plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/A1/39/9F/A1399F3AFA7FFFFD3CFD4FBFFA7DDF7D.xml

524 lines
55 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="4DD4F3D885753788E7DBA177F0403A84" ID-DOI="10.11646/zootaxa.5519.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13915657" ID-ZooBank="73D15FB1-651B-41D7-A47A-B002A20E9262" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1728584965993" checkinUser="plazi" docAuthor="Zhu, Pingzhou, Shi, Hongliang, Liu, Shengdong &amp; Liang, Hongbin" docDate="2024" docId="A1399F3AFA7FFFFD3CFD4FBFFA7DDF7D" docLanguage="en" docName="zootaxa.5519.1.4.pdf" docOrigin="Zootaxa 5519 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dolichina Brulle 1834" docType="treatment" docVersion="1" lastPageNumber="93" masterDocId="5D00E742FA7EFFFE3C6A4A51FFE1DE21" masterDocTitle="A new genus and a new species of Sphodrini (Coleoptera: Carabidae) from Changbai Mountain, China, with notes on the tribe" masterLastPageNumber="102" masterPageNumber="90" pageNumber="91" updateTime="1729000129229" updateUser="carolina" zenodo-license-document="CLOSED">
<mods:mods id="072B678CE321D887BC9BD7849AF498B6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B14FAB6E10DD925D44ACCF308C3E3B7A">
<mods:title id="E5F1D85D5C9C324C85D0BBD312961112">A new genus and a new species of Sphodrini (Coleoptera: Carabidae) from Changbai Mountain, China, with notes on the tribe</mods:title>
</mods:titleInfo>
<mods:name id="B7531F4B4063A6A0228723A8CDD8A9F6" type="personal">
<mods:role id="3F3E0E9C4E6E5B3C3F9E192F7D8EA96A">
<mods:roleTerm id="F410EA2137C788ABC65D8DA7055AC9CF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A70367A03D31F0E9455A0D8942D2BB7">Zhu, Pingzhou</mods:namePart>
<mods:affiliation id="2CE449291292F6C77EA2DE7E6B46C63F">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; College of Plant Protection, China Agricultural University, Beijing 100193, China</mods:affiliation>
</mods:name>
<mods:name id="7E9D7CB6C32DC0D814222CC5BFA4614B" type="personal">
<mods:role id="7BF8F5EDBC3DE35E0F4919D460266140">
<mods:roleTerm id="0C9B2CDCEB4C51A22D17A3DAB2ED00D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB4F5647BC27293697A389B6C777BF41">Shi, Hongliang</mods:namePart>
<mods:nameIdentifier id="DA91C584C5641B2298E7DBDD436A8AEF" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
<mods:affiliation id="8022792D51B1B32E8ADE4DCC1B32C857">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:nameIdentifier id="8527D0E437CC7CF036EB2A8A6D5491A4" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="4437CAF684C550A43587ABC8CC06C617" type="personal">
<mods:role id="981F8737206612202B61161D5A9E097E">
<mods:roleTerm id="753F0C09BCAE225690749040EFD4144D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F703DFD7A4BB28247218D48E25FF5614">Liu, Shengdong</mods:namePart>
<mods:nameIdentifier id="EFCFD1300B3EDEFCA8AFD4FA5547E689" type="ORCID">0000-0002-4699-2469</mods:nameIdentifier>
<mods:affiliation id="F1DF66F415A864CB037D98D155DB2897">Jilin Provincial Key Laboratory of Insect Biodiversity and Ecosystem Function of Changbai Mountains, Beihua University, Jilin 132013, China</mods:affiliation>
<mods:nameIdentifier id="9C9DEAF7DA08AE33E8464632D69ADD3E" type="email">liushengdong@beihua.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="F570D036997D8595623433BCB3CCD6D4" type="personal">
<mods:role id="0141F8A89CE106B016A9AC5886FBB968">
<mods:roleTerm id="5E41CDFFD8F82D6679E3182DDB26FBBB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C18BDCAD0FF73C54454E29BF50A26041">Liang, Hongbin</mods:namePart>
<mods:affiliation id="0242C628F8E1EAC622E95026F493F5E5">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D5B8DA7EBDA32BD0861475E39B55E007">text</mods:typeOfResource>
<mods:relatedItem id="A4FD2F2960A31C0E564071FE40F6A6D3" type="host">
<mods:titleInfo id="F20CB7F5BE5FD8E00983F8D8AD1CE2CF">
<mods:title id="99CC458830A09F8C72F39DB9FC22D869">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CC97FD75428FB1EEAFB447DFA983378D">
<mods:date id="1936FB6754BD65B1138671769895B087">2024</mods:date>
<mods:detail id="40DDFBDB2C9B507B5287D0559EB32F42" type="pubDate">
<mods:number id="9932464CE321B9856FFB1011F43465D6">2024-10-07</mods:number>
</mods:detail>
<mods:detail id="90C6529FD93C8D8369A188E632B5EE32" type="volume">
<mods:number id="8C4806C63885E20DA09FD5417B511456">5519</mods:number>
</mods:detail>
<mods:detail id="0E0AF2EB4BAB542B3682F72F154BCB43" type="issue">
<mods:number id="4583FB6AC31A58F8316E165199BCE005">1</mods:number>
</mods:detail>
<mods:extent id="4776BCF3DEC0FE9B3C24E719B6661187" unit="page">
<mods:start id="3B85F69BEA6A533F91804F129B68A382">90</mods:start>
<mods:end id="13A61EACCFE362B90401AC3F0D9E8B3F">102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C55CF016DE411F5FA1D88AD8DF48D18C">
<mods:url id="3936C17C781ED7425B2808047F981474">http://dx.doi.org/10.11646/zootaxa.5519.1.4</mods:url>
</mods:location>
<mods:classification id="FB3140E5101310EF65060FB49CEE8583">journal article</mods:classification>
<mods:identifier id="E208EBB92BCB6F8A475E261BA669C75B" type="DOI">10.11646/zootaxa.5519.1.4</mods:identifier>
<mods:identifier id="08C3E43914085E2A1FFA11A24F6454F7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="67B1A9E4341EE6B2296171086662D7B4" type="Zenodo-Dep">13915657</mods:identifier>
<mods:identifier id="EF71DD5EEA3433E6FCC11D5257B79D7D" type="ZooBank">73D15FB1-651B-41D7-A47A-B002A20E9262</mods:identifier>
</mods:mods>
<treatment id="A1399F3AFA7FFFFD3CFD4FBFFA7DDF7D" LSID="urn:lsid:plazi:treatment:A1399F3AFA7FFFFD3CFD4FBFFA7DDF7D" httpUri="http://treatment.plazi.org/id/A1399F3AFA7FFFFD3CFD4FBFFA7DDF7D" lastPageId="3" lastPageNumber="93" pageId="1" pageNumber="91">
<subSubSection id="618A7DA7FA7FFFFF3CFD4FBFFDC5D829" box="[151,548,1518,1545]" pageId="1" pageNumber="91" type="nomenclature">
<paragraph id="292F2E2CFA7FFFFF3CFD4FBFFDC5D829" blockId="1.[151,548,1518,1545]" box="[151,548,1518,1545]" pageId="1" pageNumber="91">
<heading id="72679940FA7FFFFF3CFD4FBFFDC5D829" bold="true" box="[151,548,1518,1545]" fontSize="11" level="1" pageId="1" pageNumber="91" reason="1">
<emphasis id="1BE4F23EFA7FFFFF3CFD4FBFFDC5D829" bold="true" box="[151,548,1518,1545]" pageId="1" pageNumber="91">
Subtribe
<taxonomicName id="EE9055AFFA7FFFFF3D664FBFFDC5D829" ID-CoL="8LTMK" authority="Brulle, 1834" authorityName="Brulle" authorityYear="1834" box="[268,548,1518,1545]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina Brullé, 1834</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="618A7DA7FA7FFFFF3CFD4C62FCCCD86A" box="[151,813,1587,1612]" pageId="1" pageNumber="91" type="type_taxon">
<paragraph id="292F2E2CFA7FFFFF3CFD4C62FCCCD86A" blockId="1.[151,813,1587,1612]" box="[151,813,1587,1612]" pageId="1" pageNumber="91">
<taxonomicName id="EE9055AFFA7FFFFF3CFD4C62FE57D86A" authority="Brulle, 1834: 295" authorityName="Brulle" authorityPageNumber="295" authorityYear="1834" box="[151,438,1587,1611]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina Brullé, 1834: 295</taxonomicName>
.
<typeStatus id="F62B908EFA7FFFFF3DAA4C65FE13D86D" box="[448,498,1588,1612]" pageId="1" pageNumber="91">Type</typeStatus>
genus:
<taxonomicName id="EE9055AFFA7FFFFF3E2B4C62FCCCD86A" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[577,813,1587,1611]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7FFFFF3E2B4C62FD7DD86A" box="[577,668,1587,1611]" italics="true" pageId="1" pageNumber="91">Dolichus</emphasis>
Bonelli, 1810
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="618A7DA7FA7FFFFF3CFD4C23FDC3D9A8" pageId="1" pageNumber="91" type="diagnosis">
<paragraph id="292F2E2CFA7FFFFF3CFD4C23FD91D8D8" blockId="1.[151,1437,1650,2037]" pageId="1" pageNumber="91">
<emphasis id="1BE4F23EFA7FFFFF3CFD4C23FEF3D8AD" bold="true" box="[151,274,1650,1676]" pageId="1" pageNumber="91">Diagnosis.</emphasis>
Adults of this subtribe can be recognized by combination of the following characters: (1) stridulatory organ absent on base of head; (2) lateral bead of prosternal process absent in most species; (3) claws pectinate; (4) aedeagus left side up in repose; (5) right paramere of aedeagus styloid; and (6) gonocoxite II with sensory pit absent or represented by a very small depression.
</paragraph>
<paragraph id="292F2E2CFA7FFFFF3CAD4D52FDC3D9A8" blockId="1.[151,1437,1650,2037]" pageId="1" pageNumber="91">
Based on the pectinate claws and styloid right paramere,
<taxonomicName id="EE9055AFFA7FFFFF3F234D52FC56D93C" authorityName="Brulle" authorityYear="1834" box="[841,951,1795,1821]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
may be most similar to subtribe
<taxonomicName id="EE9055AFFA7FFFFF39404D52FA79D93C" authorityName="Laporte" authorityYear="1834" box="[1322,1432,1795,1821]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="subTribe" subTribe="Calathina">Calathina</taxonomicName>
, but can be distinguished from the latter by the markedly reduced sensory pit of gonocoxite II. Whereas,
<taxonomicName id="EE9055AFFA7FFFFF39444D76FA7DD960" authorityName="Brulle" authorityYear="1834" box="[1326,1436,1831,1857]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
shares the markedly reduced sensory pit of gonocoxite II with the subtribe
<taxonomicName id="EE9055AFFA7FFFFF3FBB4D1AFBABD944" authorityName="Lindroth" authorityYear="1956" box="[977,1098,1867,1893]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="subTribe" subTribe="Synuchina">Synuchina</taxonomicName>
, but differs by the shape of the styloid right paramere of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="618A7DA7FA7FFFFC3CAD4DC3FC33DC41" lastPageId="2" lastPageNumber="92" pageId="1" pageNumber="91" type="distribution">
<paragraph id="292F2E2CFA7FFFFF3CAD4DC3FBCED9D4" blockId="1.[151,1437,1650,2037]" pageId="1" pageNumber="91">
<emphasis id="1BE4F23EFA7FFFFF3CAD4DC3FE80D98D" bold="true" box="[199,353,1938,1964]" pageId="1" pageNumber="91">Distribution.</emphasis>
Most of the genera and species of subtribe
<taxonomicName id="EE9055AFFA7FFFFF3F084DC2FC31D98C" authorityName="Brulle" authorityYear="1834" box="[866,976,1939,1965]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
are distributed in the eastern Palearctic Region, except for
<taxonomicName id="EE9055AFFA7FFFFF3D044DE6FDD7D9F0" baseAuthorityName="Schaller" baseAuthorityYear="1783" box="[366,566,1975,2001]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="halensis">
<emphasis id="1BE4F23EFA7FFFFF3D044DE6FDD7D9F0" box="[366,566,1975,2001]" italics="true" pageId="1" pageNumber="91">Dolichus halensis</emphasis>
</taxonomicName>
(a widespread species in the Palearctic Region, from Eastern Asia to Western Europe), and the genus
<taxonomicName id="EE9055AFFA7FFFFF3DCA4D8AFDACD9D4" baseAuthorityName="Wrase &amp; Assmann" baseAuthorityYear="2001" box="[416,589,2011,2037]" class="Insecta" family="Carabidae" genus="Anchomenidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7FFFFF3DCA4D8AFDACD9D4" box="[416,589,2011,2037]" italics="true" pageId="1" pageNumber="91">Anchomenidius</emphasis>
</taxonomicName>
(two species endemic to northwest
<collectingCountry id="51876EBCFA7FFFFF3FB44D8AFBC2D9D4" box="[990,1059,2011,2037]" name="Spain" pageId="1" pageNumber="91">Spain</collectingCountry>
).
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD4AC7FEE6DF45" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
<emphasis id="1BE4F23EFA7CFFFC3CAD4AC7FDD2DE91" bold="true" box="[199,563,150,176]" pageId="2" pageNumber="92">Monophyly and relationships.</emphasis>
From the morphological respect, the only character that might support the monophyly of
<taxonomicName id="EE9055AFFA7CFFFC3D294AEBFE50DEF5" authorityName="Brulle" authorityYear="1834" box="[323,433,186,212]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
is the reduced sensory pit on female gonocoxite II. However, the single origin of the character state remains unproven (Schmidt &amp; Will 2020;
<bibRefCitation id="4D0153DDFA7CFFFC3F7D4A8FFC16DED9" author="Schmidt, J. &amp; Scholz, S. &amp; Will, K." box="[791,1015,222,248]" pageId="2" pageNumber="92" pagination="19 - 44" refId="ref10021" refString="Schmidt, J., Scholz, S. &amp; Will, K. (2022) Character analysis and descriptions of Eocene sphodrine fossils (Coleoptera, Carabidae) using light microscopy, micro-CT scanning, and 3 D imaging. Deutsche Entomologische Zeitschrift, 69 (1), 19 - 44. https: // doi. org / 10.3897 / dez. 69.79931" type="journal article" year="2022">
Schmidt
<emphasis id="1BE4F23EFA7CFFFC3F114A8EFC52DED9" box="[891,947,222,248]" italics="true" pageId="2" pageNumber="92">et al.</emphasis>
2022
</bibRefCitation>
). In the molecular research, the genus
<taxonomicName id="EE9055AFFA7CFFFC3CFD4B53FEA5DF3D" baseAuthorityName="Wrase &amp; Assmann" baseAuthorityYear="2001" box="[151,324,258,284]" class="Insecta" family="Carabidae" genus="Anchomenidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3CFD4B53FEA5DF3D" box="[151,324,258,284]" italics="true" pageId="2" pageNumber="92">Anchomenidius</emphasis>
</taxonomicName>
is always separated from other genera of
<taxonomicName id="EE9055AFFA7CFFFC3F644B53FC9DDF3D" authorityName="Brulle" authorityYear="1834" box="[782,892,258,284]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
but close to either
<taxonomicName id="EE9055AFFA7CFFFC38264B53FB5EDF3D" authorityName="Laporte" authorityYear="1834" box="[1100,1215,258,284]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Sphodrina">Sphodrina</taxonomicName>
or
<taxonomicName id="EE9055AFFA7CFFFC38884B53FAB9DF3D" authorityName="Lindroth" authorityYear="1956" box="[1250,1368,258,284]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Pristosiina">Pristosiina</taxonomicName>
(Ruiz e
<emphasis id="1BE4F23EFA7CFFFC3CCE4B76FF31DF61" box="[164,208,294,320]" italics="true" pageId="2" pageNumber="92">t al.</emphasis>
2009). In a word, the monophyly of
<taxonomicName id="EE9055AFFA7CFFFC3E1A4B77FD3FDF61" authorityName="Brulle" authorityYear="1834" box="[624,734,294,320]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
and its relationships with other subtribes of
<taxonomicName id="EE9055AFFA7CFFFC38B94B77FAA0DF61" authorityName="Laporte" authorityYear="1834" box="[1235,1345,294,320]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="tribe" tribe="Sphodrini">Sphodrini</taxonomicName>
are still unknown.
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD4B3EFC3BDFF1" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
By the way, the “
<taxonomicName id="EE9055AFFA7CFFFC3DE34B3EFDF4DFA9" box="[393,533,366,392]" class="Insecta" family="Carabidae" genus="Xestopus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="1BE4F23EFA7CFFFC3DE34B3EFE0FDFA9" box="[393,494,367,392]" italics="true" pageId="2" pageNumber="92">Xestopus</emphasis>
sp.
</taxonomicName>
” in Ruiz’s study is more likely a misidentification of
<taxonomicName id="EE9055AFFA7CFFFC38044B3FFA9EDFA9" authorityName="Semenov" authorityYear="1889" box="[1134,1407,366,392]" class="Insecta" family="Carabidae" genus="Morphodactyla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="1BE4F23EFA7CFFFC38044B3FFA9EDFA9" box="[1134,1407,366,392]" italics="true" pageId="2" pageNumber="92">Morphodactyla potanini</emphasis>
</taxonomicName>
in my (PZ Zhu) point of view, for the former is a genus endemic to the southeastern edge of the Tibetan Plateau and the latter is a very common species in Qinling Mountains,
<collectingRegion id="EB54E0CEFA7CFFFC3F4D4BE7FC65DFF1" box="[807,900,438,464]" country="China" name="Shaanxi" pageId="2" pageNumber="92">Shaanxi</collectingRegion>
,
<collectingCountry id="51876EBCFA7CFFFC3FE54BE7FC37DFF1" box="[911,982,438,464]" name="China" pageId="2" pageNumber="92">China</collectingCountry>
.
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD4B8BFC33DC41" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
<emphasis id="1BE4F23EFA7CFFFC3CAD4B8BFE79DFD5" bold="true" box="[199,408,474,500]" pageId="2" pageNumber="92">Genera included.</emphasis>
<taxonomicName id="EE9055AFFA7CFFFC3DC94B8BFDF5DFD5" authorityName="Brulle" authorityYear="1834" box="[419,532,474,500]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
, a subtribe within
<taxonomicName id="EE9055AFFA7CFFFC3E9B4B8BFC83DFD5" authorityName="Laporte" authorityYear="1834" box="[753,866,474,500]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="tribe" tribe="Sphodrini">Sphodrini</taxonomicName>
, is represented by 35 species (including the new species described in the present paper). Eight genera are presently included in
<taxonomicName id="EE9055AFFA7CFFFC38594BAFFB47DC39" authorityName="Brulle" authorityYear="1834" box="[1075,1190,510,536]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
:
<taxonomicName id="EE9055AFFA7CFFFC38DE4BAFFA79DC39" authority="Semenov" authorityName="Semenov" authorityYear="1889" box="[1204,1432,510,536]" class="Insecta" family="Carabidae" genus="Acalathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC38DE4BAFFAC5DC39" box="[1204,1316,510,536]" italics="true" pageId="2" pageNumber="92">Acalathus</emphasis>
Semenov
</taxonomicName>
,
<taxonomicName id="EE9055AFFA7CFFFC3CFD4873FE43DC1D" authority="Heyden" authorityName="Heyden" authorityYear="1880" box="[151,418,546,572]" class="Insecta" family="Carabidae" genus="Anchomenidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3CFD4873FEA5DC1D" box="[151,324,546,572]" italics="true" pageId="2" pageNumber="92">Anchomenidius</emphasis>
Heyden
</taxonomicName>
,
<taxonomicName id="EE9055AFFA7CFFFC3DC04873FD2EDC1C" authority="Sciaky &amp; Wrase" authorityName="Sciaky &amp; Wrase" authorityYear="1998" box="[426,719,546,573]" class="Insecta" family="Carabidae" genus="Casaleius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3DC04873FDF9DC1D" box="[426,536,546,572]" italics="true" pageId="2" pageNumber="92">Casaleius</emphasis>
Sciaky &amp;Wrase
</taxonomicName>
,
<taxonomicName id="EE9055AFFA7CFFFC3EB24873FC77DC1D" authority="Bonelli" authorityName="Bonelli" authorityYear="1810" box="[728,918,546,572]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3EB24873FCDCDC1D" box="[728,829,546,572]" italics="true" pageId="2" pageNumber="92">Dolichus</emphasis>
Bonelli
</taxonomicName>
,
<taxonomicName id="EE9055AFFA7CFFFC3FF44873FB06DC1C" authority="Sciaky &amp; Wrase" authorityName="Sciaky &amp; Wrase" authorityYear="1998" box="[926,1255,546,573]" class="Insecta" family="Carabidae" genus="Doliodactyla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3FF44873FBD1DC1D" box="[926,1072,546,572]" italics="true" pageId="2" pageNumber="92">Doliodactyla</emphasis>
Sciaky &amp;Wrase
</taxonomicName>
,
<taxonomicName id="EE9055AFFA7CFFFC389A4873FEE1DC41" authority="Semenov" authorityName="Semenov" authorityYear="1889" class="Insecta" family="Carabidae" genus="Morphodactyla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC389A4873FA7CDC1D" box="[1264,1437,546,572]" italics="true" pageId="2" pageNumber="92">Morphodactyla</emphasis>
Semenov
</taxonomicName>
,
<taxonomicName id="EE9055AFFA7CFFFC3D664816FE0BDC41" authority="Andrewes" authorityName="Andrewes" authorityYear="1937" box="[268,490,582,608]" class="Insecta" family="Carabidae" genus="Xestopus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3D664816FE90DC41" box="[268,369,583,608]" italics="true" pageId="2" pageNumber="92">Xestopus</emphasis>
Andrewes
</taxonomicName>
, and the below described
<taxonomicName id="EE9055AFFA7CFFFC3F664816FC83DC41" authorityName="Zhu &amp; Shi &amp; Liu &amp; Liang" authorityYear="2024" box="[780,866,583,608]" class="Insecta" family="Carabidae" genus="Peiyuia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BE4F23EFA7CFFFC3F664816FC83DC41" box="[780,866,583,608]" italics="true" pageId="2" pageNumber="92">Peiyuia</emphasis>
</taxonomicName>
<emphasis id="1BE4F23EFA7CFFFC3F034817FC33DC41" bold="true" box="[873,978,582,608]" pageId="2" pageNumber="92">
<taxonomicNameLabel id="00D74F45FA7CFFFC3F034817FC33DC41" box="[873,978,582,608]" pageId="2" pageNumber="92" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="618A7DA7FA7CFFFC3CAD483BFC13DBBD" pageId="2" pageNumber="92" type="discussion">
<paragraph id="292F2E2CFA7CFFFC3CAD483BFADDDCD1" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
<emphasis id="1BE4F23EFA7CFFFC3CAD483BFEDADCA5" bold="true" box="[199,315,618,644]" pageId="2" pageNumber="92">Remarks.</emphasis>
Schmidt and Will (2020) provided a summary of important morphological characters for all the genera and subgenera of the subtribes
<taxonomicName id="EE9055AFFA7CFFFC3E2048DFFD54DC89" authorityName="Laporte" authorityYear="1834" box="[586,693,654,680]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Calathina">Calathina</taxonomicName>
and
<taxonomicName id="EE9055AFFA7CFFFC3E8448DFFCBEDC89" authorityName="Brulle" authorityYear="1834" box="[750,863,654,680]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
, which have benefitted the continued taxonomy of
<taxonomicName id="EE9055AFFA7CFFFC3CFD48E3FEE8DCED" authorityName="Laporte" authorityYear="1834" box="[151,265,690,716]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="tribe" tribe="Sphodrini">Sphodrini</taxonomicName>
. However, some of these characters are inaccurate or incorrect for
<taxonomicName id="EE9055AFFA7CFFFC387948E3FB65DCED" authorityName="Brulle" authorityYear="1834" box="[1043,1156,690,716]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
, based on the numerous specimens we have examined in this research. Herein, we would like to revise these characters as follows:
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD48ABFB58DD7D" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
(1) The shape of mentum tooth shows intraspecific variation in some species. It is simple in most specimens of
<taxonomicName id="EE9055AFFA7CFFFC3CDF494FFE9CDD19" baseAuthorityName="Schaller" baseAuthorityYear="1783" box="[181,381,798,824]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="halensis">
<emphasis id="1BE4F23EFA7CFFFC3CDF494FFE9CDD19" box="[181,381,798,824]" italics="true" pageId="2" pageNumber="92">Dolichus halensis</emphasis>
</taxonomicName>
and
<taxonomicName id="EE9055AFFA7CFFFC3DD9494EFDC7DD19" baseAuthorityName="Fairmaire" baseAuthorityYear="1889" box="[435,550,798,824]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="davidis">
<emphasis id="1BE4F23EFA7CFFFC3DD9494EFDC7DD19" box="[435,550,798,824]" italics="true" pageId="2" pageNumber="92">D. davidis</emphasis>
</taxonomicName>
, but can be slightly emarginate or bifid in a few specimens. While, it is bifid in most specimens of
<taxonomicName id="EE9055AFFA7CFFFC3D054913FD9EDD7D" authorityName="Semenov" authorityYear="1889" box="[367,639,834,860]" class="Insecta" family="Carabidae" genus="Morphodactyla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="1BE4F23EFA7CFFFC3D054913FD9EDD7D" box="[367,639,834,860]" italics="true" pageId="2" pageNumber="92">Morphodactyla potanini</emphasis>
</taxonomicName>
, but a few specimens are only slightly emarginate.
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD4937FAA7DD84" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
(2) The pronotal laterobasal setae show interspecific variation in the subgenus
<taxonomicName id="EE9055AFFA7CFFFC380D4937FB0EDDA1" authorityName="Jedlicka" authorityYear="1954" box="[1127,1263,870,896]" class="Insecta" family="Carabidae" genus="Procalathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC380D4937FB0EDDA1" box="[1127,1263,870,896]" italics="true" pageId="2" pageNumber="92">Procalathus</emphasis>
</taxonomicName>
and the genus
<taxonomicName id="EE9055AFFA7CFFFC3CFD49DBFEA5DD85" authorityName="Semenov" authorityYear="1889" box="[151,324,906,932]" class="Insecta" family="Carabidae" genus="Morphodactyla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3CFD49DBFEA5DD85" box="[151,324,906,932]" italics="true" pageId="2" pageNumber="92">Morphodactyla</emphasis>
</taxonomicName>
. They can be removed from lateral margins or situated along lateral margins in two groups.
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD49FFFF08DDCD" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
(3) Prosternal process of the lateral bead is present in
<taxonomicName id="EE9055AFFA7CFFFC3F2E49FFFC48DDE9" authorityName="Bonelli" authorityYear="1810" box="[836,937,942,968]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3F2E49FFFC48DDE9" box="[836,937,942,968]" italics="true" pageId="2" pageNumber="92">Dolichus</emphasis>
</taxonomicName>
as observed by
<bibRefCitation id="4D0153DDFA7CFFFC380149FFFAC3DDE8" author="Lindroth, C. H." box="[1131,1314,942,969]" pageId="2" pageNumber="92" pagination="485 - 576" refId="ref9494" refString="Lindroth, C. H. (1956) A revision of the genus Synuchus Gyllenhal (Coleoptera: Carabidae) in the widest sense, with notes on Pristosia Motschulsky (Eucalathus Bates) and Calathus Bonelli. The Transactions of the Royal Entomological Society of London, 108, 485 - 576. https: // doi. org / 10.1111 / j. 1365 - 2311.1956. tb 01274. x" type="journal article" year="1956">Lindroth (1956)</bibRefCitation>
and
<bibRefCitation id="4D0153DDFA7CFFFC390A49FFFF07DDCD" author="Habu, A." pageId="2" pageNumber="92" refId="ref9141" refString="Habu, A. (1978) Fauna Japonica. Carabidae: Platynini (Insecta: Coleoptera). Tokyo: Keigaku Publishing Co., vii + 447 pp., xxxvi pls." type="book" year="1978">Habu (1978)</bibRefCitation>
.
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD49A7FF3EDA5D" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
(4) The number of setigerous pores on elytral interval 3 shows both interspecific and intraspecific variation in
<taxonomicName id="EE9055AFFA7CFFFC3CFD4E4BFEE6DA15" authorityName="Semenov" authorityYear="1889" box="[151,263,1050,1076]" class="Insecta" family="Carabidae" genus="Acalathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3CFD4E4BFEE6DA15" box="[151,263,1050,1076]" italics="true" pageId="2" pageNumber="92">Acalathus</emphasis>
</taxonomicName>
and
<taxonomicName id="EE9055AFFA7CFFFC3D574E4BFE0BDA15" authorityName="Semenov" authorityYear="1889" box="[317,490,1050,1076]" class="Insecta" family="Carabidae" genus="Morphodactyla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3D574E4BFE0BDA15" box="[317,490,1050,1076]" italics="true" pageId="2" pageNumber="92">Morphodactyla</emphasis>
</taxonomicName>
. Two is common number, but one, three, four or absent occurs in a few specimens of different species. For example,
<taxonomicName id="EE9055AFFA7CFFFC3D934E6FFCD0DA79" authorityName="Lassalle" authorityYear="2011" box="[505,817,1086,1112]" class="Insecta" family="Carabidae" genus="Morphodactyla" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="yulongensis">
<emphasis id="1BE4F23EFA7CFFFC3D934E6FFCD0DA79" box="[505,817,1086,1112]" italics="true" pageId="2" pageNumber="92">Morphodactyla yulongensis</emphasis>
</taxonomicName>
has 3–4 setigerous pores on elytral interval 3 (
<bibRefCitation id="4D0153DDFA7CFFFC39554E6FFF32DA5D" author="Lassalle, B." pageId="2" pageNumber="92" pagination="291 - 296" refId="ref9462" refString="Lassalle, B. (2011) Nouvelles especes de Morphodactyla, de Procalathus et d' Acalathus de Chine (Coleoptera Carabidae). L'Entomologiste, 67, 291 - 296." type="journal article" year="2011">Lassalle 2011</bibRefCitation>
).
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD4ED7FEE5DAE5" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
(5) The metepisternum is always elongated in genus
<taxonomicName id="EE9055AFFA7CFFFC3F744ED7FC62DA81" authorityName="Bonelli" authorityYear="1810" box="[798,899,1158,1184]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3F744ED7FC62DA81" box="[798,899,1158,1184]" italics="true" pageId="2" pageNumber="92">Dolichus</emphasis>
</taxonomicName>
but in
<taxonomicName id="EE9055AFFA7CFFFC3FBF4ED6FBDBDA81" authorityName="Andrewes" authorityYear="1937" box="[981,1082,1159,1184]" class="Insecta" family="Carabidae" genus="Xestopus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3FBF4ED6FBDBDA81" box="[981,1082,1159,1184]" italics="true" pageId="2" pageNumber="92">Xestopus</emphasis>
</taxonomicName>
it is elongated or short (
<bibRefCitation id="4D0153DDFA7CFFFC393B4ED7FF16DAE5" author="Zhu, P. Z. &amp; Kavanaugh, D. H. &amp; Liang, H. B." pageId="2" pageNumber="92" pagination="139 - 151" refId="ref10215" refString="Zhu, P. Z., Kavanaugh, D. H. &amp; Liang, H. B. (2021) Notes on the genus Xestopus from China, with description of a new species (Carabidae, Sphodrini, Dolichina). ZooKeys, 1009, 139 - 151. https: // doi. org / 10.3897 / zookeys. 1009.61515" type="journal article" year="2021">
Zhu
<emphasis id="1BE4F23EFA7CFFFC39E24ED6FF55DAE5" italics="true" pageId="2" pageNumber="92">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CAD4E9FFC13DBBD" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="92">
Although not listed in Schmidt and Will (2020), the form of claws is a very important character in
<taxonomicName id="EE9055AFFA7CFFFC394D4E9FFA78DAC9" authorityName="Laporte" authorityYear="1834" box="[1319,1433,1230,1256]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="tribe" tribe="Sphodrini">Sphodrini</taxonomicName>
.
<taxonomicName id="EE9055AFFA7CFFFC3CFD4EA3FEE4DB2D" authorityName="Sciaky &amp; Wrase" authorityYear="1998" box="[151,261,1266,1292]" class="Insecta" family="Carabidae" genus="Casaleius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3CFD4EA3FEE4DB2D" box="[151,261,1266,1292]" italics="true" pageId="2" pageNumber="92">Casaleius</emphasis>
</taxonomicName>
is the only genus with smooth claws in all genera of both
<taxonomicName id="EE9055AFFA7CFFFC3FF64EA3FBEBDB2D" authorityName="Brulle" authorityYear="1834" box="[924,1034,1266,1292]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
and
<taxonomicName id="EE9055AFFA7CFFFC38284EA3FB51DB2D" authorityName="Laporte" authorityYear="1834" box="[1090,1200,1266,1292]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Calathina">Calathina</taxonomicName>
, according to
<bibRefCitation id="4D0153DDFA7CFFFC393A4EA3FE81DB11" author="Sciaky, R. &amp; Wrase, D. W." pageId="2" pageNumber="92" pagination="221 - 232" refId="ref10126" refString="Sciaky, R. &amp; Wrase, D. W. (1998) Two new genera of Sphodrini Dolichina from China (Coleoptera Carabidae). Bolletino della Societa Entomologica Italiana, 130 (3), 221 - 232." type="journal article" year="1998">Sciaky and Wrase (1998)</bibRefCitation>
. But in
<specimenCount id="3F96E5A5FA7CFFFC3DDE4F46FD8FDB11" box="[436,622,1302,1329]" count="7" pageId="2" pageNumber="92" type="generic">seven specimens</specimenCount>
we examined, there are always tiny but visible teeth on their claws. More materials are needed to judge whether it is an intraspecific variation or misinterpretation. On the other hand, the claws are not mentioned in the description of
<taxonomicName id="EE9055AFFA7CFFFC3ECC4F0FFB1EDB59" authority="(Wrase &amp; Assmann 2001)" baseAuthorityName="Wrase &amp; Assmann" baseAuthorityYear="2001" box="[678,1279,1374,1400]" class="Insecta" family="Carabidae" genus="Anchomenidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="feldmanni">
<emphasis id="1BE4F23EFA7CFFFC3ECC4F0FFC2DDB59" box="[678,972,1374,1400]" italics="true" pageId="2" pageNumber="92">Anchomenidius feldmanni</emphasis>
(
<bibRefCitation id="4D0153DDFA7CFFFC3FB64F0FFB16DB59" author="Wrase, D. W. &amp; Assmann, T." box="[988,1271,1374,1400]" pageId="2" pageNumber="92" pagination="1 - 9" refId="ref10168" refString="Wrase, D. W. &amp; Assmann, T. (2001) Description of a new species of Anchomenidius Heyden from the Montes de Leon in northwest Spain (Coleoptera: Carabidae, Carabinae). Koleopterologische Rundschau, 71, 1 - 9." type="journal article" year="2001">Wrase &amp; Assmann 2001</bibRefCitation>
)
</taxonomicName>
, of which the freehand sketching seems to show smooth claws but still need to be verified.
</paragraph>
</subSubSection>
<subSubSection id="618A7DA7FA7CFFFD3CFD4FBFFA7DDF7D" lastPageId="3" lastPageNumber="93" pageId="2" pageNumber="92" type="key">
<paragraph id="292F2E2CFA7CFFFC3CFD4FBFFD65D829" blockId="2.[151,644,1518,1544]" box="[151,644,1518,1544]" pageId="2" pageNumber="92">
<heading id="72679940FA7CFFFC3CFD4FBFFD65D829" bold="true" box="[151,644,1518,1544]" fontSize="11" level="1" pageId="2" pageNumber="92" reason="1">
<emphasis id="1BE4F23EFA7CFFFC3CFD4FBFFD65D829" bold="true" box="[151,644,1518,1544]" pageId="2" pageNumber="92">
Key to genera of the subtribe
<taxonomicName id="EE9055AFFA7CFFFC3E614FBFFD65D829" ID-CoL="8LTMK" authorityName="Brulle" authorityYear="1834" box="[523,644,1518,1544]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Dolichina">Dolichina</taxonomicName>
</emphasis>
</heading>
</paragraph>
<keyStep id="92643689FA7CFFFC3CFD4C62FA7DD89F" pageId="2" pageNumber="92">
<paragraph id="292F2E2CFA7CFFFC3CFD4C62FA7DD847" blockId="2.[151,1437,1587,2020]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4C62FA7DD847" pageId="2" pageNumber="92">
1. Size small, usually BL &lt;
<quantity id="EE6883C9FA7CFFFC3DB24C62FDF1D868" box="[472,528,1587,1609]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="92" unit="mm" value="9.0">9 mm</quantity>
(only
<taxonomicName id="EE9055AFFA7CFFFC3E3B4C62FCADD869" ID-CoL="DKVP" baseAuthorityName="Wrase &amp; Assmann" baseAuthorityYear="2001" box="[593,844,1587,1608]" class="Insecta" family="Carabidae" genus="Anchomenidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="feldmanni">
<emphasis id="1BE4F23EFA7CFFFC3E3B4C62FCADD869" box="[593,844,1587,1608]" italics="true" pageId="2" pageNumber="92">Anchomenidius feldmanni</emphasis>
</taxonomicName>
(
<collectingCountry id="51876EBCFA7CFFFC3F304C62FC72D868" box="[858,915,1587,1609]" name="Spain" pageId="2" pageNumber="92">Spain</collectingCountry>
) is slightly larger, BL =
<quantity id="EE6883C9FA7CFFFC38E94C62FAE6D869" box="[1155,1287,1587,1608]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.03" metricValueMin="0.97" pageId="2" pageNumber="92" unit="mm" value="10.0" valueMax="10.3" valueMin="9.7">9.7–10.3 mm</quantity>
); body more or less depigmented; apical hook of the right paramere present (though very small in
<taxonomicName id="EE9055AFFA7CFFFC3F934C01FBB9D844" ID-CoL="8KVW8" authorityName="Sciaky &amp; Wrase" authorityYear="1998" box="[1017,1112,1616,1637]" class="Insecta" family="Carabidae" genus="Casaleius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC3F934C01FBB9D844" box="[1017,1112,1616,1637]" italics="true" pageId="2" pageNumber="92">Casaleius</emphasis>
</taxonomicName>
and
<taxonomicName id="EE9055AFFA7CFFFC38EC4C01FAFBD844" ID-CoL="8KV6M" authorityName="Heyden" authorityYear="1880" box="[1158,1306,1616,1637]" class="Insecta" family="Carabidae" genus="Anchomenidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC38EC4C01FAFBD844" box="[1158,1306,1616,1637]" italics="true" pageId="2" pageNumber="92">Anchomenidius</emphasis>
</taxonomicName>
)......... 2
</keyLead>
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CFD4C3FFA7DD89F" blockId="2.[151,1437,1587,2020]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4C3FFA7DD89F" pageId="2" pageNumber="92">
- Size large, usually BL&gt;
<quantity id="EE6883C9FA7CFFFC3DBE4C3FFDF9D8A2" box="[468,536,1646,1667]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="92" unit="mm" value="10.0">10 mm</quantity>
(only
<taxonomicName id="EE9055AFFA7CFFFC3E334C3FFC56D8A5" ID-CoL="8QLB" authority="(Qilian Mountains)" baseAuthorityName="Qilian Mountains" box="[601,951,1646,1668]" class="Insecta" family="Carabidae" genus="Acalathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="1BE4F23EFA7CFFFC3E334C3FFD16D8A2" box="[601,759,1646,1667]" italics="true" pageId="2" pageNumber="92">Acalathus fallax</emphasis>
(Qilian Mountains)
</taxonomicName>
is slightly smaller, BL =
<quantity id="EE6883C9FA7CFFFC38DA4C3FFAD5D8A2" box="[1200,1332,1646,1667]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.21" metricValueMin="0.84" pageId="2" pageNumber="92" unit="mm" value="10.25" valueMax="12.1" valueMin="8.4">8.4–12.1 mm</quantity>
); body not depigmented; apical hook of the right paramere absent (body slightly depigmented and apical hook of the right paramere distinctly present in some
<taxonomicName id="EE9055AFFA7CFFFC3DB44CF8FD63D89F" box="[478,642,1704,1726]" pageId="2" pageNumber="92">
<emphasis id="1BE4F23EFA7CFFFC3DB44CF8FDD4D89F" box="[478,565,1705,1726]" italics="true" pageId="2" pageNumber="92">Xestopus</emphasis>
species
</taxonomicName>
, but their BL&gt;
<quantity id="EE6883C9FA7CFFFC3F704CF9FCBED89F" box="[794,863,1704,1726]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="92" unit="mm" value="16.0">16 mm</quantity>
).............................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="92643689FA7CFFFC3CFD4C97FA7DD8D9" pageId="2" pageNumber="92">
<paragraph id="292F2E2CFA7CFFFC3CFD4C97FA7CD8FA" blockId="2.[151,1437,1587,2020]" box="[151,1437,1734,1756]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4C97FA7CD8FA" box="[151,1437,1734,1756]" pageId="2" pageNumber="92">
2. Elytra striae obliterated; interval 3 without setigerous pore;
<collectingCountry id="51876EBCFA7CFFFC3F754C97FCB9D8FD" box="[799,856,1734,1756]" name="China" pageId="2" pageNumber="92">China</collectingCountry>
(
<collectingRegion id="EB54E0CEFA7CFFFC3F0F4C97FC4CD8FD" box="[869,941,1734,1756]" country="China" name="Xizang" pageId="2" pageNumber="92">Xizang</collectingRegion>
)................................
<taxonomicName id="EE9055AFFA7CFFFC39544C97FA7CD8FA" ID-CoL="8KVW8" authorityName="Sciaky &amp; Wrase" authorityYear="1998" box="[1342,1437,1734,1755]" class="Insecta" family="Carabidae" genus="Casaleius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC39544C97FA7CD8FA" box="[1342,1437,1734,1755]" italics="true" pageId="2" pageNumber="92">Casaleius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CFD4CB5FA7DD8D9" blockId="2.[151,1437,1587,2020]" box="[151,1436,1763,1785]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4CB5FA7DD8D9" box="[151,1436,1763,1785]" pageId="2" pageNumber="92">
- Elytra striae distinct; interval 3 with 2 setigerous pores;
<collectingCountry id="51876EBCFA7CFFFC3E964CB2FCD4D8D8" box="[764,821,1763,1785]" name="China" pageId="2" pageNumber="92">China</collectingCountry>
(
<collectingRegion id="EB54E0CEFA7CFFFC3F284CB2FC8FD8D8" box="[834,878,1763,1785]" country="China" name="Jilin" pageId="2" pageNumber="92">Jilin</collectingRegion>
) and
<collectingCountry id="51876EBCFA7CFFFC3FC94CB2FC3BD8D8" box="[931,986,1763,1785]" name="Spain" pageId="2" pageNumber="92">Spain</collectingCountry>
.................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="92643689FA7CFFFC3CFD4D50FA7DD94C" pageId="2" pageNumber="92">
<paragraph id="292F2E2CFA7CFFFC3CFD4D50FA7DD912" blockId="2.[151,1437,1587,2020]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4D50FA7DD912" pageId="2" pageNumber="92">
3. Pronotal laterobasal setae removed from lateral margins; apical hook of the right paramere large and distinct; parascutellar setae always present;
<collectingCountry id="51876EBCFA7CFFFC3D164D4FFE54D915" box="[380,437,1822,1844]" name="China" pageId="2" pageNumber="92">China</collectingCountry>
(
<collectingRegion id="EB54E0CEFA7CFFFC3DA94D4FFE0ED915" box="[451,495,1822,1844]" country="China" name="Jilin" pageId="2" pageNumber="92">Jilin</collectingRegion>
)...............................................................
<taxonomicName id="EE9055AFFA7CFFFC38994D4FFADDD912" authorityName="Zhu &amp; Shi &amp; Liu &amp; Liang" authorityYear="2024" box="[1267,1340,1822,1843]" class="Insecta" family="Carabidae" genus="Peiyuia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1BE4F23EFA7CFFFC38994D4FFADDD912" box="[1267,1340,1822,1843]" italics="true" pageId="2" pageNumber="92">Peiyuia</emphasis>
</taxonomicName>
<emphasis id="1BE4F23EFA7CFFFC39284D4FFA7DD912" bold="true" box="[1346,1436,1822,1843]" pageId="2" pageNumber="92">
<taxonomicNameLabel id="00D74F45FA7CFFFC39284D4FFA7DD912" box="[1346,1436,1822,1843]" pageId="2" pageNumber="92" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CFD4D6DFA7DD94C" blockId="2.[151,1437,1587,2020]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4D6DFA7DD94C" pageId="2" pageNumber="92">
- Pronotal laterobasal setae situated at lateral margins; apical hook of the right paramere very small; parascutellar setae absent sometimes;
<collectingCountry id="51876EBCFA7CFFFC3D3F4D09FE6DD94F" box="[341,396,1880,1902]" name="Spain" pageId="2" pageNumber="92">Spain</collectingCountry>
..........................................................................
<taxonomicName id="EE9055AFFA7CFFFC39624D09FA7DD94C" ID-CoL="8KV6M" authorityName="Heyden" authorityYear="1880" box="[1288,1436,1880,1901]" class="Insecta" family="Carabidae" genus="Anchomenidius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC39624D09FA7DD94C" box="[1288,1436,1880,1901]" italics="true" pageId="2" pageNumber="92">Anchomenidius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="92643689FA7CFFFC3CFD4D27FA7DD988" pageId="2" pageNumber="92">
<paragraph id="292F2E2CFA7CFFFC3CFD4D27FA7DD9AA" blockId="2.[151,1437,1587,2020]" box="[151,1436,1910,1932]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4D27FA7DD9AA" box="[151,1436,1910,1932]" pageId="2" pageNumber="92">
4. Parascutellar setae absent........................................................................
<taxonomicName id="EE9055AFFA7CFFFC39564D27FA7DD9AA" ID-CoL="8KTVZ" authorityName="Semenov" authorityYear="1889" box="[1340,1436,1910,1931]" class="Insecta" family="Carabidae" genus="Acalathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC39564D27FA7DD9AA" box="[1340,1436,1910,1931]" italics="true" pageId="2" pageNumber="92">Acalathus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="292F2E2CFA7CFFFC3CFD4DC5FA7DD988" blockId="2.[151,1437,1587,2020]" box="[151,1436,1939,1961]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4DC5FA7DD988" box="[151,1436,1939,1961]" pageId="2" pageNumber="92">- Parascutellar setae present.............................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="92643689FA7CFFFD3CFD4DE1FA7DDEEA" lastPageId="3" lastPageNumber="93" pageId="2" pageNumber="92">
<paragraph id="292F2E2CFA7CFFFC3CFD4DE1FA7CD9C2" blockId="2.[151,1437,1587,2020]" pageId="2" pageNumber="92">
<keyLead id="92618D19FA7CFFFC3CFD4DE1FA7CD9C2" pageId="2" pageNumber="92">
5. Elytral interval 3 without setigerous pore; lateral grooves absent on metatarsomeres I–IV; apical hook of the right paramere present; Himalaya (
<collectingCountry id="51876EBCFA7CFFFC3DF44D9FFE3AD9C5" box="[414,475,1998,2020]" name="China" pageId="2" pageNumber="92">China</collectingCountry>
,
<collectingCountry id="51876EBCFA7CFFFC3D8E4D9FFDA2D9C2" box="[484,579,1998,2019]" name="Myanmar" pageId="2" pageNumber="92">Myanmar</collectingCountry>
,
<collectingCountry id="51876EBCFA7CFFFC3E274D9FFD68D9C5" box="[589,649,1998,2020]" name="Nepal" pageId="2" pageNumber="92">Nepal</collectingCountry>
, and
<collectingCountry id="51876EBCFA7CFFFC3ED14D9FFCE3D9C5" box="[699,770,1998,2020]" name="Bhutan" pageId="2" pageNumber="92">Bhutan</collectingCountry>
)...............................................
<taxonomicName id="EE9055AFFA7CFFFC392C4D9FFA7CD9C2" ID-CoL="8KZJC" authorityName="Andrewes" authorityYear="1937" box="[1350,1437,1998,2019]" class="Insecta" family="Carabidae" genus="Xestopus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7CFFFC392C4D9FFA7CD9C2" box="[1350,1437,1998,2019]" italics="true" pageId="2" pageNumber="92">Xestopus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="292F2E2CFA7DFFFD3CFD4AC9FA7DDEEA" blockId="3.[151,1437,151,349]" pageId="3" pageNumber="93">
<keyLead id="92618D19FA7DFFFD3CFD4AC9FA7DDEEA" pageId="3" pageNumber="93">- Elytral interval 3 with at least one setigerous pore; lateral grooves present on metatarsomeres I–IV; apical hook of the right paramere absent...................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="92643689FA7DFFFD3CFD4A83FA7DDF24" pageId="3" pageNumber="93">
<paragraph id="292F2E2CFA7DFFFD3CFD4A83FA7CDEC6" blockId="3.[151,1437,151,349]" box="[151,1437,210,232]" pageId="3" pageNumber="93">
<keyLead id="92618D19FA7DFFFD3CFD4A83FA7CDEC6" box="[151,1437,210,232]" pageId="3" pageNumber="93">
6. Lateral bead of prosternal process present; almost all the Palearctic Region, from Eastern Asia to Western Europe...
<taxonomicName id="EE9055AFFA7DFFFD392C4A83FA7CDEC6" ID-CoL="8KWR2" authorityName="Bonelli" authorityYear="1810" box="[1350,1437,210,231]" class="Insecta" family="Carabidae" genus="Dolichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7DFFFD392C4A83FA7CDEC6" box="[1350,1437,210,231]" italics="true" pageId="3" pageNumber="93">Dolichus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="292F2E2CFA7DFFFD3CFD4AA1FA7DDF24" blockId="3.[151,1437,151,349]" box="[151,1436,239,261]" pageId="3" pageNumber="93">
<keyLead id="92618D19FA7DFFFD3CFD4AA1FA7DDF24" box="[151,1436,239,261]" pageId="3" pageNumber="93">
- Lateral bead of prosternal process absent; Eastern Asia (
<collectingCountry id="51876EBCFA7DFFFD3E964ABEFCD8DF24" box="[764,825,239,261]" name="China" pageId="3" pageNumber="93">China</collectingCountry>
,
<collectingCountry id="51876EBCFA7DFFFD3F294ABEFC5FDF24" box="[835,958,239,261]" name="North Korea" pageId="3" pageNumber="93">North Korea</collectingCountry>
, and
<collectingCountry id="51876EBCFA7DFFFD3F9A4ABEFB8BDF24" box="[1008,1130,239,261]" name="South Korea" pageId="3" pageNumber="93">South Korea</collectingCountry>
)....................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="92643689FA7DFFFD3CFD4B5CFA7DDF7D" pageId="3" pageNumber="93">
<paragraph id="292F2E2CFA7DFFFD3CFD4B5CFA7DDF03" blockId="3.[151,1437,151,349]" box="[151,1436,269,291]" pageId="3" pageNumber="93">
<keyLead id="92618D19FA7DFFFD3CFD4B5CFA7DDF03" box="[151,1436,269,291]" pageId="3" pageNumber="93">
7. Pronotal laterobasal seta absent;
<collectingCountry id="51876EBCFA7DFFFD3E764B5CFDB4DF02" box="[540,597,269,291]" name="China" pageId="3" pageNumber="93">China</collectingCountry>
(
<collectingRegion id="EB54E0CEFA7DFFFD3E084B5CFD53DF02" box="[610,690,269,291]" country="China" name="Shaanxi" pageId="3" pageNumber="93">Shaanxi</collectingRegion>
)...................................................
<taxonomicName id="EE9055AFFA7DFFFD39754B5CFA7DDF03" ID-CoL="8KWR3" authorityName="Sciaky &amp; Wrase" authorityYear="1998" box="[1311,1436,269,290]" class="Insecta" family="Carabidae" genus="Doliodactyla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7DFFFD39754B5CFA7DDF03" box="[1311,1436,269,290]" italics="true" pageId="3" pageNumber="93">Doliodactyla</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="292F2E2CFA7DFFFD3CFD4B7AFA7DDF7D" blockId="3.[151,1437,151,349]" pageId="3" pageNumber="93">
<keyLead id="92618D19FA7DFFFD3CFD4B7AFA7DDF7D" pageId="3" pageNumber="93">
- Pronotal laterobasal seta present;
<collectingCountry id="51876EBCFA7DFFFD3E4C4B7BFDBEDF61" box="[550,607,298,320]" name="China" pageId="3" pageNumber="93">China</collectingCountry>
(
<collectingRegion id="EB54E0CEFA7DFFFD3E064B7BFB4ADF61" box="[620,1195,298,320]" isEnumeration="true" pageId="3" pageNumber="93">Shaanxi, Gansu, Sichuan, Yunnan, Chongqing, Hubei, Jilin</collectingRegion>
),
<collectingCountry id="51876EBCFA7DFFFD38D64B7BFAD4DF61" box="[1212,1333,298,320]" name="North Korea" pageId="3" pageNumber="93">North Korea</collectingCountry>
and
<collectingCountry id="51876EBCFA7DFFFD390E4B7BFEFCDF7C" name="South Korea" pageId="3" pageNumber="93">South Korea</collectingCountry>
...................................................................................
<taxonomicName id="EE9055AFFA7DFFFD39624B16FA7DDF7D" ID-CoL="8KXVJ" authorityName="Semenov" authorityYear="1889" box="[1288,1436,327,348]" class="Insecta" family="Carabidae" genus="Morphodactyla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="1BE4F23EFA7DFFFD39624B16FA7DDF7D" box="[1288,1436,327,348]" italics="true" pageId="3" pageNumber="93">Morphodactyla</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink