense and large punctation, finer and denser around scutellum and finer along suture; abdominal tergites without visible punctation. Pronotum with very dense isodiametric microsculpture, slightly finer than that in middle portion of head; elytra with dense and coarse isodiametric microreticulation, slightly larger than that in pronotum; abdominal tergites with dense transverse or isodiametric microsculpture.
</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDFA4BEE40122" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">Head transverse, about twice as broad as long, with very short and moderately deep anteocellar foveae as long as diameter of ocellus. Ocelli moderately large, located at level of posterior margins of eyes; distance between ocelli 1.2–1.4 times as long as distance between ocellus and posterior margin of eyes. Antennomeres 6–10 distinctly transverse; antennomere 3 distinctly longer and slightly narrower than 2, 5 slightly broader than 4, 6 about as long as and about wice as broad as 5, 7–8 slightly longer and broader than 6, 9 slightly broader than 8, 10 slightly longer than 9.</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDE5CBF9901EA" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">Pronotum slightly more than twice as broad as long, broader than head, widest in middle, slightly more narrowed posteriad than anteriad.</paragraph>
<paragraph id="294A369BFFCB6D7AD68DDE84BEFE01B2" blockId="34.[151,1437,1106,2033]" pageId="34" pageNumber="285">Elytra 1.3–1.5 times as broad as long, 1.8 times to twice as long as pronotum, with narrowly impressed lateral margins.</paragraph>
<caption id="7D8A6613FFCA6D7BD6DDDC8CBD1B0058" ID-DOI="http://doi.org/10.5281/zenodo.10821267" ID-Zenodo-Dep="10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286" startId="35.[151,264,1467,1491]" targetBox="[160,1427,159,1442]" targetPageId="35" targetType="figure">
<paragraph id="294A369BFFCA6D7BD6DDDC8CBD1B0058" blockId="35.[151,1436,1467,1564]" pageId="35" pageNumber="286">
<emphasis id="1B81EA89FFCA6D7BD6DDDC8CBF630397" bold="true" box="[151,356,1467,1492]" pageId="35" pageNumber="286">FIGURES 97–103.</emphasis>
<taxonomicName id="EEF54D18FFCA6D7BD726DC8BBC960397" authorityName="Shavrin" authorityYear="2024" box="[364,657,1468,1492]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="penelopeae">
<emphasis id="1B81EA89FFCA6D7BD726DC8BBC960397" box="[364,657,1468,1492]" italics="true" pageId="35" pageNumber="286">Paraphloeostiba penelopeae</emphasis>
</taxonomicName>
(paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFCA6D7BD68DDF68BD23003A" blockId="35.[151,1437,1631,2042]" box="[199,804,1631,1658]" pageId="35" pageNumber="286">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="294A369BFFCA6D7BD68DDFB5BD23016B" blockId="35.[151,1437,1631,2042]" pageId="35" pageNumber="286">
Male. Posterior margin of abdominal tergite VIII straight (
<figureCitation id="B1CE2A1EFFCA6D7BD51BDFB5BDA200DE" box="[849,933,1666,1693]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 99</figureCitation>
). Posterior margin of abdominal sternite VIII widely concave (
<figureCitation id="B1CE2A1EFFCA6D7BD71DDF92BFBD0083" box="[343,442,1701,1728]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 100</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward wide median lobe, with widely rounded apex; parameres slightly exceeding apex of median lobe, with broadened apical portions, each with two long apical and four preapical setae; internal sac narrow and moderately long, with elongate structure in basal part (
<figureCitation id="B1CE2A1EFFCA6D7BD698DE39BF21016B" box="[210,294,1806,1832]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 97</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFCA6D7BD487DE39BD27016B" box="[717,800,1806,1832]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 98</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFCA6D7BD68DDE06BCB801D2" blockId="35.[151,1437,1631,2042]" pageId="35" pageNumber="286">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFCA6D7BD53FDE06BDD00108" box="[885,983,1841,1867]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 101</figureCitation>
). Posterior margin of abdominal sternite VIII widely rounded (
<figureCitation id="B1CE2A1EFFCA6D7BD7D8DE63BFF1012D" box="[402,502,1876,1902]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 102</figureCitation>
). Accessory sclerite with wide basal portion, gradually narrowed toward long and narrow apical part with subacute apex (
<figureCitation id="B1CE2A1EFFCA6D7BD41ADE40BCB501D2" box="[592,690,1911,1937]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="35" pageNumber="286">Fig. 103</figureCitation>
).
</paragraph>
<paragraph id="294A369BFFCA6D7BD68DDEADBA1E01B9" blockId="35.[151,1437,1631,2042]" pageId="35" pageNumber="286">
<emphasis id="1B81EA89FFCA6D7BD68DDEADBFB701F7" bold="true" box="[199,432,1946,1972]" pageId="35" pageNumber="286">Comparative notes.</emphasis>
Based on the very broad median lobe and apical portions of the parameres,
<taxonomicName id="EEF54D18FFCA6D7BD2B4DEACBB9A01F7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1278,1437,1946,1972]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFCA6D7BD2B4DEACBB0801F7" box="[1278,1295,1947,1972]" italics="true" pageId="35" pageNumber="286">P</emphasis>
.
<emphasis id="1B81EA89FFCA6D7BD357DEADBB9A01F7" box="[1309,1437,1946,1972]" italics="true" pageId="35" pageNumber="286">penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCA6D7BD6DDDE8ABED00194" bold="true" box="[151,215,1981,2007]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="00B257F2FFCA6D7BD6DDDE8ABED00194" box="[151,215,1981,2007]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFCA6D7BD720DE89BFE50194" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[362,482,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFCA6D7BD720DE89BFE50194" box="[362,482,1981,2007]" italics="true" pageId="35" pageNumber="286">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCA6D7BD7A3DE8ABC2F0194" bold="true" box="[489,552,1981,2007]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="00B257F2FFCA6D7BD7A3DE8ABC2F0194" box="[489,552,1981,2007]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFCA6D7BD47CDE89BD490194" authority="Steel, 1960" authorityName="Steel" authorityYear="1960" box="[566,846,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFCA6D7BD47CDE89BCC20194" box="[566,709,1981,2007]" italics="true" pageId="35" pageNumber="286">P. celebensis</emphasis>
<bibRefCitation id="4D644B6AFFCA6D7BD486DE8ABD490194" author="Steel, W. O." box="[716,846,1981,2007]" pageId="35" pageNumber="286" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel, 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFCA6D7BD513DE89BDD00194" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[857,983,1982,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFCA6D7BD513DE89BDD00194" box="[857,983,1982,2007]" italics="true" pageId="35" pageNumber="286">P. papuana</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFCA6D7BD5AFDE89BA420194" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[997,1093,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFCA6D7BD5AFDE89BA420194" box="[997,1093,1981,2007]" italics="true" pageId="35" pageNumber="286">P. rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCA6D7BD207DE8ABA8B0194" bold="true" box="[1101,1164,1981,2007]" pageId="35" pageNumber="286">
<taxonomicNameLabel id="00B257F2FFCA6D7BD207DE8ABA8B0194" box="[1101,1164,1981,2007]" pageId="35" pageNumber="286" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFCA6D7BD289DE89BB480194" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[1219,1359,1981,2007]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="286" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFCA6D7BD289DE89BB480194" box="[1219,1359,1981,2007]" italics="true" pageId="35" pageNumber="286">P. tonkinesis</emphasis>
</taxonomicName>
. It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD9A0BCE206F2" blockId="36.[151,1436,151,609]" box="[199,741,151,177]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD9A0BF7B06F2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,151,177]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFCD6D7CD74FD9A0BF7B06F2" box="[261,380,151,177]" italics="true" pageId="36" pageNumber="287">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD7C9D9A0BFC506F2" bold="true" box="[387,450,151,177]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD7C9D9A0BFC506F2" box="[387,450,151,177]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the shorter parameres;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD98CBA340696" blockId="36.[151,1436,151,609]" box="[199,1075,187,213]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD98CBF940696" authorityName="Steel" authorityYear="1960" box="[261,403,187,213]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="1B81EA89FFCD6D7CD74FD98CBF940696" box="[261,403,187,213]" italics="true" pageId="36" pageNumber="287">P. celebensis</emphasis>
</taxonomicName>
by slightly longer parameres with narrower apical portions;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD9E8BD4006BA" blockId="36.[151,1436,151,609]" box="[199,839,223,249]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD9E8BF8506BB" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[261,386,223,248]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFCD6D7CD74FD9E8BF8506BB" box="[261,386,223,248]" italics="true" pageId="36" pageNumber="287">P. papuana</emphasis>
</taxonomicName>
by slightly longer elytra and parameres;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD834BB60075E" blockId="36.[151,1436,151,609]" box="[199,1383,259,285]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD834BF63075E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,356,259,285]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFCD6D7CD74FD834BF63075E" box="[261,356,259,285]" italics="true" pageId="36" pageNumber="287">P. rufula</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD726D834BFAC075E" bold="true" box="[364,427,259,285]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD726D834BFAC075E" box="[364,427,259,285]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller body, the darker head, pronotum and abdomen, and longer parameres;
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD810BDCD0702" blockId="36.[151,1436,151,609]" box="[199,970,295,321]" pageId="36" pageNumber="287">
from
<taxonomicName id="EEF54D18FFCD6D7CD74FD810BF970702" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[261,400,295,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="tonkinesis">
<emphasis id="1B81EA89FFCD6D7CD74FD810BF970702" box="[261,400,295,321]" italics="true" pageId="36" pageNumber="287">P. tonkinesis</emphasis>
</taxonomicName>
by the narrower apical portion of the median lobe.
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DD87CBC1407EE" blockId="36.[151,1436,151,609]" pageId="36" pageNumber="287">
From all these species
<taxonomicName id="EEF54D18FFCD6D7CD7F4D87CBC5F0726" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[446,600,331,357]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFCD6D7CD7F4D87CBC5F0726" box="[446,600,331,357]" italics="true" pageId="36" pageNumber="287">P. penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD417D87CBC9D0726" bold="true" box="[605,666,331,357]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD417D87CBC9D0726" box="[605,666,331,357]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs by the absence of the punctation on the head and the pronotum, the presence of distinct longitudinal impressions of the more transverse pronotum, and details of the external and internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCD6D7CD68DD881BF5D07B6" pageId="36" pageNumber="287" type="distribution">
<paragraph id="294A369BFFCD6D7CD68DD881BF5D07B6" blockId="36.[151,1436,151,609]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD68DD881BF660793" bold="true" box="[199,353,438,464]" pageId="36" pageNumber="287">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFCD6D7CD722D880BCAC0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[360,683,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="287" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. nov.">
<emphasis id="1B81EA89FFCD6D7CD722D880BCAC0792" box="[360,683,439,465]" italics="true" pageId="36" pageNumber="287">Paraphloeostiba penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFCD6D7CD4F8D880BCF40792" bold="true" box="[690,755,439,465]" pageId="36" pageNumber="287">
<taxonomicNameLabel id="00B257F2FFCD6D7CD4F8D880BCF40792" box="[690,755,439,465]" pageId="36" pageNumber="287" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFCD6D7CD241D880BA3C0792" box="[1035,1083,439,465]" pageId="36" pageNumber="287">type</typeStatus>
locality in
<collectingRegion id="EB31F879FFCD6D7CD2F7D880BB4E0792" box="[1213,1353,439,465]" country="Solomon Islands" name="Guadalcanal" pageId="36" pageNumber="287">Guadalcanal</collectingRegion>
Island,
<collectingCountry id="51E2760BFFCD6D7CD6DDD8ECBF5107B6" box="[151,342,475,501]" name="Solomon Islands" pageId="36" pageNumber="287">Solomon Islands</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFCD6D7CD68DD8C8BD300422" pageId="36" pageNumber="287" type="etymology">
<paragraph id="294A369BFFCD6D7CD68DD8C8BB3F047E" blockId="36.[151,1436,151,609]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD68DD8C8BF49045A" bold="true" box="[199,334,511,537]" pageId="36" pageNumber="287">Etymology.</emphasis>
Patronymic, the species is named to honour of entomologist and naturalist Penelope Jean Macleod Greenslade, wife of P.J.M. Greenslade, who collected material on the
<collectingCountry id="51E2760BFFCD6D7CD5EFDB14BA67047E" box="[933,1120,547,573]" name="Solomon Islands" pageId="36" pageNumber="287">Solomon Islands</collectingCountry>
during 1960-1970.
</paragraph>
<paragraph id="294A369BFFCD6D7CD68DDB71BD300422" blockId="36.[151,1436,151,609]" box="[199,823,582,609]" pageId="36" pageNumber="287">
<emphasis id="1B81EA89FFCD6D7CD68DDB71BF4C0423" bold="true" box="[199,331,582,608]" pageId="36" pageNumber="287">Bionomics.</emphasis>
The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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