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<document id="F6EFB7861CE6F7CBE984B54DC915717A" ID-DOI="10.11646/zootaxa.4813.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3944139" ID-ZooBank="0ED34D69-0BC1-4E7D-A50D-6C0A31AB0374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594719214043" checkinUser="plazi" docAuthor="Jordal, Bjarte H. &amp; Smith, Sarah M." docDate="2020" docId="9E3D1C0AFB3CFFBEFF62FF731856F9D9" docLanguage="en" docName="zootaxa.4813.1.1.pdf" docOrigin="Zootaxa 4813 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scolytodes maestus Jordal and Smith 2020, sp. nov." docType="treatment" docUuid="38605E3A-7EF1-4008-88E6-52013763FAAF" docUuidSource="ZooBank" docVersion="5" lastPageNumber="51" masterDocId="62046472FB0EFF8CFFF5FFEA1918FF8A" masterDocTitle="Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences" masterLastPageNumber="67" masterPageNumber="1" pageNumber="51" updateTime="1698781377542" updateUser="plazi">
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<mods:title id="DC41A9889011ED364330973C839F31FE">Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences</mods:title>
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<mods:namePart id="A2E19C65DC67866440B8754CB3B1DAD8">Jordal, Bjarte H.</mods:namePart>
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<mods:namePart id="6647D5CA3284D114D4C093BB1A59D10C">Smith, Sarah M.</mods:namePart>
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<mods:typeOfResource id="E36579E72CDFAB172A45D5B6C91BBF1D">text</mods:typeOfResource>
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<mods:title id="495304BB4364AAE782697A4BE2B7EC8A">Zootaxa</mods:title>
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<mods:part id="DEE7CCC2139C67DB058274E6549B8DB8">
<mods:date id="D3B2A55FE6B5A6687AACEF36ABF395D4">2020</mods:date>
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<mods:number id="7C80A74CFD6CFF359E35124975B7026D">2020-07-14</mods:number>
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<mods:number id="6AB0C7A4107AEAA4F1F8EB5624150981">4813</mods:number>
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<mods:classification id="DC5AC5DCB81CAC182209A1311EAEE75A">journal article</mods:classification>
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<mods:identifier id="93FAB343A504A9CCEE722F525F34C1A4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="9E3D1C0AFB3CFFBEFF62FF731856F9D9" ID-DOI="http://doi.org/10.5281/zenodo.4338684" ID-Zenodo-Dep="4338684" LSID="urn:lsid:zoobank.org:act:38605E3A-7EF1-4008-88E6-52013763FAAF" httpUri="http://treatment.plazi.org/id/9E3D1C0AFB3CFFBEFF62FF731856F9D9" lastPageNumber="51" pageId="50" pageNumber="51">
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<paragraph id="162BAD1CFB3CFFBEFF62FF731BD5FF3E" blockId="50.[151,947,153,252]" box="[151,717,153,180]" pageId="50" pageNumber="51">
<heading id="4D631A70FB3CFFBEFF62FF731BD5FF3E" bold="true" box="[151,717,153,180]" fontSize="11" level="1" pageId="50" pageNumber="51" reason="1">
<emphasis id="24E0710EFB3CFFBEFF62FF731BD5FF3E" bold="true" box="[151,717,153,180]" pageId="50" pageNumber="51">
<taxonomicName id="D194D69FFB3CFFBEFF62FF731B7BFF39" authority="Jordal and Smith" authorityName="Jordal and Smith" authorityYear="2020" box="[151,611,153,180]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="maestus" status="sp. nov.">
<emphasis id="24E0710EFB3CFFBEFF62FF731898FF3E" bold="true" box="[151,384,153,180]" italics="true" pageId="50" pageNumber="51">Scolytodes maestus</emphasis>
Jordal and Smith
</taxonomicName>
,
<taxonomicNameLabel id="3FD3CC75FB3CFFBEFD85FF701BD5FF3E" box="[624,717,154,180]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
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<subSubSection id="5E8EFE97FB3CFFBEFF62FF0B1863FF71" box="[151,379,225,252]" pageId="50" pageNumber="51" type="description">
<paragraph id="162BAD1CFB3CFFBEFF62FF0B1863FF71" blockId="50.[151,947,153,252]" box="[151,379,225,252]" pageId="50" pageNumber="51">
(
<figureCitation id="8EAFB199FB3CFFBEFF6AFF0B186BFF71" box="[159,371,225,252]" captionStart="FIGURES 100108" captionStartId="49.[151,264,1830,1855]" captionTargetBox="[158,1434,253,1805]" captionTargetId="figure@49.[152,1435,181,1805]" captionTargetPageId="49" captionText="FIGURES 100108. Dorsal, lateral and frontal views of the female holotype of Scolytodes amictus (100, 103, 106), Scolytodes cnesinoides (101, 104, 107), and Scolytodes maestus (102, 105, 108)." figureDoi="http://doi.org/10.5281/zenodo.3944163" httpUri="https://zenodo.org/record/3944163/files/figure.png" pageId="50" pageNumber="51">Figs 102, 105, 108</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB3CFFBEFF62FEC01B65FE25" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="162BAD1CFB3CFFBEFF62FEC01B65FE25" blockId="50.[151,1437,297,1620]" pageId="50" pageNumber="51">
<emphasis id="24E0710EFB3CFFBEFF62FEC0185EFECE" bold="true" box="[151,326,297,324]" pageId="50" pageNumber="51">Type material.</emphasis>
<materialsCitation id="A6FCA741FB3CFFBEFEBBFEC01ACDFEED" collectingDate="1998-10" collectorName="T. L. Erwin" country="Ecuador" elevation="235" latitude="-0.6319445" location="Tiputini" longLatPrecision="21" longitude="-76.144165" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
<emphasis id="24E0710EFB3CFFBEFEBBFEC01B3AFECE" bold="true" box="[334,546,297,324]" pageId="50" pageNumber="51">
<typeStatus id="C92F13BEFB3CFFBEFEBBFEC018A5FECE" box="[334,445,298,324]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
, female:
</emphasis>
<collectingCountry id="6E83ED8CFB3CFFBEFDDFFEC31B94FEC9" box="[554,652,297,323]" name="Ecuador" pageId="50" pageNumber="51">Ecuador</collectingCountry>
:
<collectingRegion id="D45063FEFB3CFFBEFD62FEC01A01FECE" box="[663,793,298,324]" country="Ecuador" name="Napo" pageId="50" pageNumber="51">Napo Prov.</collectingRegion>
,
<location id="134BFBC7FB3CFFBEFCD1FEC31A65FEC9" LSID="urn:lsid:plazi:treatment:9E3D1C0AFB3CFFBEFF62FF731856F9D9:134BFBC7FB3CFFBEFCD1FEC31A65FEC9" box="[804,893,297,323]" country="Ecuador" latitude="-0.6319445" longLatPrecision="21" longitude="-76.144165" name="Tiputini" pageId="50" pageNumber="51" stateProvince="Napo">Tiputini</location>
biodiversity station,
<quantity id="D16C00F9FB3CFFBEFB99FEC31DF6FECE" box="[1132,1262,297,324]" metricMagnitude="2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="50" pageNumber="51" unit="m" value="235.0" valueMax="250.0" valueMin="220.0">
<elevation id="9DB94A2FFB3CFFBEFB99FEC31DF6FECE" box="[1132,1262,297,324]" metricMagnitude="2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="50" pageNumber="51" unit="m" value="235.0" valueMax="250.0" valueMin="220.0">220250 m</elevation>
</quantity>
,
<date id="622A8BDCFB3CFFBEFB0EFEC31C80FEC9" box="[1275,1432,297,323]" pageId="50" pageNumber="51" value="1998-10">
<collectingDate id="726E7234FB3CFFBEFB0EFEC31C80FEC9" box="[1275,1432,297,323]" pageId="50" pageNumber="51" value="1998-10">October 1998</collectingDate>
</date>
,
<geoCoordinate id="73A0CBDBFB3CFFBEFF62FEA71834FEE2" box="[151,300,333,360]" degrees="00" direction="south" minutes="37" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="55" value="-0.6319445">00° 3755S</geoCoordinate>
,
<geoCoordinate id="73A0CBDBFB3CFFBEFEC9FEA718C2FEED" box="[316,474,333,359]" degrees="76" direction="west" minutes="08" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="39" value="-76.144165">76° 0839W</geoCoordinate>
,
<collectorName id="BB61C8CAFB3CFFBEFE1CFEA41B7EFEED" box="[489,614,333,360]" pageId="50" pageNumber="51">T.L. Erwin</collectorName>
<emphasis id="24E0710EFB3CFFBEFD84FEA41BB5FEED" box="[625,685,333,359]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
collectors, indiv#001204
</materialsCitation>
.
<emphasis id="24E0710EFB3CFFBEFC11FEA71D85FEE2" bold="true" box="[996,1181,333,360]" pageId="50" pageNumber="51">
<typeStatus id="C92F13BEFB3CFFBEFC11FEA71D53FEE2" box="[996,1099,333,360]" pageId="50" pageNumber="51" type="allotype">Allotype</typeStatus>
, male:
</emphasis>
same data as HT, indiv#000758.
<materialsCitation id="A6FCA741FB3CFFBEFED9FE981A2EFE01" box="[300,822,369,396]" collectingDate="1998-06" collectorName="T. L. Erwin" country="Ecuador" elevation="235" latitude="-0.6319445" location="Tiputini" longLatPrecision="21" longitude="-76.144165" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="24E0710EFB3CFFBEFED9FE9818BAFE06" bold="true" box="[300,418,370,396]" pageId="50" pageNumber="51">
<typeStatus id="C92F13BEFB3CFFBEFED9FE981885FE06" box="[300,413,370,396]" pageId="50" pageNumber="51" type="paratype">Paratype</typeStatus>
:
</emphasis>
same data as HT, except
<date id="622A8BDCFB3CFFBEFD36FE981A2EFE01" box="[707,822,369,396]" pageId="50" pageNumber="51" value="1998-06">
<collectingDate id="726E7234FB3CFFBEFD36FE981A2EFE01" box="[707,822,369,396]" pageId="50" pageNumber="51" value="1998-06">June 1998</collectingDate>
</date>
</materialsCitation>
(1).
<materialsCitation id="A6FCA741FB3CFFBEFC98FE9B19F0FE25" collectionCode="USNM" country="Ecuador" pageId="50" pageNumber="51" specimenCount="2" stateProvince="Napo" typeStatus="holotype">
<typeStatus id="C92F13BEFB3CFFBEFC98FE9B1ACDFE01" box="[877,981,369,395]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
and
<typeStatus id="C92F13BEFB3CFFBEFBF8FE9B1D7FFE01" box="[1037,1127,369,395]" pageId="50" pageNumber="51" type="allotype">allotype</typeStatus>
temporarily held in trust at
<collectionCode id="708535D9FB3CFFBEFF62FE7F19F0FE25" box="[151,232,405,431]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="50" pageNumber="51" type="Museum">USNM</collectionCode>
</materialsCitation>
for
<collectionCode id="708535D9FB3CFFBEFEE2FE7F1873FE25" box="[279,363,405,431]" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" pageId="50" pageNumber="51">MECN</collectionCode>
,
<materialsCitation id="A6FCA741FB3CFFBEFE83FE7C1B61FE25" box="[374,633,405,432]" collectionCode="QCAZ" country="Ecuador" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
one
<typeStatus id="C92F13BEFB3CFFBEFE50FE7C1B1DFE3A" box="[421,517,406,432]" pageId="50" pageNumber="51" type="paratype">paratype</typeStatus>
in
<collectionCode id="708535D9FB3CFFBEFDDCFE7F1B61FE25" box="[553,633,405,431]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="50" pageNumber="51" type="Museum">QCAZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB3CFFBEFF32FE531806FD0D" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="162BAD1CFB3CFFBEFF32FE531806FD0D" blockId="50.[151,1437,297,1620]" pageId="50" pageNumber="51">
<emphasis id="24E0710EFB3CFFBEFF32FE53185AFE5E" bold="true" box="[199,322,441,468]" pageId="50" pageNumber="51">Diagnosis.</emphasis>
Interstriae 10 sharply elevated to level of metacoxae (continue as a faint trace to apex). Protibiae with an additional mesal tooth near tarsal insertion. Pronotum asperate on anterior half. Interstriae with small granules at base of setae, particularly on elytral disc. Belongs to the same group of species as
<taxonomicName id="D194D69FFB3CFFBEFB99FDE81C84FD91" authority="(Schedl, 1940)" baseAuthorityName="Schedl" baseAuthorityYear="1940" box="[1132,1436,513,540]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="24E0710EFB3CFFBEFB99FDE81DEAFD91" box="[1132,1266,513,540]" italics="true" pageId="50" pageNumber="51">S. rugicollis</emphasis>
(Schedl, 1940)
</taxonomicName>
and
<taxonomicName id="D194D69FFB3CFFBEFF30FDCC18FBFDB5" authority="Jordal and Smith" authorityName="Jordal and Smith" box="[197,483,549,576]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="viteus">
<emphasis id="24E0710EFB3CFFBEFF30FDCC1839FDB5" box="[197,289,550,576]" italics="true" pageId="50" pageNumber="51">S. viteus</emphasis>
Jordal and Smith
</taxonomicName>
(described below), but distinguished by the presence of erect setae on each interstriae. Also similar to
<taxonomicName id="D194D69FFB3CFFBEFEB0FDA01B2FFDE9" authority="Wood, 1982" authorityName="Wood" authorityYear="1982" box="[325,567,585,612]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pilifer">
<emphasis id="24E0710EFB3CFFBEFEB0FDA018BDFDE9" box="[325,421,585,612]" italics="true" pageId="50" pageNumber="51">S. pilifer</emphasis>
<bibRefCitation id="7205D0EDFB3CFFBEFE59FDA31B2FFDE9" author="Wood, S. L." box="[428,567,585,611]" pageId="50" pageNumber="51" pagination="1 - 1359" refId="ref46257" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood, 1982</bibRefCitation>
</taxonomicName>
, but the latter has a dense brush of setae in the female frons, and a more stout body shape.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB3CFFBEFF32FD7B1811FAD2" pageId="50" pageNumber="51" type="description">
<paragraph id="162BAD1CFB3CFFBEFF32FD7B1B36FA9A" blockId="50.[151,1437,297,1620]" pageId="50" pageNumber="51">
<emphasis id="24E0710EFB3CFFBEFF32FD7B18ADFD26" bold="true" box="[199,437,657,684]" pageId="50" pageNumber="51">Description, female.</emphasis>
Length
<quantity id="D16C00F9FB3CFFBEFDFAFD7B1B8FFD26" box="[527,663,657,684]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="50" pageNumber="51" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 mm</quantity>
, 2.52.6 × as long as wide; colour dark brown.
<emphasis id="24E0710EFB3CFFBEFB6AFD781DFEFD26" bold="true" box="[1183,1254,658,684]" pageId="50" pageNumber="51">Head.</emphasis>
Eyes almost entire, a tiny notch at level of antennal insertion, separated above by 2.5 × their width. Frons lightly impressed on little more than central half, punctured in impressed area except on lower central third which is impunctate and lightly elevated; vestiture consisting of sparse, short setae in impressed area and on epistoma. Antennal club pilose with two obliquely procurved sutures, segment 1 and 2 partly corneous. Funiculus 5-segmented.
<emphasis id="24E0710EFB3CFFBEFB59FCC81C3DFCB6" bold="true" box="[1196,1317,802,828]" pageId="50" pageNumber="51">Pronotum</emphasis>
reticulate, subshining, punctures distinct, spaced by 12 × their diameter, punctures not reaching anterior margin, rough asperities present on anterior half, anterior row of asperities indistinctly elevated. Vestiture consisting of 8 long, erect setae (4-2-2) and some additional, shorter, erect setae along anterior margin and anterior third of pronotum.
<emphasis id="24E0710EFB3CFFBEFAFDFC641C4CFC22" bold="true" box="[1288,1364,910,936]" pageId="50" pageNumber="51">Elytra</emphasis>
shiny, smooth, except a small granule at base of interstrial setae; striae mostly regular, striae 1 slightly impressed, others not, small punctures spaced by 23 × their diameter, a microscopic granule associated with each puncture; interstrial punctures microscopic, widely spaced. Interstriae 10 sharply raised to level of metacoxae, with a faint trace continuing to ventrite 3. Vestiture consisting of erect, fine, interstrial setae and much finer, semirecumbent strial setae.
<emphasis id="24E0710EFB3CFFBEFAA8FBF41C84FBB2" bold="true" box="[1373,1436,1054,1080]" pageId="50" pageNumber="51">Legs.</emphasis>
Procoxae separated by 0.40.5 × the width of one procoxa. Mesocoxae separated by 0.8 × the width of a procoxa. Protibiae narrow, distal teeth 1 similar to 2, with 34 lateral, tiny, sharp granules along a sharp flange towards tibial base; a sharp additional mesal tooth present on posterior face near tarsal insertion; protibial mucro very short to obtuse, curved posteriorly. Meso- and metatibiae with 6 and 5 socketed teeth on distal half and one-third, respectively.
<emphasis id="24E0710EFB3CFFBEFF62FB381847FB66" bold="true" box="[151,351,1233,1260]" pageId="50" pageNumber="51">Ventral vestiture</emphasis>
. Setae on mesanepisternum and metanepisternum bifid or trifid, on metasternum bifid anteriorly, simple on posterior and ventral part.
</paragraph>
<paragraph id="162BAD1CFB3CFFBEFF32FAF01811FAD2" blockId="50.[151,1437,297,1620]" pageId="50" pageNumber="51">
<emphasis id="24E0710EFB3CFFBEFF32FAF01813FABE" bold="true" box="[199,267,1306,1332]" pageId="50" pageNumber="51">Male.</emphasis>
Similar to female except frons convex, shiny, impressed on lower third, with few punctures and scant, fine setae.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB3CFFBEFF32FA8818CDFA4E" pageId="50" pageNumber="51" type="key">
<paragraph id="162BAD1CFB3CFFBEFF32FA8818CDFA4E" blockId="50.[151,1437,297,1620]" pageId="50" pageNumber="51">
<emphasis id="24E0710EFB3CFFBEFF32FA8818B8FAF6" bold="true" box="[199,416,1377,1404]" pageId="50" pageNumber="51">
Key (
<bibRefCitation id="7205D0EDFB3CFFBEFEFDFA8B188BFAF6" author="Wood, S. L." box="[264,403,1377,1404]" pageId="50" pageNumber="51" refId="ref46297" refString="Wood, S. L. (2007) Bark and ambrosia beetles of South America (Coleoptera, Scolytidae). Brigham Young University Press, Provo, Utah, 900 pp." type="book" year="2007">Wood 2007</bibRefCitation>
).
</emphasis>
Keys to couplet 87, near
<taxonomicName id="D194D69FFB3CFFBEFD3FFA881D11FAF1" ID-CoL="4VT4F" authority="Wood, 1969" authorityName="Wood" authorityYear="1969" box="[714,1033,1377,1404]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="erineophilus">
<emphasis id="24E0710EFB3CFFBEFD3FFA881A6CFAF1" box="[714,884,1377,1404]" italics="true" pageId="50" pageNumber="51">S. erineophilus</emphasis>
Wood, 1969
</taxonomicName>
, but differs by the simple, erect interstrial setae, the female frons, the more elongated and broadly rounded pronotum with sharper asperities, and the more widely separated eyes.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB3CFFBEFF32FA241B07F981" pageId="50" pageNumber="51" type="etymology">
<paragraph id="162BAD1CFB3CFFBEFF32FA241B07F981" blockId="50.[151,1437,297,1620]" pageId="50" pageNumber="51">
<emphasis id="24E0710EFB3CFFBEFF32FA241856FA62" bold="true" box="[199,334,1486,1512]" pageId="50" pageNumber="51">Etymology.</emphasis>
The Latin name
<taxonomicName id="D194D69FFB3CFFBEFDE2FA241B69FA6D" authorityName="Jordal and Smith" authorityYear="2020" box="[535,625,1486,1511]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="maestus">
<emphasis id="24E0710EFB3CFFBEFDE2FA241B69FA6D" box="[535,625,1486,1511]" italics="true" pageId="50" pageNumber="51">maestus</emphasis>
</taxonomicName>
is an adjective, meaning sad or melancholic, referring to the dark and gloomy appearance of this species.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB3CFFBEFF32F9FF1856F9D9" pageId="50" pageNumber="51" type="biology_ecology">
<paragraph id="162BAD1CFB3CFFBEFF32F9FF1856F9D9" blockId="50.[151,1437,297,1620]" pageId="50" pageNumber="51">
<emphasis id="24E0710EFB3CFFBEFF32F9FF18E8F9BA" bold="true" box="[199,496,1557,1584]" pageId="50" pageNumber="51">Biology and distribution.</emphasis>
Only known from two collecting events in the Ecuadorian Amazon lowland rainforest
<typeStatus id="C92F13BEFB3CFFBEFF48F9D019F5F9DE" box="[189,237,1594,1620]" pageId="50" pageNumber="51">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>