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<document id="7F8DED5C24F0C7D703B71C8A142F0397" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFC26D75D6DDDCD9BF390276" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba steeli Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="296" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="294" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A7B69A44C0FC0ED63EBFBE608D050D25" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7B157EA58B6498EAECD0165311915251">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
</mods:titleInfo>
<mods:name id="6A2DEAB16D7EE9D0A024C17463DC6F80" type="personal">
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<mods:namePart id="BCEF76495383826C42973930D0F7D80E">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="171C9B57C5562C5E6827A9A6A3CAC83A">Zootaxa</mods:title>
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<mods:date id="4062E8F296112234E771D5BB0B1BEEB5">2024</mods:date>
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<mods:number id="183F8570FC8540D47825B7D5D12B4A49">2024-03-14</mods:number>
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<mods:number id="EACD64980B09680DF9257DE3EF1722BF">5424</mods:number>
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<mods:number id="2EB03DD95B1DDE0ECDE178B25EF01A5F">3</mods:number>
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<mods:url id="39341D87A23CC5FD4A98C5AB2B704099">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
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<mods:classification id="4A2CD7C888D128EC8DAE784FE38AAA34">journal article</mods:classification>
<mods:identifier id="506587B1B5FD351A10BA7B5475DA455F" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="4B85C99A98AEB7B56A1F80F5235B9971" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B23CB3EF57EABC443C126536092DDC9B" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="1895CE340F3243C99ED42C0C5CBAC3FA" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFC26D75D6DDDCD9BF390276" LSID="urn:lsid:plazi:treatment:A15C878DFFC26D75D6DDDCD9BF390276" httpUri="http://treatment.plazi.org/id/A15C878DFFC26D75D6DDDCD9BF390276" lastPageId="45" lastPageNumber="296" pageId="43" pageNumber="294">
<subSubSection id="61EF6510FFC26D73D6DDDCD9BFEC004A" box="[151,491,1518,1545]" pageId="43" pageNumber="294" type="nomenclature">
<paragraph id="294A369BFFC26D73D6DDDCD9BFEC004A" blockId="43.[151,491,1518,1581]" box="[151,491,1518,1545]" pageId="43" pageNumber="294">
<heading id="720281F7FFC26D73D6DDDCD9BFEC004A" bold="true" box="[151,491,1518,1545]" fontSize="11" level="1" pageId="43" pageNumber="294" reason="1">
<emphasis id="1B81EA89FFC26D73D6DDDCD9BFEC004A" bold="true" box="[151,491,1518,1545]" pageId="43" pageNumber="294">
<taxonomicName id="EEF54D18FFC26D73D6DDDCD9BFA5004A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,418,1518,1545]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC26D73D6DDDCD9BFA5004A" bold="true" box="[151,418,1518,1545]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba steeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFC26D73D7E3DCD8BFEC004A" box="[425,491,1519,1545]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC26D73D6DDDF24BF6A006E" box="[151,365,1555,1581]" pageId="43" pageNumber="294" type="description">
<paragraph id="294A369BFFC26D73D6DDDF24BF6A006E" blockId="43.[151,491,1518,1581]" box="[151,365,1555,1581]" pageId="43" pageNumber="294">
(
<figureCitation id="B1CE2A1EFFC26D73D6D5DF24BEF0006E" box="[159,247,1555,1581]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="43" pageNumber="294">Figs 44</figureCitation>
,
<figureCitation id="B1CE2A1EFFC26D73D748DF24BF61006E" box="[258,358,1555,1581]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="43" pageNumber="294">133–140</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC26D73D6DDDF6CBA27016A" pageId="43" pageNumber="294" type="materials_examined">
<paragraph id="294A369BFFC26D73D6DDDF6CBAD600FE" blockId="43.[151,1437,1626,2013]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D6DDDF6CBF430037" bold="true" box="[151,324,1626,1653]" pageId="43" pageNumber="294">Type material.</emphasis>
<typeStatus id="F64E8839FFC26D73D700DF6CBFB50036" box="[330,434,1627,1653]" pageId="43" pageNumber="294" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFC26D73D7F3DF6CBFC90036" box="[441,462,1627,1653]" count="1" pageId="43" pageNumber="294" type="male">♂</specimenCount>
(
<figureCitation id="B1CE2A1EFFC26D73D796DF6CBC280036" box="[476,559,1627,1653]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="43" pageNumber="294">Fig. 44</figureCitation>
; dissected): ‘
<collectionCode id="4FE4AE5EFFC26D73D4F6DF6CBD070036" box="[700,768,1627,1653]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="43" pageNumber="294">NEW</collectionCode>
<collectingCountry id="51E2760BFFC26D73D54CDF6CBD770036" box="[774,880,1627,1653]" name="Guinea-Bissau" pageId="43" pageNumber="294">GUINEA</collectingCountry>
| Kamin- | dimbit, Main River, Se- | pik, Angoram sub-div., |
<date id="5D4B105BFFC26D73D741DF49BF8C00DA" box="[267,395,1662,1689]" pageId="43" pageNumber="294" value="1965-03-12">12.III.1965</date>
, running on | swamp plants in marsh’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFC26D73D5FFDF48BDC900DA" box="[949,974,1663,1689]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="294">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC26D73D6D1DF94BF3300FE" box="[155,308,1699,1725]" pageId="43" pageNumber="294" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC26D73D702DF94BC0400FE" authorityName="Steel" authorityYear="1960" box="[328,515,1699,1725]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC26D73D702DF94BC0400FE" box="[328,515,1699,1725]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC26D73D45CDF94BC5600FE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[534,593,1699,1725]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC26D73D45CDF94BC5600FE" box="[534,593,1699,1725]" italics="true" pageId="43" pageNumber="294">steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D412DF94BC9F00FE" bold="true" box="[600,664,1699,1725]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D412DF94BC9F00FE" box="[600,664,1699,1725]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC26D73D56BDF94BD3000FE" box="[801,823,1699,1725]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="43" pageNumber="294" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC26D73D220DF94BAC300FE" box="[1130,1220,1699,1725]" pageId="43" pageNumber="294">BMNH</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFC26D73D68DDFF0BA27016A" blockId="43.[151,1437,1626,2013]" pageId="43" pageNumber="294">
<typeStatus id="F64E8839FFC26D73D68DDFF0BF3F00A2" box="[199,312,1735,1761]" pageId="43" pageNumber="294" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFC26D73D702DFF0BF7100A2" box="[328,374,1735,1761]" count="1" pageId="43" pageNumber="294" type="female">1 ♀</specimenCount>
(dissected): ‘
<collectionCode id="4FE4AE5EFFC26D73D459DFF0BC5000A2" box="[531,599,1735,1761]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="43" pageNumber="294">NEW</collectionCode>
<collectingCountry id="51E2760BFFC26D73D429DFF0BCCA00A2" box="[611,717,1735,1761]" name="Guinea-Bissau" pageId="43" pageNumber="294">GUINEA</collectingCountry>
| Kamindim- | bit, Main River, Sepik, Ango- | ram sub-div.,
<date id="5D4B105BFFC26D73D6DDDFDDBF1F0146" box="[151,280,1770,1797]" pageId="43" pageNumber="294" value="1965-03-12">12.III.1965</date>
, | running on swamp plants in | marsh’ &lt;printed&gt;, ‘
<collectionCode id="4FE4AE5EFFC26D73D5D8DFDCBDAC0146" box="[914,939,1771,1797]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="294">R</collectionCode>
. Hornabrook | BM 1970-232.’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC26D73D6D1DE38BF2C016A" box="[155,299,1807,1833]" pageId="43" pageNumber="294" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC26D73D775DE38BFFD016A" authorityName="Steel" authorityYear="1960" box="[319,506,1807,1833]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC26D73D775DE38BFFD016A" box="[319,506,1807,1833]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC26D73D447DE38BC4F016A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[525,584,1807,1833]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC26D73D447DE38BC4F016A" box="[525,584,1807,1833]" italics="true" pageId="43" pageNumber="294">steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC26D73D405DE38BC88016A" bold="true" box="[591,655,1807,1833]" pageId="43" pageNumber="294">
<taxonomicNameLabel id="00B257F2FFC26D73D405DE38BC88016A" box="[591,655,1807,1833]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC26D73D552DE38BD29016A" box="[792,814,1807,1833]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="43" pageNumber="294" type="Museum">V</collectionCode>
. des. 2024’ (
<collectionCode id="4FE4AE5EFFC26D73D5F0DE38BA13016A" box="[954,1044,1807,1833]" pageId="43" pageNumber="294">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC26D75D68DDE05BF2805C2" lastPageId="45" lastPageNumber="296" pageId="43" pageNumber="294" type="description">
<paragraph id="294A369BFFC26D73D68DDE05BB8601D6" blockId="43.[151,1437,1626,2013]" pageId="43" pageNumber="294">
<emphasis id="1B81EA89FFC26D73D68DDE05BF5E010F" bold="true" box="[199,345,1842,1868]" pageId="43" pageNumber="294">Description.</emphasis>
Measurements (n=2): HW: 0.25–0.26; HL: 0.21–0.22; OL: 0.12; TL: 0.02; AL (
<typeStatus id="F64E8839FFC26D73D2ADDE04BB57010E" box="[1255,1360,1843,1869]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
): 0.49; PL: 0.32–0.34; PWmax: 0.55–0.56; PWmin: 0.50–0.52; ESL: 0.52–0.53; EW: 0.64–0.66; MTbL (
<typeStatus id="F64E8839FFC26D73D2ADDE60BB570132" box="[1255,1360,1879,1905]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFC26D73D6A2DE4CBF5601D6" box="[232,337,1915,1941]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
): 0.14 (MTrL 1–4: 0.05; MTrL 5: 0.09); AW: 0.60–0.62; AedL: 0.37; BL: 1.78–1.90 (
<typeStatus id="F64E8839FFC26D73D346DE4CBB7201D6" box="[1292,1397,1915,1941]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="294A369BFFC26D74D68DDEA8BCE0075E" blockId="43.[151,1437,1626,2013]" lastBlockId="44.[151,1437,151,645]" lastPageId="44" lastPageNumber="295" pageId="43" pageNumber="294">
Habitus as in
<figureCitation id="B1CE2A1EFFC26D73D716DEA8BFAA01FA" box="[348,429,1951,1977]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="43" pageNumber="294">Fig. 44</figureCitation>
. Body reddish-brown (
<typeStatus id="F64E8839FFC26D73D4E0DEA8BD0901FA" box="[682,782,1951,1977]" pageId="43" pageNumber="294" type="holotype">holotype</typeStatus>
with slightly darker abdomen); antennomeres 6–11 brown; mouthparts, antennomeres 1–4 and legs yellow. Head and neck without visible punctures; pronotum with very sparse and fine indistinct punctation, invisible in middle; elytra with dense punctation, distinctly larger and deeper than that on pronotum, slightly finer and sparser in middle; abdominal tergites without visible punctation. Head with very dense and coarse isodiametric microsculpture; microsculpture of pronotum about as that in middle portion of head; elytra with fine isodiametric microreticulation.
</paragraph>
<paragraph id="294A369BFFC56D74D68DD810BCB40792" blockId="44.[151,1437,151,645]" pageId="44" pageNumber="295">Head with moderately deep and wide anteriomedian depressions, slightly broader than long, with short and deep narrow anteocellar foveae about as long as diameter of ocellus; postocular carina absent. Distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Antennomeres 6–10 distinctly transverse; antennomere 5 slightly broader than 4, 6 distinctly broader than 5, 7 longer and broader than 6, 8–9 slightly shorter than 7, 10 slightly longer than 9.</paragraph>
<paragraph id="294A369BFFC56D74D68DD8ECBF99045A" blockId="44.[151,1437,151,645]" pageId="44" pageNumber="295">Pronotum 1.4–1.6 times as broad as long, twice as broad as head, widest in middle, distinctly more narrowed posteriad than anteriad.</paragraph>
<paragraph id="294A369BFFC56D74D68DDB14BC7B0422" blockId="44.[151,1437,151,645]" pageId="44" pageNumber="295">Elytra 1.2 times as broad as long, 1.5–1.6 times as long as pronotum; surface between punctures slightly and longitudinally elevated, stronger in middle.</paragraph>
<paragraph id="294A369BFFC56D74D68DDB5CBC8504C6" blockId="44.[151,1437,151,645]" box="[199,642,619,645]" pageId="44" pageNumber="295">Abdomen slightly narrower than elytra.</paragraph>
<caption id="7D8A6613FFC56D74D6DDDEA0BDDD01B4" ID-DOI="http://doi.org/10.5281/zenodo.10821281" ID-Zenodo-Dep="10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="44" pageNumber="295" startId="44.[151,264,1943,1967]" targetBox="[185,1403,678,1912]" targetPageId="44" targetType="figure">
<paragraph id="294A369BFFC56D74D6DDDEA0BDDD01B4" blockId="44.[151,1436,1943,2040]" pageId="44" pageNumber="295">
<emphasis id="1B81EA89FFC56D74D6DDDEA0BF6A01EC" bold="true" box="[151,365,1943,1967]" pageId="44" pageNumber="295">FIGURES 133–140.</emphasis>
<taxonomicName id="EEF54D18FFC56D74D738DEA0BC5201EC" authorityName="Shavrin" authorityYear="2024" box="[370,597,1943,1967]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="295" phylum="Arthropoda" rank="species" species="steeli">
<emphasis id="1B81EA89FFC56D74D738DEA0BC5201EC" box="[370,597,1943,1967]" italics="true" pageId="44" pageNumber="295">Paraphloeostiba steeli</emphasis>
</taxonomicName>
: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="294A369BFFC46D75D68DD9A0BF300726" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC46D75D521D9A0BDC806F2" box="[875,975,150,177]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 135</figureCitation>
). Posterior margin of abdominal sternite VIII widely and deeply concave (
<figureCitation id="B1CE2A1EFFC46D75D45CD98CBC7D0696" box="[534,634,187,213]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 136</figureCitation>
). Aedeagus with wide basal portion, strongly narrowed toward narrow elongate median lobe with very small rounded apex; parameres slightly exceeding apex of median lobe, moderately wide, distinctly broadened apically, with widely rounded apices, with two moderately short apical and preapical setae; internal sac long and moderately wide, without sclerotized structures (
<figureCitation id="B1CE2A1EFFC46D75D595D810BA470702" box="[991,1088,295,321]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 133</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC46D75D698D87CBF340726" box="[210,307,331,357]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 134</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC46D75D68DD858BC8F0792" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
Female. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC46D75D53FD858BDDF07CA" box="[885,984,367,393]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 137</figureCitation>
). Posterior margin of abdominal sternite VIII rounded (
<figureCitation id="B1CE2A1EFFC46D75D774D8A4BFA507EE" box="[318,418,403,429]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 138</figureCitation>
). Accessory sclerite short, from widest basal portion gradually narrowed toward rounded apex (
<figureCitation id="B1CE2A1EFFC46D75D691D880BF390792" box="[219,318,439,465]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 139</figureCitation>
). Spermatheca as in
<figureCitation id="B1CE2A1EFFC46D75D46ED880BC830792" box="[548,644,439,465]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="45" pageNumber="296">Fig. 140</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC46D75D68DD8EDBCFC047E" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DD8EDBFB607B6" bold="true" box="[199,433,474,501]" pageId="45" pageNumber="296">Comparative notes.</emphasis>
Based on the very narrow median lobe and the parameres,
<taxonomicName id="EEF54D18FFC46D75D21BD8ECBAAF07B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1105,1192,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D21BD8ECBAAF07B6" box="[1105,1192,475,501]" italics="true" pageId="45" pageNumber="296">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D2E5D8ECBAF707B6" bold="true" box="[1199,1264,475,501]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D2E5D8ECBAF707B6" box="[1199,1264,475,501]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFC46D75D3CED8ECBF2C045B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D3CED8ECBB9207B7" box="[1412,1429,475,500]" italics="true" pageId="45" pageNumber="296">P</emphasis>
.
<emphasis id="1B81EA89FFC46D75D6DDD8C8BF2C045B" box="[151,299,511,536]" italics="true" pageId="45" pageNumber="296">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D778D8C8BF75045A" bold="true" box="[306,370,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D778D8C8BF75045A" box="[306,370,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFC46D75D7CAD8C8BFFF045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[384,504,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D7CAD8C8BFFF045A" box="[384,504,511,537]" italics="true" pageId="45" pageNumber="296">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D44AD8C8BC47045A" bold="true" box="[512,576,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D44AD8C8BC47045A" box="[512,576,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="EEF54D18FFC46D75D404D8C8BCDD045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[590,730,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D404D8C8BCDD045A" box="[590,730,511,537]" italics="true" pageId="45" pageNumber="296">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D4A8D8C8BD26045A" bold="true" box="[738,801,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D4A8D8C8BD26045A" box="[738,801,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="EEF54D18FFC46D75D512D8C8BDE9045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[856,1006,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D512D8C8BDE9045A" box="[856,1006,511,537]" italics="true" pageId="45" pageNumber="296">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D5BCD8C8BA31045A" bold="true" box="[1014,1078,511,537]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D5BCD8C8BA31045A" box="[1014,1078,511,537]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from all these species by the following morphological features:
</paragraph>
<paragraph id="294A369BFFC46D75D68DDB70BB0C0422" blockId="45.[151,1437,150,1077]" box="[199,1291,583,609]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D74FDB70BFBF0423" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,440,583,609]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D74FDB70BF110423" box="[261,278,583,608]" italics="true" pageId="45" pageNumber="296">P</emphasis>
.
<emphasis id="1B81EA89FFC46D75D76EDB70BFBF0423" box="[292,440,583,608]" italics="true" pageId="45" pageNumber="296">atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7F4DB70BFF90422" bold="true" box="[446,510,583,609]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7F4DB70BFF90422" box="[446,510,583,609]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the smaller and slightly paler body, and slightly longer parameres;
</paragraph>
<paragraph id="294A369BFFC46D75D68DDB5CBDD404C6" blockId="45.[151,1437,150,1077]" box="[199,979,619,645]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D74FDB5CBF7B04C6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,619,645]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D74FDB5CBF7B04C6" box="[261,380,619,645]" italics="true" pageId="45" pageNumber="296">P. barclayi</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7C9DB5CBFC404C6" bold="true" box="[387,451,619,645]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7C9DB5CBFC404C6" box="[387,451,619,645]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the darker elytra and the shorter parameres;
</paragraph>
<paragraph id="294A369BFFC46D75D68DDBB8BD47048E" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D749DBB8BF8B04EA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[259,396,655,681]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D749DBB8BF8B04EA" box="[259,396,655,681]" italics="true" pageId="45" pageNumber="296">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7DBDBB8BFC904EA" bold="true" box="[401,462,655,681]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7DBDBB8BFC904EA" box="[401,462,655,681]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the larger body, the presence of the punctation on the head and the stronger punctation of the pronotum, the shorter elytra and the longer parameres;
</paragraph>
<paragraph id="294A369BFFC46D75D68DDBE0BDC50556" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
from
<taxonomicName id="EEF54D18FFC46D75D749DBE0BF9F04B2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[259,408,727,753]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D749DBE0BF9F04B2" box="[259,408,727,753]" italics="true" pageId="45" pageNumber="296">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D7D7DBE0BFDB04B2" bold="true" box="[413,476,727,753]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D7D7DBE0BFDB04B2" box="[413,476,727,753]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the slightly darker body, the presence of the punctation on the head and the stronger punctation on the pronotum, the shorter elytra and the longer parameres.
</paragraph>
<paragraph id="294A369BFFC46D75D68DDA28BF2805C2" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
From all these species,
<taxonomicName id="EEF54D18FFC46D75D786DA28BC21057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[460,550,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFC46D75D786DA28BFDA057B" box="[460,477,799,824]" italics="true" pageId="45" pageNumber="296">P</emphasis>
.
<emphasis id="1B81EA89FFC46D75D7A1DA28BC21057A" box="[491,550,799,825]" italics="true" pageId="45" pageNumber="296">steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D467DA28BC6B057A" bold="true" box="[557,620,799,825]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D467DA28BC6B057A" box="[557,620,799,825]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the presence of elevations between punctures on the elytra, distinctly broader apical portions of the parameres and details of the external and internal morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC46D75D68DDABDBB4B05E6" box="[199,1356,906,933]" pageId="45" pageNumber="296" type="distribution">
<paragraph id="294A369BFFC46D75D68DDABDBB4B05E6" blockId="45.[151,1437,150,1077]" box="[199,1356,906,933]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DDABDBF6605E7" bold="true" box="[199,353,906,932]" pageId="45" pageNumber="296">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFC46D75D72DDABCBC6305E6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[359,612,907,933]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="296" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="1B81EA89FFC46D75D72DDABCBC6305E6" box="[359,612,907,933]" italics="true" pageId="45" pageNumber="296">Paraphloeostiba steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC46D75D421DABCBCAC05E6" bold="true" box="[619,683,907,933]" pageId="45" pageNumber="296">
<taxonomicNameLabel id="00B257F2FFC46D75D421DABCBCAC05E6" box="[619,683,907,933]" pageId="45" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F64E8839FFC46D75D5F4DABCBDE905E6" box="[958,1006,907,933]" pageId="45" pageNumber="296">type</typeStatus>
locality in
<collectingCountry id="51E2760BFFC46D75D227DABCBB4E05E6" box="[1133,1353,907,933]" name="Papua New Guinea" pageId="45" pageNumber="296">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC46D75D68DDA98BF390276" pageId="45" pageNumber="296" type="etymology">
<paragraph id="294A369BFFC46D75D68DDA98BD2B05AE" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DDA98BF49058A" bold="true" box="[199,334,943,969]" pageId="45" pageNumber="296">Etymology.</emphasis>
The species is named in the memory of the English coleopterologist William Owen Steel (1917– 1969) for his contributions to the knowledge of
<taxonomicName id="EEF54D18FFC46D75D4FADAE4BD2F05AE" box="[688,808,979,1005]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
.
</paragraph>
<paragraph id="294A369BFFC46D75D68DDAC1BF390276" blockId="45.[151,1437,150,1077]" pageId="45" pageNumber="296">
<emphasis id="1B81EA89FFC46D75D68DDAC1BF4C0253" bold="true" box="[199,331,1014,1040]" pageId="45" pageNumber="296">Bionomics.</emphasis>
<specimenCount id="3FF3FD12FFC46D75D705DAC0BFFD0252" box="[335,506,1015,1041]" count="2" pageId="45" pageNumber="296" type="generic">Two specimens</specimenCount>
were collected in swamp plants in marsh, and
<specimenCount id="3FF3FD12FFC46D75D5B9DAC0BA8D0252" box="[1011,1162,1015,1041]" count="1" pageId="45" pageNumber="296" type="generic">one specimen</specimenCount>
(
<typeStatus id="F64E8839FFC46D75D2DCDAC0BAFD0252" box="[1174,1274,1015,1041]" pageId="45" pageNumber="296" type="paratype">paratype</typeStatus>
) was collected in rotting fruit.
</paragraph>
</subSubSection>
</treatment>
</document>
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